NORTHERN SUDAN

WADI MILK FORMATION

?CENOMANIAN STAGE

GALLIC SUBEPOCH

LOWER GULF EPOCH

MIDDLE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990

COHORT: CROCODYLOTARSI BENTON & CLARK, 1988

MAGNORDER: SUCHIA KREBS, 1974

SUPERORDER: CROCODYLOMORPHA WALKER, 1968

GRANDORDER: CROCODILIFORMES HAY, 1930

MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983

PARVORDER: METASUCHIA BENTON & CLARK, 1988

GONDWANASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

NOTOSUCHIMORPHA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

NOTOSUCHIFORMES DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

TERRIASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

PEIROSAURIMORPHA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

Family: CANDIDODONTIDAE de Souza Carvalho, Ribeiro & dos Santos Avilla, 2004

PEIROSAURIFORMES DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004

Superfamily: PEIROSAURIDEA Gasparini, 1982 emend de Souza Carvalho, Ribeiro & dos Santos Avilla, 2004

Family: PEIROSAURIDAE Gasparini, 1982

Gen. sp indet.

_____________________________________________________________________________________

NORTHERN SUDAN

ABOUT 200 KM NORTHWEST OF KHARTOUM

F1/89, F2/89

UPPERMOST ALBIAN-LOWERMOST CENOMANIAN STAGE

UPPER GALLIC SUBEPOCH

LOWEST GULF EPOCH

LATE EARLY TO EARLY MIDDLE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: CHELONOMORPHA KUHN, 1960

SUPERCOHORT: TESTUDINES LINNAEUS, 1758

INCERTAE SEDIS

Gen. sp indet.

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990

COHORT: CROCODYLOTARSI BENTON & CLARK, 1988

MAGNORDER: SUCHIA KREBS, 1974

SUPERORDER: CROCODYLOMORPHA WALKER, 1968

GRANDORDER: CROCODILIFORMES HAY, 1930

MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983

ORDER: MESOSUCHIA EMEND HUXLEY, 1875

PARVORDER: METASUCHIA BENTON & CLARK, 1988

INFRAORDER: TETHYSUCHIA BUFFETAUT, 1982

Family: DYROSAURIDAE de STEFANO, 1903

Gen. sp indet.

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

INCERTAE SEDIS

Gen. sp indet.

COHORT: ORNITHISCHIFORMES COOPER, 1985

SUPERORDER: SAUROPODOMORPHA HUENE, 1932

GRANDORDER: BRONTOSAURIA OLSHEVSKY, 1991 (emend)

MICRORDER: SAUROPODA MARSH, 1878

ORDER: EUSAUROPODA

SUBORDER: NEOSAUROPODA BONAPARTE, 1986

INFRAORDER: MACRONARIA WILSON & SERENO, 1998

PARVORDER: TITANOSAURIFORMES SALGADO ET AL, 1997

Superfamily: TITANOSAUROIDEA Lydekker, 1885 (sensu Upchurch, 1999)

Family: TITANOSAURIDAE Lydekker, 1885

Gen. sp indet.

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: CHELONOMORPHA KUHN, 1960

SUPERCOHORT: TESTUDINES LINNAEUS, 1758

Incertae sedis

Genus sp indet.

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990

COHORT: CROCODYLOTARSI BENTON & CLARK, 1988

MAGNORDER: SUCHIA KREBS, 1974

SUPERORDER: CROCODYLOMORPHA WALKER, 1968

ORDER: MESOSUCHIA EMEND HUXLEY, 1875

SUBORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983

Incertae sedis

Gen. sp indet.

 

WADI ABU HASHIM

SHENDI FORMATION

CENOMANIAN

MIDDLE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

ORDER: Incertae sedis

Gen. sp indet.

 

SUDAN

C. 200 KM NORTH-WEST OF KHARTOUM

WADI ABU HASHIM

HASHIM MEMBER

WADI MILK FORMATION

CENOMANIAN STAGE

UPPER GALIC SUBEPOCH

LOWER UPPER UPPER GULF EPOCH

MIDDLE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: AMPHIBIA LINNAEUS, 1758

SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?

ORDER: GYMNOPHIONA RAFINESQUE, 1814

Incertae sedis

Gen. sp indet.

SUPERORDER: BATRACHIA BRONGNIART, 1800

ORDER: CAUDATA SCOPOLI, 1777

Incertae sedis

Gen. sp indet.

SUBORDER: SALAMANDROIDEA NOBLE, 1931

Family: SIRENIDAE Gray, 1825

*Kababisha sudanensis
*Kababisha humarensis
Kababisha sp

SUPERORDER: SALIENTIA LAURENTI, 1768

Incertae sedis

Gen. sp indet.

ORDER: ANURA RAFINESQUE, 1815

SUBORDER: NEOBATRACHIA REIG, 1958

Superfamily: RANOIDEA Gray, 1825 (sensu Fitzinger, 1826)

Family: RANIDAE GRAY, 1825

Subfamly: RANINAE Gray, 1825

Rana sp

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: CHELONOMORPHA KUHN, 1960

SUPERCOHORT: TESTUDINES LINNAEUS, 1758

COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988

MAGNORDER: PLEURODIRA GAFFNEY & MEYLAN, 1988

ORDER: PLEURODIRIDA COPE, 1864

SUBORDER: EUPLEURODIRA GAFFNEY & MEYLAN, 1988

Superfamily: PELOMEDUSOIDES Cope, 1868 Cope, 1868

Family: PELOMEDUSIDAE Cope, 1868

Gen. sp indet.

 

SUDAN

WADI ABU HASHIM

WADI MAILK FORMATION

CENOMANIAN STAGE

UPPER GALLIC SUBEPOCH

LOWERMOST GULF EPOCH

MIDDLE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: SERPENTES LINNAEUS, 1759

ORDER: OPHIDIA BRONGNIART, 1800

SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923

Superfamily: ACROCHORDOIDEA Bonaparte, 1838

Family: PALAEOPHIDAE LYDEKKER, 1888

Gen. sp indet.

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990

COHORT: CROCODYLOTARSI BENTON & CLARK, 1988

MAGNORDER: SUCHIA KREBS, 1974

SUPERORDER: CROCODYLOMORPHA WALKER, 1968

GRANDORDER: CROCODILIFORMES HAY, 1930

MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983

Incertae sedis

Gen. sp indet

ORDER: THALATTOSUCHIA EMEND FRAAS, 1901

Family: TELEOSAURIDAE Geoffroy, 1831

Machimosaurus sp

ORDER: MESOSUCHIA EMEND HUXLEY, 1875

PARVORDER: METASUCHIA BENTON & CLARK, 1988

INFRAORDER: METAMESOSUCHIA HULKE, 1878

NEOSUCHIA CLARK, IN BENTON & CLARK, 1988

Family: TREMATOCHAMPSIDAE Buffetaut, 1974

Incertae sedis

Gen. sp indet.

INFRAORDER: TETHYSUCHIA BUFFETAUT, 1982

Family: DYROSAURIDAE de STEFANO, 1903

Incertae sedis

Gen. sp indet.

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

Incertae sedis

Gen. sp indet.

COHORT: THEROPODA EMEND MARSH, 1881

ORDER: Incertae sedis

Gen. sp indet.

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: TETANURAE GAUTHIER, 1986

ORDER: NEOTETANURAE SERENO, WILSON, LARSSON, DUTHEIL &

SUBORDER: AVETHEROPODA PAUL, 1988

Superfamily: ALLOSAUROIDEA Marsh, 1878 (sensu Currie & Zhao, 1994)

Family: CARCHARODONTOSAURIDAE Stromer, 1931

Gen. sp indet.

GRANDORDER: TETANURAE GAUTHIER, 1986

MICRORDER: COELUROSAURIA HUENE, 1914

ORDER: MANIRAPTORFORMES HOLTZ, 1996

AVETHEROPOA PAUL, 2002

SUBORDER: COELURIA MARSH, 1881

Incertae sedis

Gen. sp indet.

AVEPECTORA PAUL, 2002

SUBORDER: ORNITHOMIMOSAURIA BARSBOLD, 1976

Family: ORNITHOMIMIDAE Marsh, 1890

Incertae sedis

Gen. sp indet.

SUBORDER: MANIRAPTORA GAUTHIER, 1986

INFRAORDER: PARVES SERENO, 1997

PARVORDER: DEINONYCHOSAURIA COLBERT & D. A. RUSSELL, 1969

Family: DROMAEOSAURIDAE Russell, 1969

Incerate sedis

Gen. sp indet.

COHORT: ORNITHISCHIFORMES COOPER, 1985

SUPERORDER: SAUROPODOMORPHA HUENE, 1932

Incertae sedis

Gen. sp indet.

GRANDORDER: BRONTOSAURIA OLSHEVSKY, 1991 (emend)

MICRORDER: SAUROPODA MARSH, 1878

ORDER: EUSAUROPODA

SUBORDER: NEOSAUROPODA BONAPARTE, 1986

INFRAORDER: HOMALOSAUROPIA JANENSCH, 1929

PARVORDER: DIPLODOCIMORPHA CALVO & SALGADO, 1995

Superfamily: DIPLODOCOIDEA Upchurch, 1995

Family: DIPLODOCIDAE Marsh, 1884

Subfamily: DICRAEOSAURINAE Janensch, 1929

Gen. sp indet.

INFRAORDER: MACRONARIA WILSON & SERENO, 1998

PARVORDER: TITANOSAURIFORMES SALGADO ET AL, 1997

Superfamily: TITANOSAUROIDEA Lydekker, 1885 (sensu Upchurch, 1999)

Family: TITANOSAURIDAE Lydekker, 1885

Subfamily: TITANOSAURINAE Lydekker, 1885 (sensu Nopcsa, 1928)

Gen. sp indet.

MAGNORDER: ORNITHISCHIA SEELEY, 1888

Incertae sedis

Gen. sp indet.

SUPERORDER: NEORNITHISCHIA COOPER, 1985

GRANDORDER: ORNITHOPODA MARSH, 1871

MICRORDER: EUORNITHOPODA SERENO, 1986

ORDER: HYPSILOPHODONTIA COOPER, 1983

Family: HYPSILOPHODONTIDAE Dollo, 1882

Incertae sedis? Or new families? (pending further research)

Gen. sp indet.

ORDER: IGUANODONTIA DOLLO, 1888

SUBORDER: ANKYLOPOLLEXIA SERENO, 1986

INFRAORDER: STYRACOSTERNA SERENO, 1986

PARVORDER: HADROSAURIFORMES SERENO, 1997

Superfamily: IGUANODONTIDEA Cope, 1869 (sensu Sereno, 1986)

Family: IGUANODONTIDAE Cope, 1869

Incertae sedis

Gen. sp indet.

Superfamily: OURANOSAURIDEA Nova

Family: OURANOSAURIDAE nova

cf. Ouranosaurus sp

WADI ABU HASHIM

200 KM NW OF KHARTOUM

WADI ABU HASHIM MEMBER

WADI MILK FORMATION

CENOMANIAN STAGE

UPPERMIDDLE GALLIC SUBEPOCH

LOWERMOST GULF EPOCH

LATE MIDDLE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: SERPENTES LINNAEUS, 1759

Incertae sedis

Gen. sp indet.

ORDER: OPHIDIA BRONGNIART, 1800

SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923

Superfamily: SIMOLIOPHEOIDEA Nopcsa, 1925

Family: LAPPARENTOPHEIDAE Hoffstetter, 1968

Genus a

Genus b

Superfamily: ANILIODEA Fitzinger, 1826

Family: ANILIIDAE Fitzenger, 1826

*Coniophis dabiebus
Coniophis cf. C. dabiebus

Superfamily: BOOIDEA Gray, 1825

Family: BOIDAE Gray, 1825

Subfamily: MADTSOINAE Hoffstetter, 1961

Gen. sp indet.

Superfamily: ACROCHORDOIDEA Bonaparte, 1838

Family: ?NIGEROPHEIDAE Rage, 1975

*Nubianophis afaahus
Nubianophis cf. N. afaahus

Family: PALAEOPHIDAE LYDEKKER, 1888

Gen. sp indet.

Superfamily: COLUBROIDEA Oppel, 1811

Incerte sedis

Gen. sp indet.

Family: RUSSELLOPHEIDAE Rage, 1978

*Krebsophis thobanus

SUPERORDER: LACERTILIA OWEN, 1842

Incertae sedis

Gen. sp indet.

SUDAN

WADI ABU HASHIM

SHENDI FORMATION

CENOMANIAN STAGE

UPPERMIDDLE GALLIC SUBEPOCH

LOWERMOST GULF EPOCH

MIDDLE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

ORDER: Incertae sedis

Gen. sp indet.

____________________________________________________________________________________

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: CHELONOMORPHA KUHN, 1960

SUPERCOHORT: TESTUDINES LINNAEUS, 1758

Incertae sedis

Genus sp indet.

SUBCLASS: DIAPSIDA OSBORN, 1903

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990

COHORT: CROCODYLOTARSI BENTON & CLARK, 1988

MAGNORDER: SUCHIA KREBS, 1974

SUPERORDER: CROCODYLOMORPHA WALKER, 1968

ORDER: MESOSUCHIA EMEND HUXLEY, 1875

SUBORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983

Incertae sedis

Gen. sp indet.

____________________________________________________________________________________

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: CHELONOMORPHA KUHN, 1960

SUPERCOHORT: TESTUDINES LINNAEUS, 1758

COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988

MAGNORDER: PLEURODIRA GAFFNEY & MEYLAN, 1988

ORDER: PLEURODIRIDA COPE, 1864

SUBORDER: EUPLEURODIRA GAFFNEY & MEYLAN, 1988

Superfamily: PELOMEDUSOIDES Cope, 1868 Cope, 1868

Family: PELOMEDUSIDAE Cope, 1868

Subfamily: PELOMEDUSINAE Cope, 1868

Bothremyid gen sp indet.

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990

COHORT: CROCODYLOTARSI BENTON & CLARK, 1988

MAGNORDER: SUCHIA KREBS, 1974

SUPERORDER: CROCODYLOMORPHA WALKER, 1968

GRANDORDER: CROCODILIFORMES HAY, 1930

MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983

ORDER: EUSUCHIA emend HUXLEY, 1875

INFRAORDER: CROCODYLIA emend GMELIN, 1877

PARVORDER: BREVIROSTRES

Superfamily: CROCODILOIDEA Cuvier, 1807 (sensu Fitzinger, 1826)

Family: CROCODYLIDAE Cuvier, 1807

Subfamily: CROCODYLIDINAE Cuvier, 1807

*Crocodilus checchiai
= Crocodilus checchiai var depressa