JAPAN

HONSHU ISLAND

OSAKA

HANNAN CITY

HAKOTSUKURI

SHINTACHI FORMATION

IZUMI GROUP

MAASTRICHTIAN STAGE

UPPER SENONIAN SUBEPOCH

UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: LACERTILIA OWEN, 1842

GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961

MICRORDER: SCINCOMORPHA CAMP, 1923

ORDER: ANGUIMORPHA FURBRINGER, 1900

SUBORDER: DIPLOGLOSSA COPE, 1864

INFRAORDER: THECAGLOSSA COPE, 1890

PARVORDER: MONSTERSAURIA NORELL & GAO, 1977

Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)

Family: MOSASAURIDAE Gervais, 1853

Infrafamily: NATANIA

Subfamily: MOSASAURINAE Gervais, 1853 (sensu Williston, 1897)

Gen. sp indet.

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SOUTHWEST JAPAN

HONSHU ISLAND

OSAKA PREFECTURE

SENNAN CITY

EAST SIDE OF SHOWAIKE POND

AZENOTANI MUDSTONE MEMBER

MUTSUO FORMATION

IZUMI GROUP

LOWER MAASTRICHTIAN STAGE

UPPER SENONIAN SUBEPOCH

UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: LACERTILIA OWEN, 1842

GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961

MICRORDER: SCINCOMORPHA CAMP, 1923

ORDER: ANGUIMORPHA FURBRINGER, 1900

SUBORDER: DIPLOGLOSSA COPE, 1864

INFRAORDER: THECAGLOSSA COPE, 1890

PARVORDER: MONSTERSAURIA NORELL & GAO, 1977

Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)

Family: MOSASAURIDAE Gervais, 1853

Infrafamily: NATANIA

Subfamily: MOSASAURINAE Gervais, 1853 (sensu Williston, 1897)

Gen. sp indet.

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JAPAN

HONSHU ISLAND

OSAKA PREFECTURE

IZUMISANO CITY

TAKINOIKE

IZUMI GROUP

MAASTRICHTIAN STAGE

UPPER SENONIAN SUBEPOCH

UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: CHELONOMORPHA KUHN, 1960

SUPERCOHORT: TESTUDINES LINNAEUS, 1758

COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988

MAGANORDER: CRYPTODIRA GAFFNEY & MEYLAN, 1988

SUPERORDER: SELMACRYPTODIRA GAFFNEY, HUTCHISON, JENKINS & MEEKR, 1987

GRANDORDER: DAIOCRYPTODIRA GAFFNEY & MEYLAN, 1988

MICRORDER: CENTROCRYPTODIRA GAFFNEY & MEYLAN, 1988

ORDER: POLYCRPTODIRA GAFFNEY & MEYLAN, 1988

SUBORDER: PROCOELOCRYPTODIRA GAFFNEY & MEYLAN, 1988

INFRAORDER: CHELONOIDEA GAFFNEY & MEYLAN, 1988

Superfamily: EUCHELONIOIDEA Gaffney & Meylan, 1988

Family: CHELONIIDAE Oppel, 1811

Incertae sedis

Gen sp indet.

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: LACERTILIA OWEN, 1842

GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961

MICRORDER: SCINCOMORPHA CAMP, 1923

ORDER: ANGUIMORPHA FURBRINGER, 1900

SUBORDER: DIPLOGLOSSA COPE, 1864

INFRAORDER: THECAGLOSSA COPE, 1890

PARVORDER: MONSTERSAURIA NORELL & GAO, 1977

Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)

Family: MOSASAURIDAE Gervais, 1853

Infrafamily: NATANIA

Subfamily: HALISAURINAE Bardet, Pereda Suberbiola, Iarochene, Bouya & Amaghzaz, 2005

Gen. sp indet.

Subfamily: MOSASAURINAE Gervais, 1853 (sensu Williston, 1897)

Tribe: MOSASAURINI Gervais, 1853 (sensu Russell, 1967)

Mosasaurus cf. prismaticus

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SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: LACERTILIA OWEN, 1842

GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961

MICRORDER: SCINCOMORPHA CAMP, 1923

ORDER: ANGUIMORPHA FURBRINGER, 1900

SUBORDER: DIPLOGLOSSA COPE, 1864

INFRAORDER: THECAGLOSSA COPE, 1890

PARVORDER: MONSTERSAURIA NORELL & GAO, 1977

Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)

Family: MOSASAURIDAE Gervais, 1853

Incertae sedis

Gen. sp indet.

Infrafamily: NATANIA

Subfamily: MOSASAURINAE Gervais, 1853 (sensu Williston, 1897)

Tribe: MOSASAURINI Gervais, 1853 (sensu Russell, 1967)

Mosasaurus sp

Tribe: LEIODONTINI Lingham-Soliar, 1993

Kourisodon sp

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SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: LACERTILIA OWEN, 1842

GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961

MICRORDER: SCINCOMORPHA CAMP, 1923

ORDER: ANGUIMORPHA FURBRINGER, 1900

SUBORDER: DIPLOGLOSSA COPE, 1864

INFRAORDER: THECAGLOSSA COPE, 1890

PARVORDER: MONSTERSAURIA NORELL & GAO, 1977

Superfamily: VARANOIDEA Gray, 1827 (Sensu Camp, 1923)

Family: MOSASAURIDAE Gervais, 1853

Infrafamily: NATANIA

Subfamily: MOSASAURINAE Williston, 1897

Tribe: LEIODONTINI Lingham-Soliar, 1993

Kourisodon sp

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SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: LACERTILIA OWEN, 1842

GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961

MICRORDER: SCINCOMORPHA CAMP, 1923

ORDER: ANGUIMORPHA FURBRINGER, 1900

SUBORDER: DIPLOGLOSSA COPE, 1864

INFRAORDER: THECAGLOSSA COPE, 1890

PARVORDER: MONSTERSAURIA NORELL & GAO, 1977

Superfamily: VARANOIDEA Gray, 1827 (Sensu Camp, 1923)

Family: MOSASAURIDAE Gervais, 1853

Incertae sedis

Gen. sp indet.

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: LACERTILIA OWEN, 1842

GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961

MICRORDER: SCINCOMORPHA CAMP, 1923

ORDER: ANGUIMORPHA FURBRINGER, 1900

SUBORDER: DIPLOGLOSSA COPE, 1864

INFRAORDER: THECAGLOSSA COPE, 1890

PARVORDER: MONSTERSAURIA NORELL & GAO, 1977

Superfamily: VARANOIDEA Gray, 1827 (Sensu Camp, 1923)

Family: MOSASAURIDAE Gervais, 1853

Subfamily: RUSSELLOSAURINAE Bell, 1993

Tribe: PLIOPLATECARPINI Dollo, 1884 (sensu Russell, 1967)

Cf. Platecarpus sp

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: LACERTILIA OWEN, 1842

GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961

MICRORDER: SCINCOMORPHA CAMP, 1923

ORDER: ANGUIMORPHA FURBRINGER, 1900

SUBORDER: DIPLOGLOSSA COPE, 1864

INFRAORDER: THECAGLOSSA COPE, 1890

PARVORDER: MONSTERSAURIA NORELL & GAO, 1977

Superfamily: VARANOIDEA Gray, 1827 (Sensu Camp, 1923)

Family: MOSASAURIDAE Gervais, 1853

Incertae sedis

Gen. sp indet.
Note: Crocodile like teeth.

Infrafamily: NATANIA

Subfamily: MOSASAURINAE Gervais, 1853 (sensu Williston, 1897)

Tribe: MOSASAURINI Gervais, 1853 (sensu Russell, 1967)

Mosasaurus cf. hoffmanni

HONSHU ISLAND

OSAKA PREFECTURE

OSAKA GROUP

PLIOCENE EPOCH

UPPER NEOGENE PERIOD

LOWER QUATERNARY SUBERA-PLEISTOCENE EPOCH

UPPER QUATERNARY SUBERA

UPPER CENOZOIC ERA

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990

COHORT: CROCODYLOTARSI BENTON & CLARK, 1988

MAGNORDER: SUCHIA KREBS, 1974

SUPERORDER: CROCODYLOMORPHA WALKER, 1968

GRANDORDER: CROCODILIFORMES HAY, 1930

MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983

ORDER: EUSUCHIA emend HUXLEY, 1875

INFRAORDER: CROCODYLIA emend GMELIN, 1877

Family: GAVIALIDAE Cuvier, 1807

Subfamily: THORACOSAURINAE Cope, 1875 (sensu Nopcsa, 1928)

*Toyotamphimeia machikanensis
= Tomistoma machikanense

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CENTRAL JAPAN

HONSHU ISLAND

OSAKAPREFECTURE

KISHIWADA CITY

NAGAREKI-CHO

SENPOKU FORMATION

OSAKA GROUP

PLIOCENE EPOCH

UPPER NEOGENE PERIOD

LOWER QUATERNARY SUBERA-PLEISTOCENE EPOCH

UPPER QUATERNARY SUBERA

UPPER CENOZOIC ERA

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990

COHORT: CROCODYLOTARSI BENTON & CLARK, 1988

MAGNORDER: SUCHIA KREBS, 1974

SUPERORDER: CROCODYLOMORPHA WALKER, 1968

GRANDORDER: CROCODILIFORMES HAY, 1930

MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983

ORDER: EUSUCHIA emend HUXLEY, 1875

INFRAORDER: CROCODYLIA emend GMELIN, 1877

Family: GAVIALIDAE Cuvier, 1807

Subfamily: THORACOSAURINAE Cope, 1875 (sensu Nopcsa, 1928)

Toyotamphimeia dcf. machikanensis

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