HAINAULT PROVINCE
A VILLAGE OF HAINAULT
BERNISSART
WEALDEN FORMATION
BERNISSARTIAN
BARREMIAN-LOWER APTIAN
LATE NEOCOMIAN SUBEPOCH-LOWER BARREMIAN STAGE
EARLY GALLIC SUBEPOCH, EARLY CRETACEOUS EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
____________________________________________________________________________________SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGANORDER: CRYPTODIRA GAFFNEY & MEYLAN, 1988
Incertae sedis
Gen. sp indet.
SUPERORDER: SELMACRYPTODIRA GAFFNEY, HUTCHISON, JENKINS & MEEKR, 1987
GRANDORDER: DAIOCRYPTODIRA GAFFNEY & MEYLAN, 1988
MICRORDER: CENTROCRYPTODIRA GAFFNEY & MEYLAN, 1988
ORDER: POLYCRPTODIRA GAFFNEY & MEYLAN, 1988
SUBORDER: PROCOELOCRYPTODIRA GAFFNEY & MEYLAN, 1988
INFRAORDER: CHELOMACRYPTODIRA GAFFNEY & MEYLAN, 1988
PARVORDER: TRIONYCHIDA GRAY, 1870
Superfamily: TRIONYCHIDEA Bell, 1828 (sensu Gray, 1870)
Family: TRIONYCHIDAE Bell, 1828
*Peltochelys duchasteli
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
ORDER: MESOSUCHIA EMEND HUXLEY, 1875
PARVORDER: METASUCHIA BENTON & CLARK, 1988
INFRAORDER: METAMESOSUCHIA HULKE, 1878
NEOSUCHIA CLARK, IN BENTON & CLARK, 1988
Family: BERNISSARTIIDAE Dollo, 1883
*Bernissartia fagesii
Family: GONIOPHOLIDIDAE Cope, 1875
Anteophthalmosuchus hooleyi
Goniopholis simus
BELGIUM
HAINAULT PROVINCE
PERHAPS BERNISSART
NOT KNOWN
WEALDEN
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
ORDER: NEOTETANURAE SERENO, WILSON, LARSSON, DUTHEIL & SUES, 1994
SUBORDER: AVETHEROPODA PAUL, 1988
Superfamily: ALLOSAUROIDEA Marsh, 1878 (sensu Currie & Zhao, 1994)
Family: ALLOSAURIDAE Marsh, 1878
Incertae sedis
Becklespinax altispinax
= Acrocanthosaurus altispinax
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BELGIUM
HAINAULT PROVINCE
BERNISSART
BERNISSARTIAN
HAUTERIVIAN-LOWER APTIAN
LATE NEOCOMIAN SUBEPOCH-LOWER BARREMIAN STAGE
EARLY GALLIC SUBEPOCH, EARLY CRETACEOUS EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
SUBORDER: ANKYLOPOLLEXIA SERENO, 1986
PARVORDER: HADROSAURIFORMES SERENO, 1997
Superfamily: IGUANODONTIDEA Cope, 1869 (sensu Sereno, 1986)
Family: IGUANODONTIDAE Cope, 1869
*Dollodon bampingi
= Iguanodon? atherfieldensisIguanodon anglicus
= Iguanodon anglicum
= Iguanodon mantelli
= Therosaurus mantelli
= Iguanodon mantelli
*Iguanodon? bernissartensis
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BELGIUM
HAINAULT PROVINCE
MONS BASIN
BOIS DE BAUDOUR CLAY QUARRIES
BERNISSARTIAN
HAUTERIVIAN STAGE
UPPER NEOCOMIAN SUBEPOCH-LOWER APTIAN STAGE
MIDDLE GALLIC SUBEPOCH
EARLY CRETACEOUS EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
SUPERORDER: SAUROPODOMORPHA HUENE, 1932
GRANDORDER: BRONTOSAURIA OLSHEVSKY, 1991 (emend)
MICRORDER: SAUROPODA MARSH, 1878
Incetae sedis
Gen. sp indet.
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
SUBORDER: ANKYLOPOLLEXIA SERENO, 1986
PARVORDER: HADROSAURIFORMES SERENO, 1997
Superfamily: IGUANODONTIDEA Cope, 1869 (sensu Sereno, 1986)
Family: IGUANODONTIDAE Cope, 1869
Iguanodon bernissartensis
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BELGIUM
HAINAULT PROVINCE
BERNISSART
COAL TIPS (‘TERRIL”)
BERNISSARTIAN
HAUTERIVIAN-LOWER APTIAN
LATE NEOCOMIAN SUBEPOCH-LOWER BARREMIAN STAGE
EARLY GALLIC SUBEPOCH, EARLY CRETACEOUS EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
SUBORDER: ANKYLOPOLLEXIA SERENO, 1986
PARVORDER: HADROSAURIFORMES SERENO, 1997
Superfamily: IGUANODONTIDEA Cope, 1869 (sensu Sereno, 1986)
Family: IGUANODONTIDAE Cope, 1869
Iguanodon? bernissartensis
Mantellisaurus atherfieldensis
= Iguanodon? atherfieldensis
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BELGIUM
HAINAULT PROVINCE
BETWEEN MONS AND TOURNAI
BERNISSART
SAINTE-BARBE CLAYS FORMATION
BERNISSARTIAN
LOWER APTIAN
LATE NEOCOMIAN SUBEPOCH-LOWER BARREMIAN STAGE
EARLY GALLIC SUBEPOCH, EARLY CRETACEOUS EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
____________________________________________________________________________________SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
GRANDORDER: DAIOCRYPTODIRA GAFFNEY & MEYLAN, 1988
MICRORDER: EUCRYPTODIRA GAFFNEY, 1975
Incertae sedis
*Chitracephalus dumonii
BELGIUM
HAINAULT PROVINCE
CRAIE D’OBOURG
CRAIE DE NOUVELLES
UPPER CAMPANIAN STAGE
MIDDLE SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: LACERTILIA OWEN, 1842
GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961
MICRORDER: SCINCOMORPHA CAMP, 1923
ORDER: ANGUIMORPHA FURBRINGER, 1900
SUBORDER: DIPLOGLOSSA COPE, 1864
INFRAORDER: THECAGLOSSA COPE, 1890
PARVORDER: MONSTERSAURIA NORELL & GAO, 1977
Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)
Family: MOSASAURIDAE Gervais, 1853
Infrafamily: NATANIA
Subfamily: MOSASAURINAE Gervais, 1853 (sensu Williston, 1897)
Tribe: GLOBIDENTINI Dollo, 1924 (sensu Russell, 1967)
Globidens alabamaensis
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BELGIUM
HAINAULT PROVINCE
MONS BASIN
CIPLY AREA
SPIENNES CHALK
UPPER CAMPANIAN STAGE
MIDDLE SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: EURYAPSIDA COLBERT, 1945
SUPERORDER: SAUROPTERYGIA OWEN 1860
GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994
ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
INFRAORDER: PLESIOSAUROMORPHA NOVA
EUPLESIOSAURIA O’KEEFE, 2001
Family: PLESIOSAURIDAE Gray, 1825
Incertae sedis
non Plesiosaurus, *P. houzeaui
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BELGIUM
HAINAULT PROVINCE
HARMIGNIES
SENONIAN
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904 OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: LACERTILIA OWEN, 1842
GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961
MICRORDER: SCINCOMORPHA CAMP, 1923
ORDER: ANGUIMORPHA FURBRINGER, 1900
SUBORDER: DIPLOGLOSSA COPE, 1864
INFRAORDER: THECAGLOSSA COPE, 1890
PARVORDER: MONSTERSAURIA NORELL & GAO, 1977
Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)
Family: MOSASAURIDAE Gervais, 1853
Infrafamily: NATANIA
Subfamily: MOSASAURINAE Gervais, 1853 (sensu Williston, 1897)
Tribe: GLOBIDENTINI Dollo, 1924 (sensu Russell, 1967)
Globidens alabamaensis
Globidens dakotensis
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BELGIUM
HAINAULT PROVINCE
CIPLY
BROWN PHOSPHATIC CHALK
LATE MAASTRICHTIAN STAGE
UPPERMOST SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: LACERTILIA OWEN, 1842
GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961
MICRORDER: SCINCOMORPHA CAMP, 1923
ORDER: ANGUIMORPHA FURBRINGER, 1900
SUBORDER: DIPLOGLOSSA COPE, 1864
INFRAORDER: THECAGLOSSA COPE, 1890
PARVORDER: MONSTERSAURIA NORELL & GAO, 1977
Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)
Family: MOSASAURIDAE Gervais, 1853
Infrafamily: NATANIA
Subfamily: RUSSELLOSAURINAE Bell, 1993
Tribe: PLIOPLATECARPINI Dollo, 1884 (sensu Russell, 1967)
*Plioplatecarpus houzeaui
= Oterognathus houzeaui
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