EUROPEAN RUSSIA
ARKHANGELSK REGION
MEZENSKII DISTRICT
GLYADNAYA SHCHEL’YA LOCALITY
KRASNOSHCHEL’YA FORMATION
URZHUMIAN HORIZON
MEZEN FAUNAL ASSEMBLAGE
LOWER TATARIAN AGE
CAPITANIAN STAGE
MIDDLE ZECHSTEIN EPOCH
LATE MIDDLE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
NEOTETRAPODA
SUBCLASS: REPTILIOMORPHA SAVE-SODERBERGH, 1934
ORDER: SEYMOURIAMORPHA WATSON, 1917
SUBORDER: NYCTEROLETERIDAE TATARINOV, 1972
Family: NYCTEROLETERIDAE Romer, 1956
Subfamily: NYCTEROLECTERINAE Romer, 1956
Macroleter poezicus
Subfamily: BASHKYROLETERINAE Ivachnenko, 1997
*Bashkyroleter mesensis
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: PARAREPTILIA OLSON 1947
INFRACLASS: SAUROPSIDA HUXLEY, 1864
ORDER: PROCOLOPHONOMORPHA ROMER, 1964
SUBORDER: PROCOLPHOMORPHA SEELEY, 1888
Family: NYCTIPHRURETIDAE Efremov, 1938
*Nyctiphruretus acudens
SUBCLASS: DIAPSIDA OSBORN, 1904
Incertae sedis
*Lanthanolania ivakhenennkoi
CLADE NAME: NEOVARANOPSIA SPINDLER, WERNEBURG, SCHNEIDER, LUTHARDT, ANNACKER & RÖßLER, 2019
Family: VARANOPSEIDAE Romer & Price, 1940
Subfamily: MESENOSAURINAE Romer 1956
Mesenosaurus romeri
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPIDA BROOM, 1905
ORDER: EOTHERIODONTIA OLSON, 1962
Incertae sedis
Family: NIKKASAURIDAE Ivakhnenko, 2000
*Nikkasaurus tatarinovi
*Reiszia gubini
INFRAORDER: EOTITANOSUCHIA BOONSTRA, 1963
Family: NIIAFTASUCHIDAE Ivakhenko, 1990
Niaftasuchus zekkeli
Family: ALRAUSUCHIDAE Ivakhenko, 2008
Alrausuchus tagax
_____________________________________________________________________________________
EUROPEAN RUSSIA
ARKHANGELSK REGION
MEZENSKII DISTRICT
DOROGAYA GORA LOCALITY
BIARMIAN SERIES
URZHUMIAN STAGE
URZHUMIAN HORIZON
MEZEN FAUNAL ASSEMBLAGE
LOWER TATARIAN AGE
CAPITANIAN STAGE
MIDDLE ZECHSTEIN EPOCH
LATE MIDDLE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
NEOTETRAPODA
SUBCLASS: REPTILIOMORPHA SAVE-SODERBERGH, 1934
ORDER: SEYMOURIAMORPHA WATSON, 1917
SUBORDER: NYCTEROLETERIDAE TATARINOV, 1972
Family: NYCTEROLETERIDAE Romer, 1956
Subfamily: NYCTEROLECTERINAE Romer, 1956
Macroleter poezicus
Subfamily: BASHKYROLETERINAE Ivachnenko, 1997
Bashkyroleter mesensis
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: PARAREPTILIA OLSON 1947
INFRACLASS: SAUROPSIDA HUXLEY, 1864
ORDER: PROCOLOPHONOMORPHA ROMER, 1964
SUBORDER: PROCOLPHOMORPHA SEELEY, 1888
Family: NYCTIPHRURETIDAE Efremov, 1938
Nyctiphruretus acudens
SUBCLASS: DIAPSIDA OSBORN, 1904
CLADE NAME: NEOVARANOPSIA SPINDLER, WERNEBURG, SCHNEIDER, LUTHARDT, ANNACKER & RÖßLER, 2019
Family: VARANOPSEIDAE Romer & Price, 1940
Subfamily: MESENOSAURINAE Romer 1956
Mesenosaurus romeri
SUBCLASS: SYNAPSIDA OSBORN, 1903
INFRACLASS DINOMORPHA IVAKNENKO, 2008
SUPERORDER: GORGONTIA IVAKNENKO, 2008
ORDER: EOTHERIODONTIA OLSON, 1962
INFRAORDER: EOTITANOSUCHIA BOONSTRA, 1963
Family: NIIAFTASUCHIDAE Ivakhenko, 1990
Niaftasuchus zekkeli
_____________________________________________________________________________________
EUROPEAN RUSSIA
ARKHANGELSK REGION
MEZENSKII DISTRICT
KOSMOGORODSKO'E LOCALITY
BIARMIAN SERIES
URZHUMIAN STAGE
URZHUMIAN HORIZON
MEZEN FAUNAL ASSEMBLAGE
LOWER TATARIAN AGE
CAPITANIAN STAGE
MIDDLE ZECHSTEIN EPOCH
LATE MIDDLE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
NEOTETRAPODA
SUBCLASS: REPTILIOMORPHA SAVE-SODERBERGH, 1934
ORDER: SEYMOURIAMORPHA WATSON, 1917
SUBORDER: NYCTEROLETERIDAE TATARINOV, 1972
Family: NYCTEROLETERIDAE Romer, 1956
Subfamily: NYCTEROLECTERINAE Romer, 1956
Macroleter poezicus
Subfamily: BASHKYROLETERINAE Ivachnenko, 1997
Bashkyroleter mesensis
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: PARAREPTILIA OLSON 1947
INFRACLASS: SAUROPSIDA HUXLEY, 1864
ORDER: PROCOLOPHONOMORPHA ROMER, 1964
SUBORDER: PROCOLPHOMORPHA SEELEY, 1888
Family: NYCTIPHRURETIDAE Efremov, 1938
Nyctiphruretus acudens
SUBCLASS: DIAPSIDA OSBORN, 1904
CLADE NAME: NEOVARANOPSIA SPINDLER, WERNEBURG, SCHNEIDER, LUTHARDT, ANNACKER & RÖßLER, 2019
Family: VARANOPSEIDAE Romer & Price, 1940
Subfamily: MESENOSAURINAE Romer 1956
Mesenosaurus romeri
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPIDA BROOM, 1905
ORDER: EOTHERIODONTIA OLSON, 1962
INFRAORDER: EOTITANOSUCHIA BOONSTRA, 1963
Family: ALRAUSUCHIDAE Ivakhenko, 2008
Alrausuchus tagax
_____________________________________________________________________________________
EUROPEAN RUSSIA
ARKHANGEL’SK PROVINCE
NEAR CHORVA
BASKUNCHAK SERIES
EARLY TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LABYRINTHODONTIA OWEN, 1860
SUPERORDER: TEMNOSPONDYLIA ZITTEL, 1887
MICROORDER: LIMNARCHIA YATES & WARREN, 2000
ORDER: PLAGIOSAURIA NILSSON, 1939 EMEND PANCHEN, 1959
Superfamily: PLAGIOSAUROIDEA Jaekel, 1913 (sensu Kuhn, 1965)
Family: PLAGIOSAURIDAE Jaekel, 1913
Subfamily: PLAGIOSTERNINAE Shishkin, 1986
*Melanopelta antiqua
_____________________________________________________________________________________
EUROPEAN RUSSIA
ARKHANGEL'S SK REGION
NENETS NATIONAL RESERVATION
5 KM DOWNSTREAM FROM MOUTH OF LESTANSHOR CREEK
RIGHT BANK OF KHEY-YAGA RIVER
LOWERMOST PART OF NYADEYTINSKAYA FORMATION
ANISIAN STAGE
LOWER MID TRIASSIC EPOCH
MIDDLE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LABYRINTHODONTIA OWEN, 1860
SUPERORDER: TEMNOSPONDYLIA ZITTEL, 1887
MICROORDER: LIMNARCHIA YATES & WARREN, 2000
ORDER: CAPITOSAURIA ROMER, 1947
Superfamily: CAPITOSAUROIDEA Watson, 1919 (sensu Romer, 1947)
Incertae sedis
*Komatosuchus chalyshevi
_____________________________________________________________________________________
EUROPEAN RUSSIA
ARKHANGEL’SK REGION
6 KM FROM THE MOUTH OF THE LEFT BANK OF VYBOR RIVER
PIZHMA MEZENSKAYA FORMATION
EARLY TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LABYRINTHODONTIA OWEN, 1860
SUPERORDER: TEMNOSPONDYLIA ZITTEL, 1887
MICROORDER: LIMNARCHIA YATES & WARREN, 2000
ORDER: TREMATOSAURIA PIVETEAU, 1955
Superfamily: TREMATOSAUROIDEA Save-Soderbergh, 1935
Family: BENTHOSUCHIDAE Efremov, 1931
Subfamily: BENTHOSUCHINAE Efremov, 1931 (sensu Efremov, 1937)
*Vyborosaurus mirus
_____________________________________________________________________________________
EUROPEAN RUSSIA
ARKHANGEL’SK REGION
6 KM FROM THE MOUTH OF THE LEFT BANK OF VYBOR RIVER
NI’MA-III LOCALITY
PIZHMAN MEZENSKAYA FORMATION
EARLY TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LABYRINTHODONTIA OWEN, 1860
SUPERORDER: TEMNOSPONDYLIA ZITTEL, 1887
MICROORDER: LIMNARCHIA YATES & WARREN, 2000
ORDER: TREMATOSAURIA PIVETEAU, 1955
Superfamily: TREMATOSAUROIDEA Save-Soderbergh, 1935
Family: BENTHOSUCHIDAE Efremov, 1931
Subfamily: BENTHOSUCHINAE Efremov, 1931 (sensu Efremov, 1937)
Vyborosaurus mirus
_____________________________________________________________________________________
EUROPEAN RUSSIA
ARKHANGEL’SK REGION
6 KM FROM THE MOUTH OF THE LEFT BANK OF VYBOR RIVER
NIZHNYAYA SYAMZHEN’ GA-III LOCALITY
PIZHMAN MEZENSKAYA FORMATION
EARLY TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LABYRINTHODONTIA OWEN, 1860
SUPERORDER: TEMNOSPONDYLIA ZITTEL, 1887
MICROORDER: LIMNARCHIA YATES & WARREN, 2000
ORDER: TREMATOSAURIA PIVETEAU, 1955
Superfamily: TREMATOSAUROIDEA Save-Soderbergh, 1935
Family: BENTHOSUCHIDAE Efremov, 1931
Subfamily: BENTHOSUCHINAE Efremov, 1931 (sensu Efremov, 1937)
Vyborosaurus mirus
_____________________________________________________________________________________
EUROPEAN RUSSIA
ARKHANGEL’SK REGION
6 KM FROM THE MOUTH OF THE LEFT BANK OF VYBOR RIVER
PIZHMA’L LOCALITY
PIZHMAN MEZENSKAYA FORMATION
EARLY TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LABYRINTHODONTIA OWEN, 1860
SUPERORDER: TEMNOSPONDYLIA ZITTEL, 1887
MICROORDER: LIMNARCHIA YATES & WARREN, 2000
ORDER: TREMATOSAURIA PIVETEAU, 1955
Superfamily: TREMATOSAUROIDEA Save-Soderbergh, 1935
Family: BENTHOSUCHIDAE Efremov, 1931
Subfamily: BENTHOSUCHINAE Efremov, 1931 (sensu Efremov, 1937)
Vyborosaurus mirus
_____________________________________________________________________________________
EUROPEAN RUSSIA
ARKHANGEL’SK PROVINCE
PECHORA BASIN
HEI YAG RIVER
TIMON-URAL
LOWER LESTANSHORSKAYA SVITA
YARENSKIAN GORIZONT
EARLY TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LABYRINTHODONTIA OWEN, 1860
SUPERORDER: TEMNOSPONDYLIA ZITTEL, 1887
MICROORDER: LIMNARCHIA YATES & WARREN, 2000
ORDER: BRACHYOPOMORPHA WARREN & MARSICANO, 2000
SUBORDER: BRACHYOPOIDEA ROMER, 1947
Superfamily: BRACHYOPOIDEA Save-Soderbergh, 1935
Family: BRACHYOPIDAE Lydekker, 1885
Incertae sedis
*Batrachosuchoides impressus
= Batrachosuchus impressus
_____________________________________________________________________________________
EUROPEAN RUSSIA
ARKHANGEL’SK REGION
PEZA RIVER
LOWER TATARIAN AGE (?)
CAPITANIAN STAGE
MIDDLE ZECHSTEIN EPOCH
LATE MIDDLE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPIDA BROOM, 1905
ORDER: EOTHERIODONTIA OLSON, 1962
SUBORDER: EOTITANOSUCHIA BOONSTRA, 1963
INFRAORDER: BIARMOSUCHIA SIGOGNEAU-RUSSELL, 1989
Superfamily: RHOPALODONTIDEA Seeley, 1894
Family: RHOPALODONTIDAE Seeley, 1894)
?Rhopalodon sp
_____________________________________________________________________________________
EUROPEAN RUSSIA
ARKHANGEL’SK REGION
6 KM FROM THE MOUTH OF THE LEFT BANK OF VYBOR RIVER
TSIL’MA-I LOCALITY
PIZHMAN MEZENSKAYA FORMATION
EARLY TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LABYRINTHODONTIA OWEN, 1860
SUPERORDER: TEMNOSPONDYLIA ZITTEL, 1887
MICROORDER: LIMNARCHIA YATES & WARREN, 2000
ORDER: TREMATOSAURIA PIVETEAU, 1955
Superfamily: TREMATOSAUROIDEA Save-Soderbergh, 1935
Family: BENTHOSUCHIDAE Efremov, 1931
Subfamily: BENTHOSUCHINAE Efremov, 1931 (sensu Efremov, 1937)
Vyborosaurus mirus
_____________________________________________________________________________________
EUROPEAN RUSSIA
ARKHANGEL’SK PROVINCE
PECHORA RIVER BASIN
TSIL'MA RIVER
TSIL'MA I LOCALITY
CHARKABOZHSKAYA FORMATION
SLUDKA HORIZON
VETLUGA SUPERHORIZON
USTMYLIAN GORIZONT
EARLY OLENEKIAN
UPPER NAMMALIAN STAGE
MIDDLE SCYTHIAN EPOCH
EARLY TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
_____________________________________________________________________________________SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
MAGNORDER: PROTEROSUCHIFORMES nova
ORDER: PROTEROSUCHIA BROOM, 1906
Family: PROTEROSUCHIDAE Huene, 1914
Tsylmosuchus jakovlevi
EUROPEAN RUSSIA
ARKHANGEL’SK PROVINCE
PECHORA RIVER BASIN
TSIL'MA RIVER
TSIL'MA II LOCALITY
CHARKABOZHSKAYA FORMATION
SLUDKA HORIZON
VETLUGA SUPERHORIZON
USTMYLIAN GORIZONT
EARLY OLENEKIAN
UPPER NAMMALIAN STAGE
MIDDLE SCYTHIAN EPOCH
EARLY TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
_____________________________________________________________________________________SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
MAGNORDER: PROTEROSUCHIFORMES nova
ORDER: PROTEROSUCHIA BROOM, 1906
Family: PROTEROSUCHIDAE Huene, 1914
Tsylmosuchus jakovlevi
EUROPEAN RUSSIA
ARKHANGEL’SK PROVINCE
PECHORA RIVER BASIN
TSIL'MA RIVER
TSIL'MA V LOCALITY
CHARKABOZHSKAYA FORMATION
SLUDKA HORIZON
VETLUGA SUPERHORIZON
USTMYLIAN GORIZONT
EARLY OLENEKIAN
UPPER NAMMALIAN STAGE
MIDDLE SCYTHIAN EPOCH
EARLY TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
_____________________________________________________________________________________SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
MAGNORDER: PROTEROSUCHIFORMES nova
ORDER: PROTEROSUCHIA BROOM, 1906
Family: PROTEROSUCHIDAE Huene, 1914
Tsylmosuchus jakovlevi
EUROPEAN RUSSIA
ARKHANGELSKAYA
RIGHT BANK OF THE YARENGA R
EARLY TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LABYRINTHODONTIA OWEN, 1860
SUPERORDER: TEMNOSPONDYLIA ZITTEL, 1887
MICROORDER: LIMNARCHIA YATES & WARREN, 2000
ORDER: TREMATOSAURIA PIVETEAU, 1955
Superfamily: TREMATOSAUROIDEA Save-Soderbergh, 1935
Family: BENTHOSUCHIDAE Efremov, 1931
Subfamily: BENTHOSUCHINAE Efremov, 1931 (sensu Efremov, 1937)
*Yarengia perplexa
_____________________________________________________________________________________
EUROPEAN RUSSIA
ARKHANGEL’SK REGION
LOWER MEZEN RIVER
ZONE III
LOWER TATARIAN AGE
LOPINGIAN SUBEPOCH
UPPER ZECHSTEIN EPOCH
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
NEOTETRAPODA
SUBCLASS: REPTILIOMORPHA SAVE-SODERBERGH, 1934
ORDER: SEYMOURIAMORPHA WATSON, 1917
SUBORDER: NYCTEROLETERIDAE TATARINOV, 1972
Family: NYCTEROLETERIDAE Romer, 1956
Subfamily: NYCTEROLECTERINAE Romer, 1956
*Nycteroleter ineptus
_____________________________________________________________________________________
EUROPEAN RUSSIA
ARKHANGELSK REGION
MEZENSKII DISTRICT
UST’-PEZA LOCALITY
BIARMIN SERIES
URZHUMINA STAGE
URZHUMIAN HORIZON
UPPER KIMZHA BEDS
MEZEN FAUNAL ASSEMBLAGE
LOWER TATARIAN AGE
CAPITANIAN STAGE
MIDDLE ZECHSTEIN EPOCH
LATE MIDDLE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
NEOTETRAPODA
SUBCLASS: REPTILIOMORPHA SAVE-SODERBERGH, 1934
ORDER: SEYMOURIAMORPHA WATSON, 1917
SUBORDER: NYCTEROLETERIDAE TATARINOV, 1972
Family: NYCTEROLETERIDAE Romer, 1956
Subfamily: NYCTEROLECTERINAE Romer, 1956
*Macroleter poezicus
Subfamily: BASHKYROLETERINAE Ivachnenko, 1997
Bashkyroleter mesensis
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: PARAREPTILIA OLSON 1947
INFRACLASS: SAUROPSIDA HUXLEY, 1864
ORDER: PROCOLOPHONOMORPHA ROMER, 1964
SUBORDER: PROCOLPHOMORPHA SEELEY, 1888
Family: NYCTIPHRURETIDAE Efremov, 1938
*Nyctiphruretus acudens
SUBCLASS: DIAPSIDA OSBORN, 1904
CLADE NAME: NEOVARANOPSIA SPINDLER, WERNEBURG, SCHNEIDER, LUTHARDT, ANNACKER & RÖßLER, 2019
Family: VARANOPSEIDAE Romer & Price, 1940
Subfamily: MESENOSAURINAE Romer 1956
Mesenosaurus romeri
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPIDA BROOM, 1905
ORDER: EOTHERIODONTIA OLSON, 1962
INFRAORDER: EOTITANOSUCHIA BOONSTRA, 1963
Family: ALRAUSUCHIDAE Ivakhenko, 2008
Alrausuchus tagax
= Biarmosuchus tagax
_____________________________________________________________________________________
EUROPEAN RUSSIA
ARKHANGELSK REGION
MEZENSKII DISTRICT
BELOKUR’E LOCALITY
KRASNOSHCHEL’YA FORMATION
URZHUMIAN STAGE
URZHUMIAN HORIZON
MEZEN FAUNAL ASSEMBLAGE
LOWER TATARIAN AGE
CAPITANIAN STAGE
MIDDLE ZECHSTEIN EPOCH
LATE MIDDLE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
NEOTETRAPODA
SUBCLASS: REPTILIOMORPHA SAVE-SODERBERGH, 1934
ORDER: SEYMOURIAMORPHA WATSON, 1917
SUBORDER: NYCTEROLETERIDAE TATARINOV, 1972
Family: NYCTEROLETERIDAE Romer, 1956
Subfamily: NYCTEROLECTERINAE Romer, 1956
Macroleter poezicus
Subfamily: BASHKYROLETERINAE Ivachnenko, 1997
Bashkyroleter mesensis
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: PARAREPTILIA OLSON 1947
INFRACLASS: SAUROPSIDA HUXLEY, 1864
ORDER: PROCOLOPHONOMORPHA ROMER, 1964SUBORDER: PROCOLPHOMORPHA SEELEY, 1888
Family: NYCTIPHRURETIDAE Efremov, 1938
Nyctiphruretus acudens
SUBCLASS: DIAPSIDA OSBORN, 1904
CLADE NAME: NEOVARANOPSIA SPINDLER, WERNEBURG, SCHNEIDER, LUTHARDT, ANNACKER & RÖßLER, 2019
Family: VARANOPSEIDAE Romer & Price, 1940
Subfamily: MESENOSAURINAE Romer 1956
Mesenosaurus romeri
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPIDA BROOM, 1905
ORDER: EOTHERIODONTIA OLSON, 1962
Incertae sedis
Family: NIKKASAURIDAE Ivakhnenko, 2000
Nikkasaurus tatarinovi
_____________________________________________________________________________________
EUROPEAN RUSSIA
ARKHAGELSK REGION
MEZEN DISTRICT
UST-PYOZA
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPIDA BROOM, 1905
ORDER: EOTHERIODONTIA OLSON, 1962
SUBORDER: EOTITANOSUCHIA BOONSTRA, 1963
INFRAORDER: BIARMOSUCHIA SIGOGNEAU-RUSSELL, 1989
Family: BIARMOSUCHIDAE Olsen, 1962
Biarmosuchus tagax
_____________________________________________________________________________________
EUROPEAN RUSSIA
ARKHANGELSK REGION
MENZEN DISTRICT
UST-NYAFTA
BIARMIAN SERIES
UPPER KIMZHA BEDS
URZHUMIAN STAGE
UPPER URZHUMIAN HORIZON
MEZEN FAUNAL ASSEMBLAGE
LOWER TATARIAN AGE
CAPITANIAN STAGE
UPPER GUADELUPIAN STAGE
MIDDLE ZECHSTEIN EPOCH
LATE MIDDLE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
NEOTETRAPODA
SUBCLASS: REPTILIOMORPHA SAVE-SODERBERGH, 1934
ORDER: SEYMOURIAMORPHA WATSON, 1917
SUBORDER: NYCTEROLETERIDAE TATARINOV, 1972
Family: NYCTEROLETERIDAE Romer, 1956
Subfamily: NYCTEROLECTERINAE Romer, 1956
Macroleter poezicus
Subfamily: BASHKYROLETERINAE Ivachnenko, 1997
Bashkyroleter mesensis
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: PARAREPTILIA OLSON 1947
INFRACLASS: SAUROPSIDA HUXLEY, 1864
ORDER: PROCOLOPHONOMORPHA ROMER, 1964
SUBORDER: PROCOLPHOMORPHA SEELEY, 1888
Family: NYCTIPHRURETIDAE Efremov, 1938
Nyctiphruretus acudens
SUBCLASS: DIAPSIDA OSBORN, 1904
CLADE NAME: NEOVARANOPSIA SPINDLER, WERNEBURG, SCHNEIDER, LUTHARDT, ANNACKER & RÖßLER, 2019
Family: VARANOPSEIDAE Romer & Price, 1940
Subfamily: MESENOSAURINAE Romer 1956
Mesenosaurus romeri
SUBCLASS: SYNAPSIDA OSBORN, 1903
ORDER: PELYCOSAURIA COPE, 1878
SUBORDER: EUPELYCOSAURIA KEMP, 1982
INFRAORDER: OPHIACHODONTIA NOVA
Family: VARANOPIDAE emend Romer & Price, 1940, REISZ & DILKES, 2003
*Pyozia mesenensis
INFRACLASS DINOMORPHA IVAKNENKO, 2008
SUPERORDER: GORGONTIA IVAKNENKO, 2008
ORDER: EOTHERIODONTIA OLSON, 1962
INFRAORDER: EOTITANOSUCHIA BOONSTRA, 1963
Family: NIIAFTASUCHIDAE Ivakhenko, 1990
*Niaftasuchus zekkeli
_____________________________________________________________________________________
EUROPEAN RUSSIA
ARKHANGEL'SK REGION
MEZEN DISTRICT
KIMJA RIVER
ZONE 111
RED MARLS HORIZON
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: PARAREPTILIA OLSON 1947
INFRACLASS: SAUROPSIDA HUXLEY, 1864
ORDER: PROCOLOPHONOMORPHA ROMER, 1964
SUBORDER: PROCOLPHOMORPHA SEELEY, 1888
Family: NYCTIPHRURETIDAE Efremov, 1938
*Nyctiphruretus acudens
_____________________________________________________________________________________
EUROPEAN RUSSIA
ARKHANGELSK REGION
LESHUKONSKII DISTRICT
ON THE MEZEN RIVER
NEAR THE JUCTION WITH THE VASHKA RIVER
NYISAGORA LOCALITY
BIARMIAN SERIES
URZHUMIAN STAGE
URZHUMIAN HORIZON
ZONE II
MEZEN FAUNAL ASSEMBLAGE
EARLIEST EARLY TATARIAN AGE
GUADELUPIAN STAGE
MIDDLE ZEICHSTEIN EPOCH
MIDDLE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
NEOTETRAPODA
SUBCLASS: REPTILIOMORPHA SAVE-SODERBERGH, 1934
ORDER: SEYMOURIAMORPHA WATSON, 1917
SUBORDER: NYCTEROLETERIDAE TATARINOV, 1972
Family: NYCTEROLETERIDAE Romer, 1956
Subfamily: NYCTEROLECTERINAE Romer, 1956
Macroleter poezicus
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: PARAREPTILIA OLSON 1947
INFRACLASS: SAUROPSIDA HUXLEY, 1864
ORDER: LANTHANOSUCHIDAE TATARINOV, 1971
Family: LANTHANOSUCHIDAE Efremov, 1946
Lanthaniscus efremovi
ORDER: PROCOLOPHONOMORPHA ROMER, 1964
SUBORDER: PROCOLPHOMORPHA SEELEY, 1888
Family: NYCTIPHRURETIDAE Efremov, 1938
Nyctiphruretus acudens
SUBCLASS: DIAPSIDA OSBORN, 1904
CLADE NAME: NEOVARANOPSIA SPINDLER, WERNEBURG, SCHNEIDER, LUTHARDT, ANNACKER & RÖßLER, 2019
Family: VARANOPSEIDAE Romer & Price, 1940
Subfamily: MESENOSAURINAE Romer 1956
Mesenosaurus romeri
SUBCLASS: SYNAPSIDA OSBORN, 1903
ORDER: PELYCOSAURIA COPE, 1878
SUBORDER: CASEASAURIA WILLISTON, 1912
Family: CASEIDAE Williston, 1912
Ennatosaurus tecton
INFRACLASS DINOMORPHA IVAKNENKO, 2008
SUPERORDER: GORGONTIA IVAKNENKO, 2008
ORDER: DEINOCEPHALIA OLSEN, 1962
SUBORDER: EUDEINOCEPHALIA APPLEBY, CHARIG, COX, KERMACK & TARLO, 1967
INFRAORDER: EOTITANOSUCHIA BOONSTRA, 1963
Family: NIIAFTASUCHIDAE Ivakhenko, 1990
Niaftasuchus zekkeli
_____________________________________________________________________________________
EUROPEAN RUSSIA
ARKHANGELSK REGION
LESHUKONSKII DISTRICT
UST-VASHKA LOCALITY
KRASNOSHCHEL’YA FORMATION
BIARMIAN SERIES
URZHUMIAN STAGE
URZHUMIAN HORIZON
MEZEN FAUNAL ASSEMBLAGE
LOWER TATARIAN AGE
CAPITANIAN STAGE
MIDDLE ZECHSTEIN EPOCH
LATE MIDDLE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
NEOTETRAPODA
SUBCLASS: REPTILIOMORPHA SAVE-SODERBERGH, 1934
ORDER: SEYMOURIAMORPHA WATSON, 1917
SUBORDER: NYCTEROLETERIDAE TATARINOV, 1972
Family: NYCTEROLETERIDAE Romer, 1956
Subfamily: BASHKYROLETERINAE Ivachnenko, 1997
Bashkyroleter mesensis
Subfamily: NYCTEROLECTERINAE Romer, 1956
Macroleter poezicus
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: PARAREPTILIA OLSON 1947
INFRACLASS: SAUROPSIDA HUXLEY, 1864
ORDER: PROCOLOPHONOMORPHA ROMER, 1964
SUBORDER: PROCOLPHOMORPHA SEELEY, 1888
Family: NYCTIPHRURETIDAE Efremov, 1938
Nyctiphruretus acudens
SUBCLASS: DIAPSIDA OSBORN, 1904
CLADE NAME: NEOVARANOPSIA SPINDLER, WERNEBURG, SCHNEIDER, LUTHARDT, ANNACKER & RÖßLER, 2019
Family: VARANOPSEIDAE Romer & Price, 1940
Subfamily: MESENOSAURINAE Romer 1956
Mesenosaurus romeri
SUBCLASS: SYNAPSIDA OSBORN, 1903
INFRACLASS DINOMORPHA IVAKNENKO, 2008
SUPERORDER: NIKKASAURIA IVAKHNENKO, 2008
ORDER: NIKKASAURIDA IVAKHNENNKO, 2008
Family: NIKKASAURIDAE Ivakhnenko, 2000
Nikkasaurus tatarinovi
*Reiszia tippula
SUPERORDER: GORGONTIA IVAKNENKO, 2008
ORDER: DEINOCEPHALIA OLSEN, 1962
SUBORDER: EUDEINOCEPHALIA APPLEBY, CHARIG, COX, KERMACK & TARLO, 1967
INFRAORDER: EOTITANOSUCHIA BOONSTRA, 1963
Family: NIIAFTASUCHIDAE Ivakhenko, 1990
Niaftasuchus zekkeli
Family: ALRAUSUCHIDAE Ivakhenko, 2008
Alrausuchus tagax
_____________________________________________________________________________________
EUROPEAN RUSSIA
ARKHANGELSK REGION
PEZA-1
BIARMIAN SERIES
UPPER KIMZHA BEDS
URZHUMIAN STAGE
URZHUMIAN HORIZON
MEZEN FAUNAL ASSEMBLAGE
LOWER TATARIAN AGE
LOPINGIAN SUBEPOCH
UPPER ZECHSTEIN EPOCH
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
NEOTETRAPODA
SUBCLASS: REPTILIOMORPHA SAVE-SODERBERGH, 1934
ORDER: SEYMOURIAMORPHA WATSON, 1917
SUBORDER: NYCTEROLETERIDAE TATARINOV, 1972
Family: NYCTEROLETERIDAE Romer, 1956
Subfamily: NYCTEROLECTERINAE Romer, 1956
Macroleter poezicus
Nycteroleter ineptus
Subfamily: BASHKYROLETERINAE Ivachnenko, 1997
Bashkyroleter mesensis
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: PARAREPTILIA OLSON 1947
INFRACLASS: SAUROPSIDA HUXLEY, 1864
ORDER: LANTHANOSUCHIDAE TATARINOV, 1971
Family: LANTHANOSUCHIDAE Efremov, 1946
Lanthaniscus efremovi
ORDER: PROCOLOPHONOMORPHA ROMER, 1964
SUBORDER: PROCOLPHOMORPHA SEELEY, 1888
Family: NYCTIPHRURETIDAE Efremov, 1938
Nyctiphruretus acudens
SUBCLASS: DIAPSIDA OSBORN, 1904
CLADE NAME: NEOVARANOPSIA SPINDLER, WERNEBURG, SCHNEIDER, LUTHARDT, ANNACKER & RÖßLER, 2019
Family: VARANOPSEIDAE Romer & Price, 1940
Subfamily: MESENOSAURINAE Romer 1956
Mesenosaurus romeri
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPIDA BROOM, 1905
ORDER: EOTHERIODONTIA OLSON, 1962
INFRAORDER: EOTITANOSUCHIA BOONSTRA, 1963
Family: NIIAFTASUCHIDAE Ivakhenko, 1990
Niaftasuchus zekkeli
Family: ALRAUSUCHIDAE Ivakhenko, 2008
*Alrausuchus tagax
= Biarmosuchus tagax
_____________________________________________________________________________________
RUSSIA
(?SAVVATII, VELIKII USTYUG DISTRICT, VOLOGDA REGION, RUSSIA)
ARKHANGEL’SK PROVINCE
MALAYA NORTH DVINA RIVER
MEDVEDKOVA
SALAREVSKAYA SVITA
MIDDLE PART OF THE VYATKIAN REGIONAL STAGE
UPPER TATARIAN AGE
LOPINGIAN SUBEPOCH
UPPER ZECHSTEIN EPOCH
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
NEOTETRAPODA
SUBCLASS: REPTILIOMORPHA SAVE-SODERBERGH, 1934
ORDER: SEYMOURIAMORPHA WATSON, 1917
SUBORDER: CHRONIOSUCHIDA TATARINOV, 1971
Family: CHRONIOSUCHIDAE Vjuschkov, 1957
*Chroniosuchus licharevi
= Jugosuchus licharevi
_____________________________________________________________________________________
EUROPEAN RUSSIA
ARKHANGELSK REGION
KOTLAS DISTRICT
MALAYA SEVERNAYA DVINA RIVER
ZAVARZHYE LOCALITY
VYATKA REGIONAL STAGE
UPPER TATARIAN AGE
LOPINGIAN SUBEPOCH
UPPER ZECHSTEIN EPOCH
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPID BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: ANOMODONTIA WATSON & ROMER, 1956
SUBORDER: DICYNODONTIA OWEN, 1860
Superfamily: PRISTERODONTIA Cluver & King, 1983
Family: DICYNODONTOIDAE Owen, 1859
Subfamily: CRYPTODONTINAE Owen, 1859
Tribe: DICYNODONTINI Owen, 1859
Dicynodon trautscholdi
= *Elph borealis
_____________________________________________________________________________________
EUROPEAN RUSSIA
ARKANGELSK PROVINCE
KOTLASSKII (KOTLAS) DISTRICT
BASIN OF THE NORTHERN DVINA RIVER
SOKOLKI LOCALITY
TATARIAN SERIES
UPPER VYATKIAN SUBSTAGE
VYATKIAN HORIZON (NORTHERN DVINA HORIZON, BOTTOM OF THE VYATKIAN HORIZON)
SOKOLKI FAUNAL ASSEMBLAGE
SOKOLKI FAUNAL SUBASSEMBLAGE
SCUTOSAURUS KARPINSKII ZONE (DICYNODON ASSEMBLAGE ZONE, LUCAS, 2005)
ZONE IV
UPPER TATARIAN AGE
LOPINGIAN SUBEPOCH
UPPER ZECHSTEIN EPOCH
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LABYRINTHODONTIA OWEN, 1860
SUPERORDER: TEMNOSPONDYLIA ZITTEL, 1887
MICROORDER: LIMNARCHIA YATES & WARREN, 2000
ORDER: DVINOSAURIA YATES & WARREN, 2000
INFRAORDER: TRIMERORHACHOIDA ROMER, 1947
Superfamily: DVINOSAUROIDEA Watson, 1919
Family: DVINOSAURIDAE Amalitzky, 1921
*Dvinosaurus primus
= *Dvinosaurus secundus
= *Dvinosaurus tertiusNEOTETRAPODA
SUBCLASS: REPTILIOMORPHA SAVE-SODERBERGH, 1934
ORDER: SEYMOURIAMORPHA WATSON, 1917
SUBORDER: CHRONIOSUCHIDA TATARINOV, 1971
Family: CHRONIOSUCHIDAE Vjuschkov, 1957
Chroniosuchus licharevi
SUBORDER: SEYMOURIDA TATARINOV, 1971
Superfamily: KOTLASSOIDEA Romer, 1933 (sensu Vjuschkov, 1957)
Family: KOTLASSIDAE Romer, 1933
Subfamily: KOTLASSIINAE Romer, 1933
*Kotlassia prima
Superfamily: SEYMOURIOIDEA Williston, 1911 (sensu Tatarinov, 1971)
Family: KARPINSKIOSAURIDAE Sushkin, 1925
Subfamily: KARPINSKIOSAURINAE Sushkin, 1925
*Karpinskiosaurus secunda
= Kotlassia secunda
= *Karpinskiosaurus neglectus
= Karpinskiosaurus secundus
= Kotlassia primaCLASS: REPTILIA LAURENTI, 1768
SUBCLASS: PARAREPTILIA OLSON 1947
INFRACLASS: SAUROPSIDA HUXLEY, 1864
ORDER: PROCOLOPHONOMORPHA ROMER, 1964
SUBORDER: PAREIASAUROIDEA SELEY, 1888
Family: PARAEIASAURIDAE Cope, 1869
Subfamily: PAREIASAURINAE Nopcsa, 1923
Scutosaurus karpinskii
= Scutosaurus tuberculatusSUBCLASS: SYNAPSIDA OSBORN, 1903
INFRACLASS DINOMORPHA IVAKNENKO, 2008
SUPERORDER: GORGONTIA IVAKNENKO, 2008
ORDER: GORGONOPSIA SEELEY, 1895
SUBORDER: GORGONOPIA SEELEY, 1895
Superfamily: GORGONOPIOIDEA Lydekker, 1890
Family: GORGONOPIDAE Lydekker, 1890
Subfamily: GORGONOPINAE Sigogneau, 1970
*Pravoslavelvia parva
= ?Inostrancevia parvaSuperfamily: INOSTRANCEVIOIDEA Huene, 1948
Family: INOSTRANCEVIIDAE Pravoslavlev, 1927
*Inostrancevia alexandri
*Inostrancevia latifronsNEOTHERAPSIDA RUBIDGE & SIDOR, 2001
SUPERORDER: ANOMODONTIA WATSON & ROMER, 1956
ORDER: DICYNODONTIA OWEN, 1860
SUBORDER: DICYNODONTIA OWEN, 1860
Superfamily: PRISTERODONTIA Cluver & King, 1983
Family: DICYNODONTOIDAE Owen, 1859
Subfamily: CRYPTODONTINAE Owen, 1859
Tribe: DICYNODONTINI Owen, 1859
*Dicynodon trautscholdi
= *Dicynodon amalitzkii
= Elph borealisINFRACLASS: THERIODONTIA EMEND OWEN, 1881
ORDER: EUTHERIODONTIA EMEND RUBIDGE & SIDOR, 2001
SUBORDER: THEROCEPHALIA BROOM, 1903
Superfamily: WHAITSIOIDEA Haughton, 1918
Family: ANNATHERAPSIDAE Kuhn, 1961
*Annatherapsidus petri
= Anna petriSUBORDER: CYNODONTIA OWEN, 1860
INFRAORDER: PROCYNOSUCHIA
Family: DVINIIDAE Tatarinov, 1968
*Dvinia prima
_____________________________________________________________________________________
EUROPEAN RUSSIA
ARKHANGEL’SK REGION
NORTHERN DVINA
UPPER TATARIAN AGE
LOPINGIAN SUBEPOCH
UPPER ZECHSTEIN EPOCH
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPIDA BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: THERIODONTIA OWEN, 1881
EUTHERIODONTIA
SUBORDER: CYNODONTIA OWEN, 1860
INFRAORDER: PROCYNOSUCHIA
Family: DVINIIDAE Tatarinov, 1968
Dvinia prima
= *Permocynodon sushkini
_____________________________________________________________________________________
EUROPEAN RUSSIA
ARKHANGEL'SK REGION
MEZEN DISTRICT
KIMJA RIVER
ZONE 111
RED MARLS HORIZON
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
NEOTETRAPODA
SUBCLASS: REPTILIOMORPHA SAVE-SODERBERGH, 1934
ORDER: SEYMOURIAMORPHA WATSON, 1917
SUBORDER: SEYMOURIDA TATARINOV, 1971
Superfamily: SEYMOURIOIDEA Williston, 1911 (sensu Tatarinov, 1971)
Family: NYCTIPHRURETIDAE Efremov, 1938
*Nyctiphruretus acudens
_____________________________________________________________________________________
EUROPEAN RUSSIA
LOWER MEZEN' RIVER AREA
ARKHANGEL'SK REGION
ZONE III
LOWER TATARIAN AGE
LOPINGIAN SUBEPOCH
UPPER ZECHSTEIN EPOCH
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
NEOTETRAPODA
SUBCLASS: REPTILIOMORPHA SAVE-SODERBERGH, 1934
ORDER: SEYMOURIAMORPHA WATSON, 1917
SUBORDER: NYCTEROLETERIDAE TATARINOV, 1972
Family: NYCTEROLETERIDAE Romer, 1956
Subfamily: NYCTEROLECTERINAE Romer, 1956
*Macroleter poezicus
_____________________________________________________________________________________
EUROPEAN RUSSIA
ARKHANGELSK REGION
UST’-PEZA
LOWER TATARIAN AGE
LOPINGIAN SUBEPOCH
UPPER ZECHSTEIN EPOCH
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
NEOTETRAPODA
SUBCLASS: REPTILIOMORPHA SAVE-SODERBERGH, 1934
ORDER: SEYMOURIAMORPHA WATSON, 1917
SUBORDER: NYCTEROLETERIDAE TATARINOV, 1972
Family: NYCTEROLETERIDAE Romer, 1956
Subfamily: BASHKYROLETERINAE Ivachnenko, 1997
Bashkyroleter mesensis
_____________________________________________________________________________________
EUROPEAN RUSSIA
ARKHANGELSK REGION
KOTLASSKII DISTRICT
BOTTOM OF VYATKIAN HORIZON
UPPER TATARIAN AGE
CAPITANIAN STAGE
MIDDLE ZECHSTEIN EPOCH
LATE MIDDLE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPID BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: ANOMODONTIA WATSON & ROMER, 1956
SUBORDER: DICYNODONTIA OWEN, 1860
Superfamily: PRISTERODONTIA Cluver & King, 1983
Family: DICYNODONTOIDAE Owen, 1859
Subfamily: CRYPTODONTINAE Owen, 1859
Tribe: DICYNODONTINI Owen, 1859
Dicynodon trautscholdi
_____________________________________________________________________________________
NORTH EUROPEAN RUSSIA
ARKHANGEL’SK PROVINCE
KOTLASSKII (KOTLAS) DISTRICT
MALAYA SEVERNAYA DVINA RIVER
ZAVARZHYE LOCALITY
TATARIAN SERIES
VYATKIAN SUBSTAGE
VYATKIAN HORIZON
VYATKA REGIONAL STAGE
SOKOLKI FAUNAL ASSEMBLAGE
SOKOLKI FAUNAL SUBSTAGE
UPPER TATARIAN AGE
LOPINGIAN SUBEPOCH
UPPER ZECHSTEIN EPOCH
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LABYRINTHODONTIA OWEN, 1860
SUPERORDER: TEMNOSPONDYLIA ZITTEL, 1887
MICROORDER: LIMNARCHIA YATES & WARREN, 2000
ORDER: DVINOSAURIA YATES & WARREN, 2000
INFRAORDER: TRIMERORHACHOIDA ROMER, 1947
Superfamily: DVINOSAUROIDEA Watson, 1919
Family: DVINOSAURIDAE Amalitzky, 1921
Dvinosaurus primus
NEOTETRAPODA
SUBCLASS: REPTILIOMORPHA SAVE-SODERBERGH, 1934
ORDER: SEYMOURIAMORPHA WATSON, 1917
SUBORDER: CHRONIOSUCHIDA TATARINOV, 1971
Family: CHRONIOSUCHIDAE Vjuschkov, 1957
Chroniosuchus licharevi
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: PARAREPTILIA OLSON 1947
INFRACLASS: SAUROPSIDA HUXLEY, 1864
ORDER: PROCOLOPHONOMORPHA ROMER, 1964
SUBORDER: PAREIASAUROIDEA SELEY, 1888
Family: PARAEIASAURIDAE Cope, 1869
Subfamily: PAREIASAURINAE Nopcsa, 1923
Scutosaurus karpinskii
SUBCLASS: SYNAPSIDA OSBORN, 1903
INFRACLASS DINOMORPHA IVAKNENKO, 2008
SUPERORDER: GORGONTIA IVAKNENKO, 2008
ORDER: GORGONOPSIA SEELEY, 1895
SUBORDER: GORGONOPIA SEELEY, 1895
Superfamily: INOSTRANCEVIOIDEA Huene, 1948
Family: INOSTRANCEVIIDAE Pravoslavlev, 1927
Inostrancevia latifrons
NEOTHERAPSIDA RUBIDGE & SIDOR, 2001
SUPERORDER: ANOMODONTIA WATSON & ROMER, 1956
ORDER: DICYNODONTIA OWEN, 1860
SUBORDER: DICYNODONTIA OWEN, 1860
Superfamily: PRISTERODONTIA Cluver & King, 1983
Family: DICYNODONTOIDAE Owen, 1859
Subfamily: CRYPTODONTINAE Owen, 1859
Tribe: DICYNODONTINI Owen, 1859
Dicynodon trautscholdi
= Elph borealis
_____________________________________________________________________________________
EUROPEAN RUSSIA
CIS-URALS
ARKHANGEL’SK PROVINCE
0.5 KM BELOW THE MOUTH OF NIOZHNYAYA SYAMZHEN’GA RIVER
PIZHMA MEZENSKAYA FORMATION
EARLY TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: PARAREPTILIA OLSON 1947
INFRACLASS: SAUROPSIDA HUXLEY, 1864
ORDER: PROCOLOPHONOMORPHA ROMER, 1964
SUBORDER: PROCOLPHOMORPHA SEELEY, 1888
Family: PROCOLOPHONIDAE, Seeley, 1888
Subfamily: PROCOLOPHONINAE Seeley, 1888 (sensu Romer, 1956)
Timanophon raridentatus
= Burtensia burtensis
*Timanophon raridentatus
_____________________________________________________________________________________
EUROPEAN RUSSIA
CIS-URALS
ARKHANGEL’S SK REGION
KHEY-YAGA RIVER BASIN
EARLY TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: PARAREPTILIA OLSON 1947
INFRACLASS: SAUROPSIDA HUXLEY, 1864
ORDER: PROCOLOPHONOMORPHA ROMER, 1964
SUBORDER: PROCOLPHOMORPHA SEELEY, 1888
Family: PROCOLOPHONIDAE, Seeley, 1888
Subfamily: PROCOLOPHONINAE Seeley, 1888 (sensu Romer, 1956)
Orenburgia bruma
_____________________________________________________________________________________
EUROPEAN RUSSIA
CIS-URALS
ARKHANGEL’S SK REGION
MESCHERYAKOVKA LOCALITY
EARLY TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: PARAREPTILIA OLSON 1947
INFRACLASS: SAUROPSIDA HUXLEY, 1864
ORDER: PROCOLOPHONOMORPHA ROMER, 1964
SUBORDER: PROCOLPHOMORPHA SEELEY, 1888
Family: PROCOLOPHONIDAE, Seeley, 1888
Subfamily: PROCOLOPHONINAE Seeley, 1888 (sensu Romer, 1956)
Burtensia burtensis
= Tichvinskia burtensis
_____________________________________________________________________________________
NORTHERN RUSSIA
ARKHANGEL’SK PROVINCE
NORTH DVINA AND ILJINSKOJE
LOWER VYATSKIAN GORIZONT
ZONE IV
UPPER TATARIAN AGE
LOPINGIAN SUBEPOCH
ZECHSTEIN EPOCH
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: PARAREPTILIA OLSON 1947
INFRACLASS: SAUROPSIDA HUXLEY, 1864
ORDER: PROCOLOPHONOMORPHA ROMER, 1964
SUBORDER: PAREIASAUROIDEA SELEY, 1888
Family: PARAEIASAURIDAE Cope, 1869
Subfamily: PAREIASAURINAE Nopcsa, 1923
Scutosaurus karpinskii
_____________________________________________________________________________________
EUROPEAN RUSSIA
CIS-URALS
ARKHANGEL’SK PROVINCE
NENETSKIY NATIONAL RESERVATION
3.5 KM ABOVE MOUTH OF LESTANSHOR CREEK
LOWER SUBFORMATION OF LESTANSHOR CREEK
EARLY TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: PARAREPTILIA OLSON 1947
INFRACLASS: SAUROPSIDA HUXLEY, 1864
ORDER: PROCOLOPHONOMORPHA ROMER, 1964
SUBORDER: PROCOLPHOMORPHA SEELEY, 1888
Family: PROCOLOPHONIDAE, Seeley, 1888
Subfamily: PROCOLOPHONINAE Seeley, 1888 (sensu Romer, 1956)
*Lestanshoria massiva
_____________________________________________________________________________________
EUROPEAN RUSSIA
CIS-URALS
ARKHANGEL’SK PROVINCE
MEZEN RIVER BASIN
RIGHT BANK OF PEZO RIVER
UPPER KAZANIAN STAGE
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: PARAREPTILIA OLSON 1947
INFRACLASS: SAUROPSIDA HUXLEY, 1864
ORDER: LANTHANOSUCHIDAE TATARINOV, 1971
Family: LANTHANOSUCHIDAE Efremov, 1946
*Lanthaniscus efremovi
_____________________________________________________________________________________
EUROPEAN RUSSIA
ARKHANGEL’SK REGION
PIZHMA-I LOCALITY
EARLY TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: PARAREPTILIA OLSON 1947
INFRACLASS: SAUROPSIDA HUXLEY, 1864
ORDER: PROCOLOPHONOMORPHA ROMER, 1964
SUBORDER: PROCOLPHOMORPHA SEELEY, 1888
Family: PROCOLOPHONIDAE, Seeley, 1888
Subfamily: PROCOLOPHONINAE Seeley, 1888 (sensu Romer, 1956)
Timanophon raridentatus
= Burtensia burtensis
_____________________________________________________________________________________
EUROPEAN RUSSIA
ARKHANGEL’SK REGION
6 KM FROM THE MOUTH OF THE LEFT BANK OF VYBOR RIVER
PIZHMAN MEZENSKAYA FORMATION
EARLY TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: PARAREPTILIA OLSON 1947
INFRACLASS: SAUROPSIDA HUXLEY, 1864
ORDER: PROCOLOPHONOMORPHA ROMER, 1964
SUBORDER: PROCOLPHOMORPHA SEELEY, 1888
Family: PROCOLOPHONIDAE, Seeley, 1888
Subfamily: PROCOLOPHONINAE Seeley, 1888 (sensu Romer, 1956)
Timanophon raridentatus
= Burtensia burtensis
_____________________________________________________________________________________
ASTRAKHANSKAYA OBLAST
EUROPEAN RUSSIA
ASTRAKHANSKAYA OBLAST
LAKE BASKUNCHAK
MT. GREAT BOGDO JUST NORTH OF THE MOUTHS OF THE VOLGA
ZONE VI
BASKUNCHAK SERIES
EARLY TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LABYRINTHODONTIA OWEN, 1860
SUPERORDER: TEMNOSPONDYLIA ZITTEL, 1887
MICROORDER: LIMNARCHIA YATES & WARREN, 2000
ORDER: CAPITOSAURIA ROMER, 1947
Superfamily: CAPITOSAUROIDEA Watson, 1919 (sensu Romer, 1947)
Incertae sedis
Non *Parotosaurus bogdoanus
= Capitosaurus sp
= Capitosaurus bogdoanus
= Parotosaurus bogdoanus
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EUROPEAN RUSSIA
ARKHANGELSK REGION
NENETS NATIONAL DISTRICT
KHEI-YAGA 4 LOCALITY
LESTANSHORSKAYA FROMATION
LOWER SUBFORMATION
LATEST OLENEKIAN STAGE (LATEST SPATHIAN STAGE)
LATEST SCYTHIAN EPOCH (LATEST EARLY TRIASSIC EPOCH)
LATEST EARLY TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
MAGNORDER: PROTOROSAURIA HUXLEY, 1871
SUPERORDER: TAPINOPLATIA PETERS EMEND NOVA
GRANDORDER: CHARACIOPODA PETERS EMEND NOVA
Family: TANYSTROPHEIDAE Geravis, 1858
Gen. sp indet.
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