ORYOL REGION
SOUTHWEST OF THE LIVNY TOWN
GORNOSTAYEVKA QUARRY
?TRIAGULARIS-CREPIDA ZONE OF THE STANDARD CODONT SCALE
ZADONSKIAN REGIONAL STAGE
LOWER FAMENNIAN STAGE
UPPER LATE DEVONIAN EPOCH
LATE DEVONIAN
STEM: TETRAPODA GOODRICH, 1930
Incertae sedis
*Jakubsonia livnensis