PENZA OBLAST
NEAR THE MIRONOSITSKOE CEMETERY
WITHIN THE CITY OF PENZA
OUTCROP IN THE PROLOM RAVINE
MAASTRICHTIAN STAGE
UPPERMOST SENONIAN SUBEPOCH
UPPERMOST GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: LACERTILIA OWEN, 1842
GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961
MICRORDER: SCINCOMORPHA CAMP, 1923
ORDER: ANGUIMORPHA FURBRINGER, 1900
SUBORDER: DIPLOGLOSSA COPE, 1864
INFRAORDER: THECAGLOSSA COPE, 1890
PARVORDER: MONSTERSAURIA NORELL & GAO, 1977
Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)
Family: MOSASAURIDAE Gervais, 1853
Infrafamily: NATANIA
Subfamily: MOSASAURINAE Gervais, 1853 (sensu Williston, 1897)
Tribe: MOSASAURINI Gervais, 1853 (sensu Russell, 1967)
Mosasaurus hoffmanni
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APTIAN STAGE
MIDDLE GALLIC SUBEPOCH
LOWER CRETACEOUS EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
NEODIAPSIDA: INCERTAE SEDIS
OR AMNIOTA: INCERTAE SEDIS
MAGANORDER: ICHTHYOPTERYGIA BAUR, 1887 emend nova
SUPERORDER: EOICHTHYOSAURIA MOTANI, 1999
GRANDORDER: ICHTHYOSAURIA EMEND BLAINVILLE, 1835
Incertae sedis
Gen. sp indet.
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EUROPEAN RUSSIA
PENZA REGION
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: EURYAPSIDA COLBERT, 1945
SUPERORDER: SAUROPTERYGIA OWEN 1860
GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994
ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
INFRAORDER: PLESIOSAUROMORPHA NOVA
EUPLESIOSAURIA O’KEEFE, 2001
Superfamily: CRYPTOCLEIDOIDEA Williston, 1925
TRICLEDIA O’KEEFE , 2001
Family: POLYCOTYLIDAE Cope, 1869
Incertae sedis
Neopliosaurus (no species given)
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RUSSIA (OLD PRUSSIA)
PENZA REGION
SERDOBA (OLD GUMBINNEN)
UPPER SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: EURYAPSIDA COLBERT, 1945
SUPERORDER: SAUROPTERYGIA OWEN 1860
GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994
ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
INFRAORDER: PLESIOSAUROMORPHA NOVA
EUPLESIOSAURIA O’KEEFE, 2001
Superfamily: ELASMOSAUROIDEA Cope, 1869
Family: ELASMOSAURIDAE Cope, 1869
Incertae sedis
non Elasmosaurus, *E. helmersenii
= Plesiosaurus helmersenii
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EUROPEAN RUSSIA
PENZA REGION
PETROVSK
SERDOBA
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: EURYAPSIDA COLBERT, 1945
SUPERORDER: SAUROPTERYGIA OWEN 1860
GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994
ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
INFRAORDER: PLESIOSAUROMORPHA NOVA
EUPLESIOSAURIA O’KEEFE, 2001
Superfamily: ELASMOSAUROIDEA Cope, 1869
Family: ELASMOSAURIDAE Cope, 1869
Incertae sedis
non Elasmosaurus, *E. serdobensis
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EUROPEAN RUSSIA
PENZA PROVINCE
ZATOLOKINO VILLAGE
SANTONIAN STAGE
SENONIAN SUBEPOCH
LOWER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: EURYAPSIDA COLBERT, 1945
SUPERORDER: SAUROPTERYGIA OWEN, 1860
GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994
ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
INFRAORDER: PLIOSAUROMORPHA NOVA
Superfamily: PLIOSAUROIDEA Seeley, 1874
Family: PLIOSAURIDAE Seeley, 1874
*Georgiasaurus penzensis
= Georgia penzensis
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EUROPEAN RUSSIA
NOW PENZA PROVINCE (OLD PROVINCE OF SARATOV)
FOLGA REGION
MARINE SEDIMENTS NEAR THE VILLAGE OF MALAYA SERDOBA
CONIACIAN?-SANTONIAN OR POSSIBLY EARLY CAMPANIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
EARLY LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
SUPERORDER: TAPINOPLATIA PETERS, 2000
GRANDORDER: CHARACIOPODA PETERS, 2000
MICRORDER: FENESTRASAURIA PETERS, 2000
ORDER: PTEROSAURIA KAUP, 1834
SUBORDER: EUPTERODACTYLOIDEA BENNETT, 1994
Superfamily: AZHDARCHIDEA Nesov, 1984
Family: AZHDARCHIDAE Nesov, 1984 emend Padian, 1986
*Bogoloubovia orientalis
= Ornithostoma orientale
= Ornithostoma orientalis
= Pteranodon orientalisGen. sp indet.
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