FRANCE
BOUCHES-DU-RHONE DEPARTMENT
AIX-EN-PROVINCE BASIN
VELAUX
LA BASTIDE NEUVE
GLAUCONITIC GRAINSTONE LIMESTONE
LATE CAMPANIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN,
1960
SUPERCOHORT: TESTUDINES LINNAEUS,
1758
COHORT: CASICHELYDIA
emend GAFFNEY & MEYLAN, 1988
MAGANORDER: CRYPTODIRA
GAFFNEY & MEYLAN, 1988
ORDER: CRYPTODIRA COPE, 1868
Family: SOLEMYDIDAE
Lapparent & Murelaga, 1997
Gen. sp indet.
MAGNORDER: PLEURODIRA GAFFNEY & MEYLAN,
1988
ORDER: PLEURODIRIDA COPE,
1864
SUBORDER: EUPLEURODIRA
GAFFNEY & MEYLAN, 1988
Superfamily: BOTHREMYDINURA Baur, 1891
Epifamily: BOTHERMYDINURA Baur, 1891
Gen. sp indet.
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON,
1985
LEGION: ARCHOSAUROMORPHA
HUENE, 1946
SUBLEGION: DINOSAUROMORPHA
BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES
NOVAS, 1992
SUPERCOHORT: DINOSAURIA
OWEN, 1842
COHORT: ORNITHISCHIFORMES
COOPER, 1985
SUPERORDER: SAUROPODOMORPHA
HUENE, 1932
GRANDORDER: BRONTOSAURIA
OLSHEVSKY, 1991 (emend)
MICRORDER:
SAUROPODA MARSH, 1878
ORDER: EUSAUROPODA
UPCHURCH, 1995
SUBORDER:
NEOSAUROPODA BONAPARTE, 1986
INFRAORDER:
MACRONARIA WILSON & SERENO, 1998
PARVORDER:
TITANOSAURIFORMES SALGADO ET AL, 1997
Superfamily:
TITANOSAUROIDEA Lydekker, 1885 (sensu
Upchurch, 1999)
Family:
TITANOSAURIDAE Lydekker, 1885
Gen. sp indet.
Subfamily: LIRAINOSAURINAE Diez Diaz, Garcia, Pereda Suberbiola, & Valentin, 2012
*Garrigatitan meridionalis
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
Family:
RHABDODONTIDAE Weishampel, Jainu, Csiki & Norman,
2003
Gen. sp indet.
_____________________________________________________________________________________
FRANCE
BOUCHES-DU-RHONE DEPARTMENT
AIX-EN-PROVENCE BASIN
VELAUX-LA BASTIDE NEUVE
BEGUDIAN (LOCAL STAGE) SANDSTONES.
LATE
CAMPANIAN STAGE
SENONIAN
SUBEPOCH
UPPERMOST
GULF EPOCH
LATE
CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL,
1866
CLASS: REPTILIA LAURENTI,
1768
SUBCLASS: DIAPSIDA OSBORN,
1904
SUPERLEGION: NEODIAPSIDA
BENTON, 1985
LEGION: ARCHOSAUROMORPHA
HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA
BENTON, 1999
COHORT: PROLACERTIFORMES
CAMP, 1945 NOVA
SUPERORDER: TAPINOPLATIA
PETERS, 2000
GRANDORDER:
CHARACIOPODA PETERS, 2000
MICRORDER:
FENESTRASAURIA PETERS, 2000
ORDER:
PTEROSAURIA KAUP, 1834
SUBORDER:
EUPTERODACTYLOIDEA BENNETT, 1994
Superfamily:
AZHDARCHIDEA Nesov, 1984
Family:
AZHDARCHIDAE Nesov, 1984 emend Padian, 1986
*Mistralazhdarcho maggii
SUBLEGION: DINOSAUROMORPHA
BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
Family:
RHABDODONTIDAE Weishampel, Jainu, Csiki & Norman,
2003
*Matheronodon provincialis
_____________________________________________________________________________________
FRANCE
FRANCE
BOUCHES-DU-RHONE DEPARTMENT
FUVELIA
BASIN
VALDONNE
LIGNITE MINE
'LA GRANDE
MENE'
FUVELIAN
LIGNITES
EARLY
CAMPANIAN STAGE
SENONIAN
SUBEPOCH
UPPERMOST
GULF EPOCH
LATE
CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI
SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI
BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA
WHETSTONE & WHYBROW, 1983
ORDER: EUSUCHIA emend HUXLEY, 1875
ORDER: EUSUCHIA emend HUXLEY, 1875
Family: ALLODAPOSUCHIDAE Narvaez, Brochu, Escaso, Perez-Garcia, & Ortega,
2015
*Massaliasuchus
affuvelensis = Crocodilus affuvelensis
_____________________________________________________________________________________
SOUTHERN FRANCE
BOUCHES DU RHONE DEPARTMENT
VELAUX-LA BASTIDE NEUVE
LATE CAMPANIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI
SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI
BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA
WHETSTONE & WHYBROW, 1983
ORDER: EUSUCHIA emend HUXLEY, 1875
ORDER: EUSUCHIA emend HUXLEY, 1875
Family: ALLODAPOSUCHIDAE Narvaez, Brochu, Escaso, Perez-Garcia, & Ortega,
2015
Allodaposuchus precedens
*Allodaposuchus iberoarmoricanus
_____________________________________________________________________________________
FRANCE
BOUCHES-DU-RHONE DEPARTMENT
AIX-EN-PROVINCE BASIN
VILLAGE OF ROUSSET
LA CAIRANNE-HIGHWAY
LATE MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: MAMMALIA LINNAEUS, 1758
SUBCLASS: PROTOTHERIA GILL, 1872
INFRACLASS: TRECHNOTHERIA McKENNA,
1975
LEGION: CLADOTHERIA McKENNA,
1975
SUBLEGION: ZATHERIA MCKENNA,
1975
INFRALEGION: TRIBOSPHENIDA
(McKENNA, 1975) McKENNA & BELL, 1997
SUPERCOHORT: THERIA
(PARKER & HASWELL, 1987) McKENNA & BELL, 1997
COHORT: PLACENTALIA
(OWEN, 1837) McKENNA & BELL, 1997
MAGNORDER: EPITHERIA
(McKENNA, 1975) McKENNA & BELL, 1997
SUPERORDER: PREPTOTHERIA
McKENNA, 1975
GRANDORDER:
UNGULATOMORPHA ARCHIBALD, 1996
Family: ZHELESTIDAE
Nesov, 1990
*Mistralestes
arcensis
_____________________________________________________________________________________
FRANCE
BOUCHES-DU-RHONE DEPARTMENT
ARC BASIN
VITROLLES LA PLAINE SITE
LATE MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN,
1960
SUPERCOHORT: TESTUDINES LINNAEUS,
1758
COHORT: CASICHELYDIA emend
GAFFNEY & MEYLAN, 1988
MAGNORDER: PLEURODIRA GAFFNEY & MEYLAN,
1988
ORDER: PLEURODIRIDA COPE,
1864
SUBORDER: EUPLEURODIRA
GAFFNEY & MEYLAN, 1988
Superfamily: BOTHREMYDINURA Baur, 1891
Epifamily: BOTHERMYDINURA Baur, 1891
Subfamily: BOTHREMYDINAE Baur, 1891
Infrafamily: BOTHREMYDODDA Baur, 1891
Tribe: FOXEMYDINA Gaffney, Tong & Meylan, 2006
Cf.
Polysternon
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON,
1985
LEGION: LEPIDOSAUROMORPHA DUMERIL & BIBRON,
1839
SUBLEGION: LEPIDOSAURIA HAECKEL,
1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA OPPEL,
1811
Incertae sedis
Gen. sp indet.
LEGION: ARCHOSAUROMORPHA HUENE,
1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI,
1990
COHORT: CROCODYLOTARSI
BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS,
1974
SUPERORDER: CROCODYLOMORPHA
WALKER, 1968
GRANDORDER: CROCODILIFORMES
HAY, 1930
MICRORDER: MESOEUROCROCODYLIA
WHETSTONE & WHYBROW, 1983
ORDER: EUSUCHIA emend HUXLEY, 1875
Family: ALLODAPOSUCHIDAE Narvaez, Brochu, Escaso, Perez-Garcia, & Ortega,
2015
Cf.
Musturzabalsuchus
SUBLEGION: DINOSAUROMORPHA
BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES
NOVAS, 1992
SUPERCOHORT: DINOSAURIA
OWEN, 1842
COHORT: THEROPODA EMEND
MARSH, 1881
Incertae sedis
Ricardoestesia sp
Gen. sp indet.
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA
BAKKER, 1986
GRANDORDER: CERATOSAURIA
NOVAS, 1992
AVEROSTRA PAUL,
2001
ORDER: CERATOSAURIA
MARSH, 1884
Gen. Sp indet.
COHORT: ORNITHISCHIFORMES
COOPER, 1985
SUPERORDER: SAUROPODOMORPHA
HUENE, 1932
GRANDORDER: BRONTOSAURIA
OLSHEVSKY, 1991 (emend)
MICRORDER: SAUROPODA
MARSH, 1878
ORDER: EUSAUROPODA
SUBORDER:
NEOSAUROPODA BONAPARTE, 1986
INFRAORDER:
MACRONARIA WILSON & SERENO, 1998
PARVORDER:
TITANOSAURIFORMES SALGADO ET AL, 1997
Family:
TITANOSAURIDAE Lydekker, 1885
Gen.
Sp indet.
MAGNORDER:
ORNITHISCHIA SEELEY, 1888
SUPERORDER:
NEORNITHISCHIA COOPER, 1985
GRANDORDER:
ORNITHOPODA MARSH, 1871
MICRORDER:
EUORNITHOPODA SERENO, 1986
ORDER:
IGUANODONTIA DOLLO, 1888
Family:
RHABDODONTIDAE Weishampel, Jainu, Csiki & Norman,
2003
Gen.
sp indet.
SUBORDER:
ANKYLOPOLLEXIA SERENO, 1986
PARVORDER:
HADROSAURIFORMES SERENO, 1997
SUPERFAMILY:
HADROSAUROIDEA COPE, 1869 (SENSU SERENO,
1986)
Family:
HADROSAURIDAE COPE, 1869
Gen.
sp indet.
CLASS: MAMMALIA LINNAEUS, 1758
SUBCLASS: PROTOTHERIA GILL, 1872
INFRACLASS: TRECHNOTHERIA McKENNA,
1975
LEGION: CLADOTHERIA McKENNA,
1975
SUBLEGION: ZATHERIA MCKENNA,
1975
INFRALEGION: TRIBOSPHENIDA
(McKENNA, 1975) McKENNA & BELL, 1997
SUPERCOHORT: THERIA
(PARKER & HASWELL, 1987) McKENNA & BELL, 1997
COHORT: PLACENTALIA
(OWEN, 1837) McKENNA & BELL, 1997
MAGNORDER: EPITHERIA
(McKENNA, 1975) McKENNA & BELL, 1997
SUPERORDER: PREPTOTHERIA
McKENNA, 1975
GRANDORDER:
UNGULATOMORPHA ARCHIBALD, 1996
Family: ZHELESTIDAE
Nesov, 1990
*Valentinella
parvus
_____________________________________________________________________________________
FRANCE
BOUCHES-DU-RHONE DEPARTMENT
AIX-EN-PROVINCE BASIN
VELAUX
LA BASTIDE NEUVE
GLAUCONITIC GRAINSTONE LIMESTONE
LATE CAMPANIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON,
1985
LEGION: ARCHOSAUROMORPHA
HUENE, 1946
SUBLEGION: DINOSAUROMORPHA
BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES
NOVAS, 1992
SUPERCOHORT: DINOSAURIA
OWEN, 1842
COHORT: ORNITHISCHIFORMES
COOPER, 1985
SUPERORDER: SAUROPODOMORPHA
HUENE, 1932
GRANDORDER: BRONTOSAURIA
OLSHEVSKY, 1991 (emend)
MICRORDER:
SAUROPODA MARSH, 1878
ORDER: EUSAUROPODA
UPCHURCH, 1995
SUBORDER:
NEOSAUROPODA BONAPARTE, 1986
INFRAORDER:
MACRONARIA WILSON & SERENO, 1998
PARVORDER:
TITANOSAURIFORMES SALGADO ET AL, 1997
Superfamily:
TITANOSAUROIDEA Lydekker, 1885 (sensu
Upchurch, 1999)
Family:
TITANOSAURIDAE Lydekker, 1885
*Atsinganosaurus
valauciensis
Genus nova
_____________________________________________________________________________________
FRANCE
BOUCHES-DU-RHONE DEPARTMENT
VITROLLES-COUPERIGNE LOCALITY
EARLY ROGNACIAN = EARLY MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI
SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI
BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA
WHETSTONE & WHYBROW, 1983
ORDER: MESOSUCHIA EMEND HUXLEY, 1875
PARVORDER:
METASUCHIA BENTON & CLARK, 1988
INFRAORDER: METAMESOSUCHIA HULKE, 1878
NEOSUCHIA
CLARK, IN BENTON & CLARK, 1988
Family: TREMATOCHAMPSIDAE Buffetaut, 1974
Ischyrochampsa meridionalis
SUBLEGION: DINOSAUROMORPHA
BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA PAUL, 2002
AVEPECTORA PAUL, 2002
SUBORDER: MANIRAPTORA GAUTHIER, 1986
INFRAORDER: PARVES SERENO, 1997
PARVORDER:
DEINONYCHOSAURIA COLBERT & D.
A. RUSSELL, 1969
Family: DROMAEOSAURIDAE Russell, 1969
Gen. sp indet.
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
Family:
RHABDODONTIDAE Weishampel, Jainu, Csiki & Norman,
2003
Rhabdodon priscus
= Rhabdodon priscum
_____________________________________________________________________________________
FRANCE
BOUCHES-DU-RHONE DEPARTMENT
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA
BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
SUPERORDER: SAUROPODOMORPHA HUENE, 1932
GRANDORDER: BRONTOSAURIA OLSHEVSKY, 1991 (emend)
MICRORDER: SAUROPODA MARSH, 1878
ORDER: EUSAUROPODA
SUBORDER: NEOSAUROPODA BONAPARTE, 1986
INFRAORDER:
MACRONARIA WILSON & SERENO, 1998
PARVORDER: TITANOSAURIFORMES SALGADO ET AL, 1997
Family: TITANOSAURIDAE Lydekker, 1885
Subfamily: TITANOSAURINAE Lydekker, 1885 (sensu Nopcsa, 1928)
Hypselosaurus priscus
_____________________________________________________________________________________
FRANCE
BOUCHES-DU-RHONE DEPARTMENT
LAMEAU DU BEAUSSET
FUVELIEN
LOWER CAMPANIAN STAGE
UPPER SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA
BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: CERATOSAURIA NOVAS, 1992
AVEROSTRA PAUL, 2001
ORDER: CERATOSAURIA MARSH, 1884
Superfamily: ABELISAUROIDEA Novas, 1989
Family:
ABELISAURIDAE Bonaparte & Novas, 1985
*Tarascosaurus salluvicus
_____________________________________________________________________________________
FRANCE
BOUCHES-DU-RHONE DEPARTMENT
TUNNEL DE LA NERTHE
LACUSTRINE MARLS
EARLY ROGNACIAN = EARLY MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA
BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
Family:
RHABDODONTIDAE Weishampel, Jainu, Csiki & Norman,
2003
*Rhabdodon priscus
= Rhabdodon priscum
_____________________________________________________________________________________
FRANCE
BOUCHES-DU-RHONE DEPARTMENT
BASSIN OF LIGNITE DE FUVEAU
MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON,
1985
LEGION: ARCHOSAUROMORPHA
HUENE, 1946
SUBLEGION: DINOSAUROMORPHA
BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES
NOVAS, 1992
SUPERCOHORT: DINOSAURIA
OWEN, 1842
COHORT: ORNITHISCHIFORMES
COOPER, 1985
MAGNORDER: ORNITHISCHIA
SEELEY, 1888
SUPERORDER: NEORNITHISCHIA
COOPER, 1985
GRANDORDER:
ORNITHOPODA MARSH, 1871
MICRORDER:
EUORNITHOPODA SERENO, 1986
ORDER:
IGUANODONTIA DOLLO, 1888
Family:
RHABDODONTIDAE Weishampel, Jainu, Csiki & Norman,
2003
Rhabdodon
priscus
= Rhabdodon priscum
_____________________________________________________________________________________
FRANCE
BOUCHES-DU-RHONE DEPARTMENT
AIX BASIN
ROGNAC PROVINCE
MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON,
1985
LEGION: ARCHOSAUROMORPHA
HUENE, 1946
SUBLEGION: DINOSAUROMORPHA
BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES
NOVAS, 1992
SUPERCOHORT: DINOSAURIA
OWEN, 1842
COHORT: ORNITHISCHIFORMES
COOPER, 1985
SUPERORDER: SAUROPODOMORPHA
HUENE, 1932
GRANDORDER: BRONTOSAURIA
OLSHEVSKY, 1991 (emend)
MICRORDER:
SAUROPODA MARSH, 1878
ORDER: EUSAUROPODA
SUBORDER:
NEOSAUROPODA BONAPARTE, 1986
INFRAORDER:
MACRONARIA WILSON & SERENO, 1998
PARVORDER:
TITANOSAURIFORMES SALGADO ET AL, 1997
Family:
TITANOSAURIDAE Lydekker, 1885
Subfamily:
TITANOSAURINAE Lydekker, 1885 (sensu
Nopcsa, 1928)
*Hypselosaurus
priscus
_____________________________________________________________________________________
SOUTHEAST FRANCE
BOUCHES-DU-RHONE DEPARTMENT
AIX-EN-PROVINCE
AIX-EN-PROVINCE BASIN
LE NEUVE
LE NEUVE FOSSIL SITE
BEGUDIEN
UPPER SENONIEN
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?
ORDER: ALLOCAUDATA FOX & NAYLOR,
1982
Family: ALBANERPETONTIDAE
Fox & Naylor, 1982
Gen. sp indet.
SUPERORDER: BATRACHIA BRONGNIART, 1800
ORDER: CAUDATA SCOPOLI, 1777
Incertae sedis
Gen. sp indet.
SUPERORDER: SALIENTIA LAURENTI, 1768
Incertae sedis
Gen. sp indet.
ORDER: ANURA RAFINESQUE, 1815
SUBORDER: DISCOGLOSSOIDEI SOKOL, 1977
Family: DISCOGLOSSIDAE Guenther, 1859
Incertae sedis
Gen. sp indet.
SUBORDER: MESOBATRACHIA LAURENT, 1979
Superfamily: PIPOIDEA Gray, 1825 (sensu Fitzinger, 1843)
Family: PALAEOBATRACHIDAE Cope, 1865
Gen. sp indet.
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA
DUMERIL & BIBRON, 1839
SUBLEGION: LEPIDOSAURIA
HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA OPPEL, 1811
SUPERORDER: LACERTILIA OWEN, 1842
Incertae sedis
Gen. sp indet.
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI
SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI
BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA
WHETSTONE & WHYBROW, 1983
ORDER: EUSUCHIA emend HUXLEY, 1875
INFRAORDER: CROCODYLIA emend GMELIN, 1877
PARVORDER: BREVIROSTRES
Superfamily: ALLIGATORIDEA Buffetaut, 1985
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
FRANCE
BOUCHES-DU-RHONE DEPARTMENT
ROGNAC
BASE OF ROGNACIEN
MAASTRICHTIAN STAGE
UPPERMOST SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA
emend GAFFNEY & MEYLAN, 1988
MAGANORDER: CRYPTODIRA
GAFFNEY & MEYLAN, 1988
ORDER: CRYPTODIRA COPE, 1868
Family: SOLEMYDIDAE
Lapparent & Murelaga, 1997
*Solemys gaudryi
_____________________________________________________________________________________
FRANCE
BOUCHES-DU-RHONE DEPARTMENT
BUVEAU-VALDONNE BASIN
FUVELIAN
CAMPANIAN STAGE
SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA
emend GAFFNEY & MEYLAN, 1988
MAGNORDER: PLEURODIRA
GAFFNEY & MEYLAN, 1988
ORDER: PLEURODIRIDA COPE, 1864
SUBORDER: EUPLEURODIRA
GAFFNEY & MEYLAN, 1988
Superfamily: BOTHREMYDINURA Baur, 1891
Epifamily: BOTHERMYDINURA Baur, 1891
Subfamily: BOTHREMYDINAE Baur, 1891
Infrafamily: BOTHREMYDODDA Baur, 1891
Tribe: FOXEMYDINA Gaffney, Tong & Meylan, 2006
*Elochelys perfecta
Polysternon provinciale
_____________________________________________________________________________________
FRANCE
BOUCHES-DU-RHONE DEPARTMENT
COUDOUX
EARLY CAMPANIAN STAGE
SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGNORDER: PLEURODIRA GAFFNEY & MEYLAN, 1988
ORDER: PLEURODIRIDA COPE, 1864
SUBORDER: EUPLEURODIRA GAFFNEY & MEYLAN, 1988
Superfamily: PELOMEDUSOIDES Cope, 1868 Cope, 1868
Family: PELOMEDUSIDAE Cope, 1868
Subfamily: PELOMEDUSINAE Cope, 1868
Taphrosphys sp
_____________________________________________________________________________________
FRANCE
BOUCHES DU RHONE DEPARTMENT
SAINT-ESTEVE-JANSON
SABLES DU ROGNACIEN
LATE CAMPANIAN OR EARLY MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI
SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI
BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER:
MESOEUROCROCODYLIA WHETSTONE & WHYBROW,
1983
ORDER: MESOSUCHIA EMEND HUXLEY, 1875
PARVORDER:
METASUCHIA BENTON & CLARK, 1988
INFRAORDER: METAMESOSUCHIA HULKE, 1878
NEOSUCHIA
CLARK, IN BENTON & CLARK, 1988
Family: TREMATOCHAMPSIDAE Buffetaut, 1974
*Ischyrochampsa meridionalis
_____________________________________________________________________________________
FRANCE
BOUCHES-DU-RHONE DEPARTMENT
AQUITANIAN OF VENELLES
UPPER OLIGOCENE EPOCH
UPPER PALEOGENE PERIOD- LOWER MIOCENE EPOCH
LOWER NEOGENE PERIOD
UPPER TERTIARY SUBERA
MIDDLE CENOZOIC ERA
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI
SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI
BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER:
MESOEUROCROCODYLIA WHETSTONE & WHYBROW,
1983
ORDER: EUSUCHIA emend HUXLEY, 1875
INFRAORDER: CROCODYLIA emend GMELIN, 1877
PARVORDER: BREVIROSTRES
Superfamily: ALLIGATORIDEA Buffetaut, 1985
Family:
DIPLOCYNODONTIDAE emend Brouchu, 1999
Diplocynodon rateli
= Crocodilus ratelii
_____________________________________________________________________________________
FRANCE
BOUCHES-DU-RHONE DEPARTMENT
COMMUNE DE TRETS
LOWER ROGNACIAN
LATE MAASTRICHTIAN STAGE
UPPERMOST SENONIAN SUBEPOCH
UPPERMOST GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW,
1983
ORDER: EUSUCHIA emend HUXLEY, 1875
INFRAORDER: CROCODYLIA emend GMELIN, 1877
Incerate sedis
Eggshells
_____________________________________________________________________________________
FRANCE
BOUCHES-DU-RHONE DEPARTMENT
2 KM SE OF TRETS
LA BOUCHARDE
BEGUDIAN
LATE CAMPANIAN-EARLY MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA
BENTON & NORMAN IN BENTON,
1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA PAUL, 2002
AVEPECTORA PAUL, 2002
SUBORDER: MANIRAPTORA GAUTHIER, 1986
INFRAORDER: PARVES SERENO, 1997
PARVORDER:
DEINONYCHOSAURIA COLBERT & D.
A. RUSSELL, 1969
Family: DROMAEOSAURIDAE Russell, 1969
*Pyroraptor olympous
_____________________________________________________________________________________
FRANCE
BOUCHES-DU-RHONE DEPARTMENT
NEGION OF TRETS
LA NEUVE
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL,
1866
CLASS: REPTILIA LAURENTI,
1768
SUBCLASS: DIAPSIDA OSBORN,
1904
SUPERLEGION: NEODIAPSIDA
BENTON, 1985
LEGION: ARCHOSAUROMORPHA
HUENE, 1946
SUBLEGION: DINOSAUROMORPHA
BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES
NOVAS, 1992
SUPERCOHORT:
DINOSAURIA OWEN, 1842
COHORT: THEROPODA
EMEND MARSH, 1881
AVEPODA PAUL,
2002
SUPERORDER:
NEOTHEROPODA BAKKER, 1986
GRANDORDER:
TETANURAE GAUTHIER, 1986
MICRORDER:
COELUROSAURIA HUENE, 1914
ORDER:
MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA
PAUL, 2002
AVEPECTORA
PAUL, 2002
SUBORDER:
MANIRAPTORA GAUTHIER, 1986
INFRAORDER:
PARVES SERENO, 1997
PARVORDER:
DEINONYCHOSAURIA COLBERT & D.
A. RUSSELL, 1969
Family:
DROMAEOSAURIDAE Russell, 1969
Gen.
sp indet.
_____________________________________________________________________________________
FRANCE
BOUCHES-DU-RHONE DEPARTMENT
MARSEILLE
DIANIAN
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA
BENTON & NORMAN IN BENTON,
1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
SUPERORDER: SAUROPODOMORPHA HUENE, 1932
GRANDORDER: BRONTOSAURIA OLSHEVSKY, 1991 (emend)
MICRORDER: SAUROPODA MARSH, 1878
ORDER: EUSAUROPODA
SUBORDER: NEOSAUROPODA BONAPARTE, 1986
INFRAORDER:
MACRONARIA WILSON & SERENO, 1998
PARVORDER: TITANOSAURIFORMES SALGADO ET AL, 1997
Family: TITANOSAURIDAE Lydekker, 1885
Subfamily: TITANOSAURINAE Lydekker, 1885 (sensu Nopcsa, 1928)
Hypselosaurus priscus
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
Family:
RHABDODONTIDAE Weishampel, Jainu, Csiki & Norman,
2003
Rhabdodon priscus
= Rhabdodon priscum
SUBORDER: ANKYLOPOLLEXIA SERENO, 1986
PARVORDER: HADROSAURIFORMES SERENO, 1997
SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)
Family: HADROSAURIDAE COPE, 1869
Subfamily: HADROSAURINAE Cope, 1869 (sensu Lambe, 1918)
Telmatosaurus transsylvanicus
____________________________________________________________________________________
FRANCE
BOUCHES-DU-RHONE DEPARTMENT
MARSEILLE
PALAIS LONGCHAMP
DIANIAN
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL,
1866
CLASS: REPTILIA LAURENTI,
1768
SUBCLASS: DIAPSIDA OSBORN,
1904
SUPERLEGION: NEODIAPSIDA
BENTON, 1985
LEGION: ARCHOSAUROMORPHA
HUENE, 1946
SUBLEGION: DINOSAUROMORPHA
BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES
NOVAS, 1992
SUPERCOHORT:
DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES
COOPER, 1985
SUPERORDER:
SAUROPODOMORPHA HUENE, 1932
GRANDORDER:
BRONTOSAURIA OLSHEVSKY, 1991 (emend)
MICRORDER:
SAUROPODA MARSH, 1878
ORDER:
EUSAUROPODA
SUBORDER:
NEOSAUROPODA BONAPARTE, 1986
INFRAORDER:
MACRONARIA WILSON & SERENO, 1998
PARVORDER:
TITANOSAURIFORMES SALGADO ET AL, 1997
Family:
TITANOSAURIDAE Lydekker, 1885
Subfamily:
TITANOSAURINAE Lydekker, 1885 (sensu
Nopcsa, 1928)
Titanosaurus
indicu
_____________________________________________________________________________________
FRANCE
BOUCHES-DU-RHONE DEPARTMENT
BANKS OF ARC N OF TRETS
BEGUDIAN
LATE CRETACEOUS
STEM: TETRAPODA
GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON,
1985
LEGION: ARCHOSAUROMORPHA
HUENE, 1946
SUBLEGION: DINOSAUROMORPHA
BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES
NOVAS, 1992
SUPERCOHORT: DINOSAURIA
OWEN, 1842
COHORT: ORNITHISCHIFORMES
COOPER, 1985
SUPERORDER: SAUROPODOMORPHA
HUENE, 1932
GRANDORDER: BRONTOSAURIA
OLSHEVSKY, 1991 (emend)
MICRORDER:
SAUROPODA MARSH, 1878
ORDER: EUSAUROPODA
SUBORDER:
NEOSAUROPODA BONAPARTE, 1986
INFRAORDER:
MACRONARIA WILSON & SERENO, 1998
PARVORDER:
TITANOSAURIFORMES SALGADO ET AL, 1997
Family:
TITANOSAURIDAE Lydekker, 1885
Subfamily:
TITANOSAURINAE Lydekker, 1885 (sensu
Nopcsa, 1928)
Hypselosaurus
priscus
_____________________________________________________________________________________
FRANCE
BOUCHES-DU-RHONE DEPARTMENT
TRETS
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON,
1985
LEGION: ARCHOSAUROMORPHA
HUENE, 1946
SUBLEGION: DINOSAUROMORPHA
BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES
NOVAS, 1992
SUPERCOHORT: DINOSAURIA
OWEN, 1842
COHORT: ORNITHISCHIFORMES
COOPER, 1985
SUPERORDER: SAUROPODOMORPHA
HUENE, 1932
GRANDORDER: BRONTOSAURIA
OLSHEVSKY, 1991 (emend)
MICRORDER:
SAUROPODA MARSH, 1878
ORDER: EUSAUROPODA
SUBORDER:
NEOSAUROPODA BONAPARTE, 1986
INFRAORDER:
MACRONARIA WILSON & SERENO, 1998
PARVORDER:
TITANOSAURIFORMES SALGADO ET AL, 1997
Family:
TITANOSAURIDAE Lydekker, 1885
Subfamily:
TITANOSAURINAE Lydekker, 1885 (sensu
Nopcsa, 1928)
Hypselosaurus
priscus
_____________________________________________________________________________________
FRANCE
BOUCHES-DU-RHONE DEPARTMENT
COMMUNE DE BEURECUEIL
LOWER ROGNACIAN
LATE MAASTRICHTIAN STAGE
UPPERMOST SENONIAN SUBEPOCH
UPPERMOST GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL,
1866
CLASS: REPTILIA LAURENTI,
1768
SUBCLASS: DIAPSIDA OSBORN,
1904
SUPERLEGION: NEODIAPSIDA
BENTON, 1985
LEGION: ARCHOSAUROMORPHA
HUENE, 1946
SUPERCOHORT: CRUROTARSI
SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI
BENTON & CLARK, 1988
MAGNORDER: SUCHIA
KREBS, 1974
SUPERORDER:
CROCODYLOMORPHA WALKER, 1968
GRANDORDER:
CROCODILIFORMES HAY, 1930
MICRORDER:
MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
ORDER:
EUSUCHIA emend HUXLEY, 1875
INFRAORDER:
CROCODYLIA emend GMELIN, 1877
Incerate
sedis
Eggshells
_____________________________________________________________________________________
FRANCE
BOUCHES-DU-RHONE DEPARTMENT
AIX-EN-PROVINCE
LATE CRETACEOUS
VETEROVATA
OOSUPERCOHORT: DINOSAURIOVUM VIALOV, 1975
OOCOHORT: ORNITHISCHIFORMES NOVA
OOSUPERORDER: SAUROPODOMORPHA NOVA
oofamily: "MEGALOOLITHIDAE" Zhao, 1979 [Sauropoda]
Megaloolithus
_____________________________________________________________________________________
FRANCE
BOUCHES-DU-RHONE DEPARTMENT
AIX BASIN
LA NEUVE
LATE CRETACEOUS
VETEROVATA
oofamily: Indet.
Gen. sp indet.
OORDER: CROCODYLINES NOVA
oofamily: KROKOLITHIDAE Kohring & Hirsch, 1996
Gen. sp indet.
OOSUPERCOHORT: DINOSAURIOVUM VIALOV, 1975
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
FRANCE
BOUCHES-DU-RHONE DEPARTMENT
AIX BASIN
COMMUNE DE BEURECUEIL
LOWER ROGNACIAN
LATE MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
VETEROVATA
OORDER: CROCODYLINES NOVA
oofamily: KROKOLITHIDAE Kohring & Hirsch, 1996
Gen. sp indet.
_____________________________________________________________________________________
FRANCE
BOUCHES-DU-RHONE DEPARTMENT
AIX BASIN
VITROLLES-COUPERIGNE
EARLY ROGNACIAN = EARLY MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
VETEROVATA
OORDER: CROCODYLINES NOVA
oofamily: KROKOLITHIDAE Kohring & Hirsch, 1996
Gen. sp indet.
OOSUPERCOHORT: DINOSAURIOVUM VIALOV, 1975
oofamily: Incertae sedis
Gen. sp indet.
OOCOHORT: THEROPODA NOVA
oofamily: PRISMATOOLITHIDAE Hirsch, 1994
Prismatoolithus tenuis
Prismatoolithus sp
= cf. Prismatoolithus sp
OOCOHORT: ORNITHISCHIFORMES NOVA
OOSUPERORDER: SAUROPODOMORPHA NOVA
oofamily: "MEGALOOLITHIDAE" Zhao, 1979 [Sauropoda]
Megaloolithus aurelinesis
Megaloolithus sp
OOCLASS: AVES NOVA
oofamily: LAEVISOOLITHIDAE Mikhailov, 1991
Gen. sp indet.
Morphotype: Ornithoid-ratite
Ageroolithus fontllongensis
Ageroolithus sp
_____________________________________________________________________________________
FRANCE
BOUCHES-DU-RHONE DEPARTMENT
AIX BASIN
COMMUNE DE BEURECUEIL
LATE CRETACEOUS
VETEROVATA
OOSUPERCOHORT: DINOSAURIOVUM VIALOV, 1975
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
FRANCE
BOUCHES-DU-RHONE DEPARTMENT
AIX EN PROVINCE BASIN
LA CAIRANNE
LOWER ROGNACIEN
LATE MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
VETEROVATA
OOSUPERCOHORT: DINOSAURIOVUM VIALOV, 1975
OOCOHORT: THEROPODA NOVA
oofamily: ELONGATOOLITHIDAE Zhao, 1975
Gen. sp indet.
_____________________________________________________________________________________
FRANCE
BOUCHES-DU-RHONE DEPARTMENT
AIX-EN-PROVINCE PROVINCE
LA CAIRANNE
LOWER ROGNACIAN
LATE MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
VETEROVATA
OOSUPERCOHORT: DINOSAURIOVUM VIALOV, 1975
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
FRANCE
BOUCHES-DU-RHONE DEPARTMENT
AIX-EN-PROVINCE PROVINCE
EAST OF ROUSSET
LA CAIRANNE
LOWER ROGNACIAN
LATE MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
VETEROVATA
OOSUPERCOHORT: DINOSAURIOVUM VIALOV, 1975
OOCOHORT: ORNITHISCHIFORMES NOVA
OOSUPERORDER: SAUROPODOMORPHA NOVA
oofamily: "MEGALOOLITHIDAE" Zhao,
1979 [Sauropoda]
*Cairanoolithus dughii
*Megaloolithus microtuberculata
_____________________________________________________________________________________
FRANCE
BOUCHES-DU-RHONE DEPARTMENT
AIX-EN-PROVINCE PROVINCE
LA CAIRANNE
UNNAMED FORMATION
MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
VETEROVATA
OORDER: CROCODYLINES NOVA
oofamily: KROKOLITHIDAE Kohring & Hirsch, 1996
Gen. sp indet.
OOSUPERCOHORT: DINOSAURIOVUM VIALOV, 1975
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
FRANCE
BOUCHES-DU-RHONE DEPARTMENT
AIX-EN-PROVINCE
NEAR LA BEGUDE
ROUSSERT-ERBEN
LATE ROGNACIAN
LATE MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
VETEROVATA
OOSUPERCOHORT: DINOSAURIOVUM VIALOV, 1975
OOCOHORT: ORNITHISCHIFORMES NOVA
OOSUPERORDER: SAUROPODOMORPHA NOVA
oofamily: "MEGALOOLITHIDAE" Zhao,
1979 [Sauropoda]
*Megaloolithus mammilare
_____________________________________________________________________________________
FRANCE
BOUCHES-DU-RHONE DEPARTMENT
COUDOUX
FUVELIEN
LATE CAMPANIAN
LATE CRETACEOUS
VETEROVATA
OOSUPERCOHORT: DINOSAURIOVUM VIALOV, 1975
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
FRANCE
BOUCHES-DU-RHONE DEPARTMENT
AIX-EN-PROVINCE
LES BREGUIERES
LATE MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
VETEROVATA
OOSUPERCOHORT: DINOSAURIOVUM VIALOV, 1975
OOCOHORT: ORNITHISCHIFORMES NOVA
OOSUPERORDER: SAUROPODOMORPHA NOVA
oofamily: "MEGALOOLITHIDAE" Zhao,
1979 [Sauropoda]
Megaloolithus mammilare
Megaloolithus baghensis
=* Megaloolithus pseudomamillare
= Megaloolithus cf. mammilare
_____________________________________________________________________________________
FRANCE
BOUCHES-DU-RHONE DEPARTMENT
AIX-EN-PROVINCE
LE NEUVE
LE NEUVE FOSSIL SITE
BEGUDIEN
UPPER SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
VETEROVATA
Eggs with gekoid basic shell type (Gekkota)
Gen. sp indet.
OOSUPERCOHORT: DINOSAURIOVUM VIALOV, 1975
OOCOHORT: THEROPODA NOVA
oofamily: PRISMATOOLITHIDAE Hirsch, 1994
Gen. sp indet.
OOCLASS: AVES NOVA
oofamily: LAEVISOOLITHIDAE Mikhailov, 1991
Gen. sp indet.
_____________________________________________________________________________________
FRANCE
BOUCHES-DU-RHONE DEPARTMENT
AIX BASIN
LA CARDELINE
LATE MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
VETEROVATA
OOSUPERCOHORT: DINOSAURIOVUM VIALOV, 1975
OOCOHORT: ORNITHISCHIFORMES NOVA
OOSUPERORDER: SAUROPODOMORPHA NOVA
oofamily: "MEGALOOLITHIDAE" Zhao,
1979 [Sauropoda]
Megaloolithus mammilare
_____________________________________________________________________________________
FRANCE
BOUCHES-DU-RHONE DEPARTMENT
LA BEGUDE
LATE CRETACEOUS
VETEROVATA
OOSUPERCOHORT: DINOSAURIOVUM VIALOV, 1975
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
FRANCE
BOUCHES-DU-RHONE DEPARTMENT
LA BEGUDE
“MIDDLE” ROGNACIAN
MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
VETEROVATA
OOSUPERCOHORT: DINOSAURIOVUM VIALOV, 1975
OOCOHORT: ORNITHISCHIFORMES NOVA
OOSUPERORDER: SAUROPODOMORPHA NOVA
oofamily: "MEGALOOLITHIDAE" Zhao,
1979 [Sauropoda]
*Megaloolithus siruguei
_____________________________________________________________________________________
FRANCE
BOUCHES-DU-RHONE DEPARTMENT
CHATEAUNEUF-LE-ROUGE
LATE CRETACEOUS
VETEROVATA
OOSUPERCOHORT: DINOSAURIOVUM VIALOV, 1975
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
FRANCE
BOUCHES-DU-RHONE DEPARTMENT
COUDOUX
FUVELIAN
EARLY CAMPANIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
VETEROVATA
OOSUPERCOHORT: DINOSAURIOVUM VIALOV, 1975
OOCOHORT: THEROPODA NOVA
oofamily: PRISMATOOLITHIDAE Hirsch, 1994
Prismatoolithus tenuis
_____________________________________________________________________________________
FRANCE
BOUCHES-DU-RHONE DEPARTMENT
LA BASTIDE BLANCHE
FUVELIAN
EARLY CAMPANIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
VETEROVATA
OOSUPERCOHORT: DINOSAURIOVUM VIALOV, 1975
OOCOHORT: THEROPODA NOVA
oofamily: PRISMATOOLITHIDAE Hirsch, 1994
Prismatoolithus tenuis
_____________________________________________________________________________________
FRANCE
BOUCHES-DU-RHONE DEPARTMENT
MARSEILLE
DIANIAN
LATE CRETACEOUS
VETEROVATA
OOSUPERCOHORT: DINOSAURIOVUM VIALOV, 1975
OOCOHORT: ORNITHISCHIFORMES NOVA
OOSUPERORDER: SAUROPODOMORPHA NOVA
oofamily: "MEGALOOLITHIDAE" Zhao, 1979 [Sauropoda]
Gen. sp indet.
Hypselosaurus priscus?
_____________________________________________________________________________________
FRANCE
BOUCHES-DU-RHONE DEPARTMENT
LES PENNES MIRABEAU
PUYLOUBIER
ROGNAICAN
EARLY LATE MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
VETEROVATA
OOSUPERCOHORT: DINOSAURIOVUM VIALOV, 1975
OOCOHORT: THEROPODA NOVA
oofamily: PRISMATOOLITHIDAE Hirsch, 1994
Prismatoolithus tenuis
_____________________________________________________________________________________
FRANCE
BOUCHES-DU-RHONE
ROQUES-HAUTES
LATE CRETACEOUS
VETEROVATA
OOSUPERCOHORT: DINOSAURIOVUM VIALOV, 1975
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
FRANCE
BOUCHES-DU-RHONE DEPARTMENT
N OF ROQUEHAUTE FARM
LATE CRETACEOUS
VETEROVATA
OOSUPERCOHORT: DINOSAURIOVUM VIALOV, 1975
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
FRANCE
BOUCHES-DU-RHONE DEPARTMENT
ROUSSET-SUR-ARC
LATE CRETACEOUS
VETEROVATA
OOSUPERCOHORT: DINOSAURIOVUM VIALOV, 1975
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
FRANCE
BOUCHES-DU-RHONE DEPARTMENT
ROUSSET-ROUTIERS
LATE CRETACEOUS
VETEROVATA
OOSUPERCOHORT: DINOSAURIOVUM VIALOV, 1975
OOCOHORT: ORNITHISCHIFORMES NOVA
OOSUPERORDER: SAUROPODOMORPHA NOVA
oofamily: "MEGALOOLITHIDAE" Zhao,
1979 [Sauropoda]
Megaloolithus siruguei
_____________________________________________________________________________________
FRANCE
BOUCHES-DU-RHONE DEPARTMENT
ROUSSET VILLAGE (RSV)
LATE CRETACEOUS
VETEROVATA
OOSUPERCOHORT: DINOSAURIOVUM VIALOV, 1975
OOCOHORT: ORNITHISCHIFORMES NOVA
OOSUPERORDER: SAUROPODOMORPHA NOVA
oofamily: "MEGALOOLITHIDAE" Zhao,
1979 [Sauropoda]
Megaloolithus siruguei
_____________________________________________________________________________________
FRANCE
BOUCHES-DU-RHONE DEPARTMENT
ROUSSET-VILLAGE
“MIDDLE” ROGNAC
LATE CRETACEOUS
VETEROVATA
OOSUPERCOHORT: DINOSAURIOVUM VIALOV, 1975
OOCOHORT: ORNITHISCHIFORMES NOVA
OOSUPERORDER: SAUROPODOMORPHA NOVA
oofamily: "MEGALOOLITHIDAE" Zhao,
1979 [Sauropoda]
Dughioolithus roussetensis
_____________________________________________________________________________________
FRANCE
BOUCHES-DU-RHONE DEPARTMENT
ROUSSET
UNNAMED FORMATION
MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
VETEROVATA
OOSUPERCOHORT: DINOSAURIOVUM VIALOV, 1975
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
FRANCE
BOUCHES-DU-RHONE DEPARTMENT
ROUSSET
UPPER ROGNACIAN
LATE MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
VETEROVATA
OOSUPERCOHORT: DINOSAURIOVUM VIALOV, 1975
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
FRANCE
BOUCHES-DU-RHONE DEPARTMENT
ROUSSET ERBEN AND A LA CARDELINE
LATE CRETACEOUS
VETEROVATA
OOSUPERCOHORT: DINOSAURIOVUM VIALOV, 1975
OOCOHORT: ORNITHISCHIFORMES NOVA
OOSUPERORDER: SAUROPODOMORPHA NOVA
oofamily: "MEGALOOLITHIDAE" Zhao,
1979 [Sauropoda]
Megaloolithus siruguei
_____________________________________________________________________________________
FRANCE
BOUCHES-DU-RHONE DEPARTMENT
LE ROUSSETT
ROGNACIEN SUPERIOR
LATE CRETACEOUS
VETEROVATA
OOSUPERCOHORT: DINOSAURIOVUM VIALOV, 1975
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
FRANCE
BOUCHES-DU-RHONE DEPARTMENT
LES GRANDES CREUX
LOWER ROGNACIEN
LATE MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
VETEROVATA
OOSUPERCOHORT: DINOSAURIOVUM VIALOV, 1975
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
FRANCE
BOUCHES-DU-RHONE DEPARTMENT
SAINT-SER
UPPER ROGNACIEN
LATE MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
VETEROVATA
OOSUPERCOHORT: DINOSAURIOVUM VIALOV, 1975
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
FRANCE
BOUCHES-DU-RHONE DEPARTMENT
PENNES-MIRABEAU
LOWER ROGNACIEN
LATE MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
VETEROVATA
OOSUPERCOHORT: DINOSAURIOVUM VIALOV, 1975
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
FRANCE
BOUCHES-DU-RHONE DEPARTMENT
PUYLOUBIER
LOWER ROGNACIEN
LATE MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
VETEROVATA
OOSUPERCOHORT: DINOSAURIOVUM VIALOV, 1975
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
FRANCE
BOUCHES-DU-RHONE DEPARTMENT
AIX BASIN
COMMUNE DE TRETS
LATE CRETACEOUS
VETEROVATA
OORDER: CROCODYLINES NOVA
oofamily: KROKOLITHIDAE Kohring & Hirsch, 1996
Gen. sp indet.
_____________________________________________________________________________________
FRANCE
BOUCHES-DU-RHONE DEPARTMENT
TRETS
ROGNACIAN
EARLY LATE MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
VETEROVATA
OOSUPERCOHORT: DINOSAURIOVUM VIALOV, 1975
Incertae sedis
Gen. sp indet.
OOCOHORT: THEROPODA NOVA
oofamily: PRISMATOOLITHIDAE Hirsch, 1994
Prismatoolithus tenuis
_____________________________________________________________________________________
FRANCE
BOUCHES-DU-RHONE DEPARTMENT
NEAR TRETS
CLOS-LA-NEUVE
BEGUDIAN (= CAMPANIAN)
LATE CRETACEOUS
VETEROVATA
OOSUPERCOHORT: DINOSAURIOVUM VIALOV, 1975
OOCOHORT: ORNITHISCHIFORMES NOVA
OOSUPERORDER: SAUROPODOMORPHA NOVA
oofamily: "MEGALOOLITHIDAE" Zhao,
1979 [Sauropoda]
*Megaloolithus aurelinesis
_____________________________________________________________________________________
FRANCE
BOUCHES-DU-RHONE DEPARTMENT
PYULOBIER
LATE CRETACEOUS
VETEROVATA
OOSUPERCOHORT: DINOSAURIOVUM VIALOV, 1975
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
FRANCE
BOUCHES-DU-RHONE DEPARTMENT
MASSIF D'ALLAUCH
ESCAOUPRES VALLEY
NEOCOMIAN TRIOLOGY
N2B FORMATIOIN
LATE VALANGINIAN STAGE
MIDDLE NEOCOMIAN
SUBEPOCH
MIDDLE EARLY CRETACEOUS EPOCH
EARLY
CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL,
1866
CLASS: REPTILIA LAURENTI,
1768
SUBCLASS: DIAPSIDA OSBORN,
1904
INFRACLASS: NEODIAPSIDA
BENTON, 1985
LEGION: ARCHOSAUROMORPHA
HUENE, 1946
SUPERCOHORT: CRUROTARSI
SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI
BENTON & CLARK, 1988
MAGNORDER: SUCHIA
KREBS, 1974
SUPERORDER:
CROCODYLOMORPHA WALKER, 1968
GRANDORDER:
CROCODILIFORMES HAY, 1930
MICRORDER:
MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
ORDER:
MESOSUCHIA EMEND HUXLEY, 1875
SUBORDER:
THALATTOSUCHIA EMEND FRAAS, 1901
Family:
METRIORHYNCHIDAE Fitzinger, 1843
Subfamily:
GEOSAURINAE Lydekker, 1889
Tribe: PLESIOSUCHINA Young, de Andrade, Cornee, Steel & Foffa, 2014
Gen. sp indet.
_____________________________________________________________________________________