SPAIN
NE HUESCA PROVINCE
NORTH OF THE PYRENEES NEAR THE VILLAGE OF SERRADUY DEL PON
AMOR 3
AREN FORMATION
TREMP GROUP
LATE CAMPANIAN-MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER UPPER GULF EPOC
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI
SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI
BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA
WHETSTONE & WHYBROW, 1983
ORDER: EUSUCHIA emend HUXLEY, 1875
ORDER: EUSUCHIA emend HUXLEY, 1875
Family: ALLODAPOSUCHIDAE Narvaez, Brochu, Escaso, Perez-Garcia, & Ortega,
2015
Allodaposuchus
sp
_____________________________________________________________________________________
SPAIN
HUESCA PROVINCE
15 KM WEST OF THE CLASSIC SITES OF AREN
AMOR 3
CONQUES FORMATION
TREMP GROUP
MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER UPPER GULF EPOC
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL,
1866
CLASS: REPTILIA LAURENTI,
1768
SUBCLASS: DIAPSIDA OSBORN,
1904
SUPERLEGION: NEODIAPSIDA
BENTON, 1985
LEGION: ARCHOSAUROMORPHA
HUENE, 1946
SUBLEGION: DINOSAUROMORPHA
BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES
NOVAS, 1992
SUPERCOHORT:
DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES
COOPER, 1985
MAGNORDER:
ORNITHISCHIA SEELEY, 1888
SUPERORDER:
NEORNITHISCHIA COOPER, 1985
GRANDORDER:
ORNITHOPODA MARSH, 1871
MICRORDER:
EUORNITHOPODA SERENO, 1986
ORDER:
IGUANODONTIA DOLLO, 1888
SUBORDER:
ANKYLOPOLLEXIA SERENO, 1986
PARVORDER:
HADROSAURIFORMES SERENO, 1997
SUPERFAMILY:
HADROSAUROIDEA COPE, 1869 (SENSU SERENO,
1986)
Family:
HADROSAURIDAE COPE, 1869
Incertae
sedis
Gen.
sp indet.
_____________________________________________________________________________________
SPAIN
HUESCA PROVINCE
15 KM WEST OF THE CLASSIC SITES OF AREN
BARRANCO SERRADUY 4
CONQUES FORMATION
TREMP GROUP
MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER UPPER GULF EPOC
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL,
1866
CLASS: REPTILIA LAURENTI,
1768
SUBCLASS: DIAPSIDA OSBORN,
1904
SUPERLEGION: NEODIAPSIDA
BENTON, 1985
LEGION: ARCHOSAUROMORPHA
HUENE, 1946
SUBLEGION: DINOSAUROMORPHA
BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES
NOVAS, 1992
SUPERCOHORT:
DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES
COOPER, 1985
MAGNORDER:
ORNITHISCHIA SEELEY, 1888
SUPERORDER:
NEORNITHISCHIA COOPER, 1985
GRANDORDER:
ORNITHOPODA MARSH, 1871
MICRORDER:
EUORNITHOPODA SERENO, 1986
ORDER:
IGUANODONTIA DOLLO, 1888
SUBORDER:
ANKYLOPOLLEXIA SERENO, 1986
PARVORDER:
HADROSAURIFORMES SERENO, 1997
SUPERFAMILY:
HADROSAUROIDEA COPE, 1869 (SENSU SERENO,
1986)
Family:
HADROSAURIDAE COPE, 1869
Incertae
sedis
Gen.
sp indet
EUHADROSAURIA WEISHAMPEL, NORMAN & GRIGORESCU, 1993
Incertae
sedis
Gen.
sp indet
_____________________________________________________________________________________
SPAIN
HUESCA PROVINCE
15 KM WEST OF THE CLASSIC SITES OF AREN
COLOR
CONQUES FORMATION
TREMP GROUP
MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER UPPER GULF EPOC
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL,
1866
CLASS: REPTILIA LAURENTI,
1768
SUBCLASS: DIAPSIDA OSBORN,
1904
SUPERLEGION: NEODIAPSIDA
BENTON, 1985
LEGION: ARCHOSAUROMORPHA
HUENE, 1946
SUBLEGION: DINOSAUROMORPHA
BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES
NOVAS, 1992
SUPERCOHORT:
DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES
COOPER, 1985
MAGNORDER:
ORNITHISCHIA SEELEY, 1888
SUPERORDER:
NEORNITHISCHIA COOPER, 1985
GRANDORDER:
ORNITHOPODA MARSH, 1871
MICRORDER:
EUORNITHOPODA SERENO, 1986
ORDER:
IGUANODONTIA DOLLO, 1888
SUBORDER:
ANKYLOPOLLEXIA SERENO, 1986
PARVORDER:
HADROSAURIFORMES SERENO, 1997
SUPERFAMILY:
HADROSAUROIDEA COPE, 1869 (SENSU SERENO,
1986)
Family:
HADROSAURIDAE COPE, 1869
Incertae
sedis
Gen.
sp indet.
_____________________________________________________________________________________
SPAIN
HUESCA PROVINCE
15 KM WEST OF THE CLASSIC SITES OF AREN
DOLOR 3
CONQUES FORMATION
TREMP GROUP
MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER UPPER GULF EPOC
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL,
1866
CLASS: REPTILIA LAURENTI,
1768
SUBCLASS: DIAPSIDA OSBORN,
1904
SUPERLEGION: NEODIAPSIDA
BENTON, 1985
LEGION: ARCHOSAUROMORPHA
HUENE, 1946
SUBLEGION: DINOSAUROMORPHA
BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES
NOVAS, 1992
SUPERCOHORT:
DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES
COOPER, 1985
MAGNORDER:
ORNITHISCHIA SEELEY, 1888
SUPERORDER:
NEORNITHISCHIA COOPER, 1985
GRANDORDER:
ORNITHOPODA MARSH, 1871
MICRORDER:
EUORNITHOPODA SERENO, 1986
ORDER:
IGUANODONTIA DOLLO, 1888
SUBORDER:
ANKYLOPOLLEXIA SERENO, 1986
PARVORDER:
HADROSAURIFORMES SERENO, 1997
SUPERFAMILY:
HADROSAUROIDEA COPE, 1869 (SENSU SERENO,
1986)
Family:
HADROSAURIDAE COPE, 1869
Incertae
sedis
Gen.
sp indet.
_____________________________________________________________________________________
NORTH-EAST SPAIN
SOUTHERN PYRENEES
HUESCA PROVINCE
MUNICIPALITY OF BERANUY
'DOLOR' FOSSIL SITE
MIDDLE PART OF THE 'LOWER RED GARUMNIAN'
UPPER MAASTRICHTIAN STAGE
UPPERMOST SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH,
1930
SUPERCLASS: AMNIOTA HAECKEL,
1866
CLASS: AVES LINNAEUS, 1758
SUBCLASS: CARINATE MERREM, 1813
INFRACLASS: ORNITHUROMORPHA CHIAPPE, 2002
Incerate
sedis
Gen. sp indet.
_____________________________________________________________________________________
NORTH-EAST SPAIN
HUESCA PROVINCE
NEAR THE VILLAGE OF SERRADUY DEL POIN
MUNICIPALITY OF BERANUY
CONQUES FORMATION
TREMP GROUP
UPPER MAASTRICHTIAN STAGE
UPPERMOST SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI
SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI
BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA
WHETSTONE & WHYBROW, 1983
ORDER: EUSUCHIA emend HUXLEY, 1875
ORDER: EUSUCHIA emend HUXLEY, 1875
Family: ALLODAPOSUCHIDAE Narvaez, Brochu, Escaso, Perez-Garcia, & Ortega,
2015
*Allodaposuchus
subjuniperus
= Agaresuchus subjuniperus
_____________________________________________________________________________________
SPAIN
NE HUESCA PROVINCE
THE CENTRAL-SOUTHERN PYRENEES
NEAR THE VILLAGES OF SERRADUY DEL POIN AND BERANUY
TREMP FORMATION
MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL,
1866
CLASS: REPTILIA LAURENTI,
1768
SUBCLASS: DIAPSIDA OSBORN,
1904
SUPERLEGION: NEODIAPSIDA
BENTON, 1985
LEGION: ARCHOSAUROMORPHA
HUENE, 1946
SUBLEGION: DINOSAUROMORPHA
BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES
NOVAS, 1992
SUPERCOHORT:
DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES
COOPER, 1985
MAGNORDER:
ORNITHISCHIA SEELEY, 1888
SUPERORDER:
NEORNITHISCHIA COOPER, 1985
GRANDORDER:
ORNITHOPODA MARSH, 1871
MICRORDER:
EUORNITHOPODA SERENO, 1986
ORDER:
IGUANODONTIA DOLLO, 1888
SUBORDER:
ANKYLOPOLLEXIA SERENO, 1986
PARVORDER:
HADROSAURIFORMES SERENO, 1997
SUPERFAMILY:
HADROSAUROIDEA COPE, 1869 (SENSU SERENO,
1986)
Family:
HADROSAURIDAE COPE, 1869
Incertae
sedis
Gen.
sp indet.
_____________________________________________________________________________________
SPAIN
HUESCA PROVINCE
ESTADA
MUSCHELKALK
MID TRIASSIC EPOCH
MIDDLE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: EURYAPSIDA COLBERT, 1945
SUPERORDER: SAUROPTERYGIA OWEN 1860
GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994
ORDER: NOTHOSAURIFORMES STORRS, 1991
SUBORDER: EUSAUROPTERYGIA TSCHANZ, 1989
INFRAORDER: NOTHOSAUROIDEA BAUR, 1889
Superfamily: NOTHOSAURIA Baur, 1889
Family: NOTHOSAURIDAE Baur, 1889
Lariosaurus balsami
Lariosaurus valceresii
_____________________________________________________________________________________
SPAIN
HUESCA PROVINCE
EOCENE EPOCH
MIDDLE PALEOGENE PERIOD
TERTIARY SUBERA
CENOZOIC ERA
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI
SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI
BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA
WHETSTONE & WHYBROW, 1983
ORDER: EUSUCHIA emend HUXLEY, 1875
INFRAORDER: CROCODYLIA emend GMELIN, 1877
PARVORDER: BREVIROSTRES
Superfamily: CROCODILOIDEA Cuvier, 1807 (sensu Fitzinger, 1826)
Family: PRISTICHAMPSIDAE Efimov, 1988
Pristichampsus rollainti
= Crocodilus bolcensis
_____________________________________________________________________________________
SPAIN
HUESCA PROVINCE
SERRADUY
TREMP FORMATION
MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA
BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
SUPERORDER: SAUROPODOMORPHA HUENE, 1932
GRANDORDER: BRONTOSAURIA OLSHEVSKY, 1991 (emend)
MICRORDER: SAUROPODA MARSH, 1878
ORDER: EUSAUROPODA
SUBORDER: NEOSAUROPODA BONAPARTE, 1986
INFRAORDER:
MACRONARIA WILSON & SERENO, 1998
PARVORDER: TITANOSAURIFORMES SALGADO ET AL, 1997
Family: TITANOSAURIDAE Lydekker, 1885
Subfamily: TITANOSAURINAE Lydekker, 1885 (sensu Nopcsa, 1928)
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
SPAIN
HUESCA PROVINCE
FIGUEROLA 2
TREMP FORMATION
EARLY MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA
BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA PAUL, 2002
AVEPECTORA PAUL, 2002
SUBORDER: MANIRAPTORA GAUTHIER, 1986
INFRAORDER: PARVES SERENO, 1997
PARVORDER:
DEINONYCHOSAURIA COLBERT & D. A.
RUSSELL, 1969
Family: DROMAEOSAURIDAE Russell, 1969
Incertae sedis
Morphotype 1
Gen. sp indet.
Morphotype 2
Gen. sp indet.
_____________________________________________________________________________________
SPAIN
HUESCA PROVINCE
FONTLLONGA 6
TREMP FORMATION
EARLY MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
LEGION: ARCHOSAUROMORPHA HUENE,
1946
SUBLEGION: DINOSAUROMORPHA
BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES
NOVAS, 1992
SUPERCOHORT: DINOSAURIA
OWEN, 1842
COHORT: THEROPODA EMEND
MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA
BAKKER, 1986
GRANDORDER: TETANURAE
GAUTHIER, 1986
MICRORDER:
COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES
HOLTZ, 1996
AVETHEROPODA
PAUL, 2002
AVEPECTORA
PAUL, 2002
SUBORDER:
MANIRAPTORA GAUTHIER, 1986
INFRAORDER:
PARVES SERENO, 1997
PARVORDER:
DEINONYCHOSAURIA COLBERT & D. A.
RUSSELL, 1969
Family:
DROMAEOSAURIDAE Russell, 1969
Incertae
sedis
Morphotype
1
Gen.
sp indet.
Morphotype
2
Gen.
sp indet.
_____________________________________________________________________________________
SPAIN
HUESCA PROVINCE
AREN
MONTREBEI
TREMP FORMATION
LATE CAMPANIAN STAGE-EARLY MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL,
1866
CLASS: REPTILIA LAURENTI,
1768
SUBCLASS: DIAPSIDA OSBORN,
1904
LEGION: ARCHOSAUROMORPHA
HUENE, 1946
SUBLEGION: DINOSAUROMORPHA
BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES
NOVAS, 1992
SUPERCOHORT: DINOSAURIA
OWEN, 1842
COHORT: THEROPODA
EMEND MARSH, 1881
Incertae sedis
Gen. sp indet.
AVEPODA PAUL,
2002
SUPERORDER:
NEOTHEROPODA BAKKER, 1986
GRANDORDER:
TETANURAE GAUTHIER, 1986
MICRORDER:
COELUROSAURIA HUENE, 1914
Incertae
sedis
Gen.
sp indet.
ORDER:
MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA
PAUL, 2002
AVEPECTORA
PAUL, 2002
SUBORDER:
MANIRAPTORA GAUTHIER, 1986
INFRAORDER:
PARVES SERENO, 1997
PARVORDER:
DEINONYCHOSAURIA COLBERT & D.
A. RUSSELL, 1969
Family:
DROMAEOSAURIDAE Russell, 1969
Incertae
sedis
Morphotype
3
Gen.
sp indet.
Morphotype
4
Gen.
sp indet.
_____________________________________________________________________________________
SPAIN
HUESCA PROVINCE
AREN
VICARI 4
TREMP FORMATION
LATE CAMPANIAN STAGE
SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL,
1866
CLASS: REPTILIA LAURENTI,
1768
SUBCLASS: DIAPSIDA OSBORN,
1904
LEGION: ARCHOSAUROMORPHA
HUENE, 1946
SUBLEGION: DINOSAUROMORPHA
BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES
NOVAS, 1992
SUPERCOHORT: DINOSAURIA
OWEN, 1842
COHORT: THEROPODA
EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER:
NEOTHEROPODA BAKKER, 1986
GRANDORDER:
TETANURAE GAUTHIER, 1986
MICRORDER:
COELUROSAURIA HUENE, 1914
Incertae
sedis
Gen.
sp indet.
ORDER:
MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA
PAUL, 2002
AVEPECTORA
PAUL, 2002
SUBORDER:
MANIRAPTORA GAUTHIER, 1986
INFRAORDER:
PARVES SERENO, 1997
PARVORDER:
DEINONYCHOSAURIA COLBERT & D. A.
RUSSELL, 1969
Family:
DROMAEOSAURIDAE Russell, 1969
Incertae
sedis
Morphotype
4
Gen.
sp indet.
_____________________________________________________________________________________
SPAIN
HUESCA PROVINCE
SOUTH-CENTRAL PYRENEES
AREN
BLASI 1
AREN SANDSTONE
UPPER TREMP FORMATION
LATE MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI,
1768
SUBCLASS: DIAPSIDA OSBORN,
1904
SUPERLEGION: NEODIAPSIDA
BENTON, 1985
LEGION: ARCHOSAUROMORPHA
HUENE, 1946
SUPERCOHORT: CRUROTARSI
SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI
BENTON & CLARK, 1988
MAGNORDER: SUCHIA
KREBS, 1974
SUPERORDER:
CROCODYLOMORPHA WALKER, 1968
GRANDORDER:
CROCODILIFORMES HAY, 1930
MICRORDER:
MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
ORDER:
EUSUCHIA emend HUXLEY, 1875
INFRAORDER:
CROCODYLIA emend GMELIN, 1877
PARVORDER:
BREVIROSTRES
Superfamily:
ALLIGATORIDEA Buffetaut, 1985
Incertae
sedis
Gen.
sp indet.
SUBLEGION: DINOSAUROMORPHA
BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES
NOVAS, 1992
SUPERCOHORT:
DINOSAURIA OWEN, 1842
COHORT: THEROPODA
EMEND MARSH, 1881
Incertae
sedis
Gen. sp
indet.
COHORT: ORNITHISCHIFORMES
COOPER, 1985
MAGNORDER:
ORNITHISCHIA SEELEY, 1888
SUPERORDER:
NEORNITHISCHIA COOPER, 1985
GRANDORDER:
ORNITHOPODA MARSH, 1871
MICRORDER:
EUORNITHOPODA SERENO, 1986
ORDER:
IGUANODONTIA DOLLO, 1888
SUBORDER:
ANKYLOPOLLEXIA SERENO, 1986
PARVORDER:
HADROSAURIFORMES SERENO, 1997
SUPERFAMILY:
HADROSAUROIDEA COPE, 1869 (SENSU SERENO,
1986)
Family:
HADROSAURIDAE COPE, 1869
Subfamily:
LAMBEOSAURINAE Parks, 1923
Tribe:
LAMBEOSAURINI Parks, 1923
Incertae
sedis
Gen.
sp indet.
Tribe: ARENYSAURINI Longrich, Pereda Suberbiola, Pyron & Jalil, 2019
*Blasisaurus
canudoi
_____________________________________________________________________________________
SPAIN
HUESCA PROVINCE
SOUTH-CENTRAL PYRENEES
AREN
BLASI 1B
AREN SANDSTONE
UPPER TREMP FORMATION
LATE MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL,
1866
CLASS: REPTILIA LAURENTI,
1768
SUBCLASS: DIAPSIDA OSBORN,
1904
SUPERLEGION: NEODIAPSIDA
BENTON, 1985
LEGION: ARCHOSAUROMORPHA
HUENE, 1946
SUBLEGION: DINOSAUROMORPHA
BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES
NOVAS, 1992
SUPERCOHORT:
DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES
COOPER, 1985
MAGNORDER:
ORNITHISCHIA SEELEY, 1888
SUPERORDER:
NEORNITHISCHIA COOPER, 1985
GRANDORDER:
ORNITHOPODA MARSH, 1871
MICRORDER:
EUORNITHOPODA SERENO, 1986
ORDER:
IGUANODONTIA DOLLO, 1888
SUBORDER:
ANKYLOPOLLEXIA SERENO, 1986
PARVORDER:
HADROSAURIFORMES SERENO, 1997
SUPERFAMILY:
HADROSAUROIDEA COPE, 1869 (SENSU SERENO,
1986)
Family:
HADROSAURIDAE COPE, 1869
Incertae
sedis
Gen.
sp indet.
_____________________________________________________________________________________
SPAIN
HUESCA PROVINCE
SOUTH-CENTRAL PYRENEES
AREN
BLASI 2
AREN SANDSTONE
ARENISCA FORMATION (UPPER TREMP FORMATION)
LATE MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL,
1866
CLASS: AMPHIBIA LINNAEUS,
1758
Incertae sedis
Gen. sp indet.
SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?
ORDER: ALLOCAUDATA FOX & NAYLOR,
1982
Family: ALBANERPETONTIDAE
Fox & Naylor,
1982
Albanerpeton aff. A. nexuosus
SUPERORDER: SALIENTIA LAURENTI, 1768
ORDER: ANURA RAFINESQUE, 1815
SUBORDER: DISCOGLOSSOIDEI SOKOL, 1977
Family: DISCOGLOSSIDAE Guenther, 1859
aff. Paradiscoglossus sp
SUBORDER: MESOBATRACHIA LAURENT, 1979
Superfamily: PIPOIDEA Gray, 1825 (sensu
Fitzinger, 1843)
Family: PALAEOBATRACHIDAE Cope, 1865
Gen. sp indet.
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA
KUHN, 1960
SUPERCOHORT: TESTUDINES
LINNAEUS, 1758
COHORT: CASICHELYDIA
emend GAFFNEY & MEYLAN, 1988
MAGNORDER: PLEURODIRA
GAFFNEY & MEYLAN, 1988
ORDER: PLEURODIRIDA
COPE, 1864
SUBORDER: EUPLEURODIRA
GAFFNEY & MEYLAN, 1988
Superfamily:
PELOMEDUSOIDES Cope, 1868 Cope, 1868
Family: PELOMEDUSIDAE
Cope, 1868
Subfamily:
PELOMEDUSINAE Cope, 1868
Cf. Polysternon sp
Gen.
sp indet.
SUBCLASS: DIAPSIDA OSBORN,
1904
INFRACLASS: NEODIAPSIDA
BENTON, 1985
LEGION: LEPIDOSAUROMORPHA
DUMERIL & BIBRON, 1839
SUBLEGION: LEPIDOSAURIA
HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA
OPPEL, 1811
SUPERORDER: LACERTILIA
OWEN, 1842
Incertae sedis
Gen. sp indet.
Iguania or Scleroglossa
Indertminate lizard 1
Scleroglossa, Scincomorpha
Indertminate lizard 2
GRANDORDER:
ASCALABOTA GILMORE, 1928
ORDER: IGUANIA
LATREILLE, 1825
SUBORDER:
DRACOMORPHA NORTHCUTT, 1978
INFRAORDER:
IGUANOMORPHA FURBRINGER, 1900
Family:
IGUANIDAE Gray, 1827
Gen.
sp indet.
GRANDORDER: SCINCOGEKKONOMORPHA
SUKHANOV, 1961
ORDER: ANGUIMORPHA
FURBRINGER, 1900
SUBORDER: DIPLOGLOSSA
COPE, 1864
INFRAORDER:
OPHISAUROMORPHA ALIFANOV, 2000
Family: ANGUIDAE
Gray, 1825
Gen. sp
indet.
SUPERORDER: SERPENTES
LINNAEUS, 1759
ORDER: OPHIDIA BRONGNIART,
1800
SUBORDER: ALETHINOPHIDIA
NOPCSCA, 1923
Incertae sedis
Gen. sp indet.
LEGION: ARCHOSAUROMORPHA
HUENE, 1946
SUPERCOHORT: CRUROTARSI
SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI
BENTON & CLARK, 1988
MAGNORDER: SUCHIA
KREBS, 1974
SUPERORDER:
CROCODYLOMORPHA WALKER, 1968
GRANDORDER:
CROCODILIFORMES HAY, 1930
Incertae
sedis
Gen.
sp indet.
MICRORDER:
MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
ORDER:
MESOSUCHIA EMEND HUXLEY, 1875
PARVORDER:
METASUCHIA BENTON & CLARK, 1988
INFRAORDER:
METAMESOSUCHIA HULKE, 1878
NEOSUCHIA
CLARK, IN BENTON & CLARK, 1988
Family:
TREMATOCHAMPSIDAE Buffetaut, 1974
Incertae
sedis
Gen.
sp indet.
ORDER:
EUSUCHIA emend HUXLEY, 1875
INFRAORDER:
CROCODYLIA emend GMELIN, 1877
Arenysuchus
gascabadiolorum
PARVORDER:
BREVIROSTRES
Superfamily:
ALLIGATORIDEA Buffetaut, 1985
Incertae
sedis
Gen.
sp indet.
GLOBIDONTA
BROUCHU, 1999
Family:
ALLIGATORIDAE Gray, 1844
Acynodon sp
SUBLEGION: DINOSAUROMORPHA
BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES
NOVAS, 1992
SUPERCOHORT:
DINOSAURIA OWEN, 1842
COHORT: THEROPODA
EMEND MARSH, 1881
Incertae
sedis
?Rhicharodoestesia
sp
Gen. sp
indet.
AVEPODA PAUL,
2002
SUPERORDER:
NEOTHEROPODA BAKKER, 1986
GRANDORDER:
TETANURAE GAUTHIER, 1986
MICRORDER:
COELUROSAURIA HUENE, 1914
Gen.
sp indet.
ORDER:
MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA
PAUL, 2002
AVEPECTORA
PAUL, 2002
SUBORDER:
MANIRAPTORA GAUTHIER, 1986
INFRAORDER:
PARVES SERENO, 1997
PARVORDER:
DEINONYCHOSAURIA COLBERT & D.
A. RUSSELL, 1969
Family:
DROMAEOSAURIDAE Russell, 1969
?Paronychodon sp = Cf. Euronychodon sp
Gen.
sp indet.
Morphotype
4
Gen.
sp indet.
Morphotype
6
Gen.
sp indet.
Morphotype
7
Gen.
sp indet.
Morphotype
8
Gen.
sp indet.
Subfamily:
DROMAEOSAURINAE Matthew & Brown,
1922
Gen.
sp indet.
Subfamily:
VELOCIRAPTORINAE Barsbold,
1978
Gen.
sp indet.
COHORT: ORNITHISCHIFORMES
COOPER, 1985
SUPERORDER:
SAUROPODOMORPHA HUENE, 1932
GRANDORDER:
BRONTOSAURIA OLSHEVSKY, 1991 (emend)
MICRORDER:
SAUROPODA MARSH, 1878
Incertae
sedis
Gen.
sp indet.
MAGNORDER:
ORNITHISCHIA SEELEY, 1888
SUPERORDER:
NEORNITHISCHIA COOPER, 1985
GRANDORDER:
ORNITHOPODA MARSH, 1871
MICRORDER:
EUORNITHOPODA SERENO, 1986
ORDER:
IGUANODONTIA DOLLO, 1888
SUBORDER:
ANKYLOPOLLEXIA SERENO, 1986
PARVORDER:
HADROSAURIFORMES SERENO, 1997
SUPERFAMILY:
HADROSAUROIDEA COPE, 1869 (SENSU SERENO,
1986)
Family:
HADROSAURIDAE COPE, 1869
Incertae
sedis
Gen.
sp indet.
Note: Euhadrosaurian.
VETEROVATA
OOSUPERCOHORT: DINOSAURIOVUM
VIALOV, 1975
Incertae sedis
Gen. sp indet.
OOCOHORT: THEROPODA NOVA
Incertae sedis
Gen. sp indet.
OOCOHORT: ORNITHISCHIFORMES
NOVA
OOSUPERORDER: SAUROPODOMORPHA
NOVA
oofamily: "MEGALOOLITHIDAE" Zhao,
1979 [Sauropoda]
Gen. sp indet.
_____________________________________________________________________________________
SPAIN
HUESCA PROVINCE
SOUTH-CENTRAL PYRENEES
AREN
BLASI 3
AREN SANDSTONE
UPPER TREMP FORMATION
LATE MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL,
1866
CLASS: REPTILIA LAURENTI,
1768
SUBCLASS: CHELONOMORPHA
KUHN, 1960
SUPERCOHORT: TESTUDINES
LINNAEUS, 1758
COHORT: CASICHELYDIA
emend GAFFNEY & MEYLAN, 1988
MAGNORDER: PLEURODIRA
GAFFNEY & MEYLAN, 1988
ORDER: PLEURODIRIDA
COPE, 1864
SUBORDER: EUPLEURODIRA
GAFFNEY & MEYLAN, 1988
Superfamily:
PELOMEDUSOIDES Cope, 1868 Cope, 1868
Family: PELOMEDUSIDAE
Cope, 1868
Subfamily:
PELOMEDUSINAE Cope, 1868
Gen.
sp indet.
Cf. Polysternon sp
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA
BENTON, 1985
LEGION: ARCHOSAUROMORPHA
HUENE, 1946
SUPERCOHORT: CRUROTARSI
SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI
BENTON & CLARK, 1988
MAGNORDER: SUCHIA
KREBS, 1974
SUPERORDER:
CROCODYLOMORPHA WALKER, 1968
GRANDORDER:
CROCODILIFORMES HAY, 1930
MICRORDER:
MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
ORDER:
MESOSUCHIA EMEND HUXLEY, 1875
PARVORDER:
METASUCHIA BENTON & CLARK, 1988
INFRAORDER:
METAMESOSUCHIA HULKE, 1878
NEOSUCHIA
CLARK, IN BENTON & CLARK, 1988
Family:
TREMATOCHAMPSIDAE Buffetaut, 1974
Incertae
sedis
Gen.
sp indet.
ORDER:
EUSUCHIA emend HUXLEY, 1875
Incertae
sedis
Gen.
sp indet.
INFRAORDER:
CROCODYLIA emend GMELIN, 1877
PARVORDER:
BREVIROSTRES
Superfamily:
ALLIGATORIDEA Buffetaut, 1985
Incertae
sedis
Gen.
sp indet.
SUBLEGION: DINOSAUROMORPHA
BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES
NOVAS, 1992
SUPERCOHORT:
DINOSAURIA OWEN, 1842
COHORT: THEROPODA
EMEND MARSH, 1881
Incertae
sedis
Gen. sp
indet.
AVEPODA PAUL,
2002
SUPERORDER:
NEOTHEROPODA BAKKER, 1986
GRANDORDER:
TETANURAE GAUTHIER, 1986
MICRORDER:
COELUROSAURIA HUENE, 1914
ORDER:
MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA
PAUL, 2002
AVEPECTORA
PAUL, 2002
SUBORDER:
MANIRAPTORA GAUTHIER, 1986
INFRAORDER:
PARVES SERENO, 1997
PARVORDER:
DEINONYCHOSAURIA COLBERT & D.
A. RUSSELL, 1969
Family:
DROMAEOSAURIDAE Russell, 1969
Morphotype
5
Gen.
sp indet.
Subfamily:
VELOCIRAPTORINAE Barsbold,
1978
Gen.
sp indet.
COHORT: ORNITHISCHIFORMES
COOPER, 1985
MAGNORDER:
ORNITHISCHIA SEELEY, 1888
SUPERORDER:
NEORNITHISCHIA COOPER, 1985
GRANDORDER:
ORNITHOPODA MARSH, 1871
MICRORDER:
EUORNITHOPODA SERENO, 1986
ORDER:
IGUANODONTIA DOLLO, 1888
SUBORDER:
ANKYLOPOLLEXIA SERENO, 1986
PARVORDER:
HADROSAURIFORMES SERENO, 1997
SUPERFAMILY:
HADROSAUROIDEA COPE, 1869 (SENSU SERENO,
1986)
Family:
HADROSAURIDAE COPE, 1869
Incertae
sedis
Gen.
sp indet.
Subfamily:
LAMBEOSAURINAE Parks, 1923
Genus
nova
Gen.
sp indet.
Tribe: ARENYSAURINI Longrich, Pereda Suberbiola, Pyron & Jalil, 2019
*Arenysaurus
ardevoli
_____________________________________________________________________________________
SPAIN
HUESCA PROVINCE
SOUTH-CENTRAL PYRENEES
AREN
BLASI 3
CONQUES FORMATION
TREMP GROUP
LATE MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL,
1866
CLASS: REPTILIA LAURENTI,
1768
SUBCLASS: DIAPSIDA OSBORN,
1904
SUPERLEGION: NEODIAPSIDA
BENTON, 1985
LEGION: ARCHOSAUROMORPHA
HUENE, 1946
SUBLEGION: DINOSAUROMORPHA
BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES
NOVAS, 1992
SUPERCOHORT:
DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES
COOPER, 1985
MAGNORDER:
ORNITHISCHIA SEELEY, 1888
SUPERORDER:
NEORNITHISCHIA COOPER, 1985
GRANDORDER:
ORNITHOPODA MARSH, 1871
MICRORDER:
EUORNITHOPODA SERENO, 1986
ORDER:
IGUANODONTIA DOLLO, 1888
SUBORDER:
ANKYLOPOLLEXIA SERENO, 1986
PARVORDER:
HADROSAURIFORMES SERENO, 1997
SUPERFAMILY:
HADROSAUROIDEA COPE, 1869 (SENSU SERENO,
1986)
Family:
HADROSAURIDAE COPE, 1869
Incertae
sedis
Gen.
sp indet.
Subfamily:
LAMBEOSAURINAE Parks, 1923
Incertae
sedis
Gen.
sp indet.
_____________________________________________________________________________________
SPAIN
HUESCA PROVINCE
SOUTH-CENTRAL PYRENEES
AREN
BLASI 3.5
AREN SANDSTONE
UPPER TREMP FORMATION
LATE MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL,
1866
CLASS: REPTILIA LAURENTI,
1768
SUBCLASS: CHELONOMORPHA
KUHN, 1960
SUPERCOHORT: TESTUDINES
LINNAEUS, 1758
COHORT: CASICHELYDIA
emend GAFFNEY & MEYLAN, 1988
MAGNORDER: PLEURODIRA
GAFFNEY & MEYLAN, 1988
ORDER: PLEURODIRIDA
COPE, 1864
SUBORDER: EUPLEURODIRA
GAFFNEY & MEYLAN, 1988
Superfamily:
PELOMEDUSOIDES Cope, 1868 Cope, 1868
Family: PELOMEDUSIDAE
Cope, 1868
Subfamily:
PELOMEDUSINAE Cope, 1868
Gen.
sp indet.
_____________________________________________________________________________________
SPAIN
HUESCA PROVINCE
SOUTH-CENTRAL PYRENEES
AREN
BLASI 4
AREN SANDSTONE
UPPER TREMP FORMATION
LATE MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL,
1866
CLASS: REPTILIA LAURENTI,
1768
SUBCLASS: CHELONOMORPHA
KUHN, 1960
SUPERCOHORT: TESTUDINES
LINNAEUS, 1758
COHORT: CASICHELYDIA
emend GAFFNEY & MEYLAN, 1988
MAGNORDER: PLEURODIRA
GAFFNEY & MEYLAN, 1988
ORDER: PLEURODIRIDA
COPE, 1864
SUBORDER: EUPLEURODIRA
GAFFNEY & MEYLAN, 1988
Superfamily:
PELOMEDUSOIDES Cope, 1868 Cope, 1868
Family: PELOMEDUSIDAE
Cope, 1868
Subfamily:
PELOMEDUSINAE Cope, 1868
Cf. Polysternon sp
Gen.
sp indet.
SUBCLASS: DIAPSIDA OSBORN,
1904
SUPERLEGION: NEODIAPSIDA
BENTON, 1985
LEGION: ARCHOSAUROMORPHA
HUENE, 1946
SUBLEGION: DINOSAUROMORPHA
BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES
NOVAS, 1992
SUPERCOHORT: DINOSAURIA
OWEN, 1842
COHORT: ORNITHISCHIFORMES
COOPER, 1985
MAGNORDER:
ORNITHISCHIA SEELEY, 1888
SUPERORDER:
NEORNITHISCHIA COOPER, 1985
GRANDORDER:
ORNITHOPODA MARSH, 1871
MICRORDER:
EUORNITHOPODA SERENO, 1986
ORDER:
IGUANODONTIA DOLLO, 1888
SUBORDER:
ANKYLOPOLLEXIA SERENO, 1986
PARVORDER:
HADROSAURIFORMES SERENO, 1997
SUPERFAMILY:
HADROSAUROIDEA COPE, 1869 (SENSU SERENO,
1986)
Family:
HADROSAURIDAE COPE, 1869
Incertae
sedis
Gen.
sp indet.
Subfamily:
LAMBEOSAURINAE Parks, 1923
Incertae
sedis
Gen.
sp indet.
_____________________________________________________________________________________
SPAIN
HUESCA PROVINCE
SOUTH-CENTRAL PYRENEES
AREN
BLASI 5
AREN SANDSTONE
UPPER TREMP FORMATION
LATE MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL,
1866
CLASS: REPTILIA LAURENTI,
1768
SUBCLASS: CHELONOMORPHA
KUHN, 1960
SUPERCOHORT: TESTUDINES
LINNAEUS, 1758
COHORT: CASICHELYDIA
emend GAFFNEY & MEYLAN, 1988
MAGNORDER: PLEURODIRA
GAFFNEY & MEYLAN, 1988
ORDER: PLEURODIRIDA
COPE, 1864
SUBORDER: EUPLEURODIRA
GAFFNEY & MEYLAN, 1988
Superfamily:
PELOMEDUSOIDES Cope, 1868 Cope, 1868
Family: PELOMEDUSIDAE
Cope, 1868
Subfamily:
PELOMEDUSINAE Cope, 1868
Cf. Polysternon sp
Gen.
sp indet.
SUBCLASS: DIAPSIDA OSBORN,
1904
SUPERLEGION: NEODIAPSIDA
BENTON, 1985
LEGION: ARCHOSAUROMORPHA
HUENE, 1946
SUPERCOHORT: CRUROTARSI
SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI
BENTON & CLARK, 1988
MAGNORDER: SUCHIA
KREBS, 1974
SUPERORDER:
CROCODYLOMORPHA WALKER, 1968
GRANDORDER:
CROCODILIFORMES HAY, 1930
Incertae
sedis
Gen.
sp indet.
SUBLEGION: DINOSAUROMORPHA
BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES
NOVAS, 1992
SUPERCOHORT:
DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES
COOPER, 1985
MAGNORDER:
ORNITHISCHIA SEELEY, 1888
SUPERORDER:
NEORNITHISCHIA COOPER, 1985
GRANDORDER:
ORNITHOPODA MARSH, 1871
MICRORDER:
EUORNITHOPODA SERENO, 1986
ORDER:
IGUANODONTIA DOLLO, 1888
SUBORDER:
ANKYLOPOLLEXIA SERENO, 1986
PARVORDER:
HADROSAURIFORMES SERENO, 1997
SUPERFAMILY:
HADROSAUROIDEA COPE, 1869 (SENSU SERENO,
1986)
Family:
HADROSAURIDAE COPE, 1869
Incertae
sedis
Gen.
sp indet.
_____________________________________________________________________________________
SPAIN
HUESCA PROVINCE
SOUTH-CENTRAL PYRENEES
AREN
BLA(R)
AREN SANDSTONE
UPPER TREMP FORMATION
LATE MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL,
1866
CLASS: REPTILIA LAURENTI,
1768
SUBCLASS: CHELONOMORPHA
KUHN, 1960
SUPERCOHORT: TESTUDINES
LINNAEUS, 1758
COHORT: CASICHELYDIA
emend GAFFNEY & MEYLAN, 1988
MAGNORDER: PLEURODIRA
GAFFNEY & MEYLAN, 1988
ORDER: PLEURODIRIDA
COPE, 1864
SUBORDER: EUPLEURODIRA
GAFFNEY & MEYLAN, 1988
Superfamily:
PELOMEDUSOIDES Cope, 1868 Cope, 1868
Family: PELOMEDUSIDAE
Cope, 1868
Subfamily:
PELOMEDUSINAE Cope, 1868
Gen. sp indet.
_____________________________________________________________________________________
SPAIN
HUESCA PROVINCE
SOUTH-CENTRAL PYRENEES
AREN
RIM2
AREN SANDSTONE
UPPER TREMP FORMATION
LATE MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL,
1866
CLASS: REPTILIA LAURENTI,
1768
SUBCLASS: CHELONOMORPHA
KUHN, 1960
SUPERCOHORT: TESTUDINES
LINNAEUS, 1758
COHORT: CASICHELYDIA
emend GAFFNEY & MEYLAN, 1988
MAGNORDER: PLEURODIRA
GAFFNEY & MEYLAN, 1988
ORDER: PLEURODIRIDA
COPE, 1864
SUBORDER: EUPLEURODIRA
GAFFNEY & MEYLAN, 1988
Superfamily:
PELOMEDUSOIDES Cope, 1868 Cope, 1868
Family: PELOMEDUSIDAE
Cope, 1868
Subfamily:
PELOMEDUSINAE Cope, 1868
Gen. sp indet.
_____________________________________________________________________________________
SPAIN
HUESCA PROVINCE
SOUTH-CENTRAL PYRENEES
AREN
RIM 2 (R)
AREN SANDSTONE
UPPER TREMP FORMATION
LATE MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL,
1866
CLASS: REPTILIA LAURENTI,
1768
SUBCLASS: CHELONOMORPHA
KUHN, 1960
SUPERCOHORT: TESTUDINES
LINNAEUS, 1758
COHORT: CASICHELYDIA
emend GAFFNEY & MEYLAN, 1988
MAGNORDER: PLEURODIRA
GAFFNEY & MEYLAN, 1988
ORDER: PLEURODIRIDA
COPE, 1864
SUBORDER: EUPLEURODIRA
GAFFNEY & MEYLAN, 1988
Superfamily:
PELOMEDUSOIDES Cope, 1868 Cope, 1868
Family: PELOMEDUSIDAE
Cope, 1868
Subfamily:
PELOMEDUSINAE Cope, 1868
Gen. sp indet.
_____________________________________________________________________________________
SPAIN
HUESCA PROVINCE
SOUTH-CENTRAL PYRENEES
AREN
BLA2
AREN SANDSTONE
UPPER TREMP FORMATION
LATE MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL,
1866
CLASS: REPTILIA LAURENTI,
1768
SUBCLASS: CHELONOMORPHA
KUHN, 1960
SUPERCOHORT: TESTUDINES
LINNAEUS, 1758
COHORT: CASICHELYDIA
EMEND GAFFNEY & MEYLAN, 1988
MAGANORDER: CRYPTODIRA
GAFFNEY & MEYLAN, 1988
ORDER: CRYPTODIRA
COPE, 1868
Family: SOLEMYDIDAE
Lapparent & Murelaga, 1997
Gen. sp indet.
_____________________________________________________________________________________
SPAIN
HUESCA PROVINCE
SOUTH-CENTRAL PYRENNES
ARENY
AREN FORMATION
MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
ICHNOSTEM: TETRAPODIPIDA NOVA
ICHNOSUPERCLASS: AMNIOTAIPIDII NOVA
ICHNOCLASS: REPTILIPEDIA VIALOV,
1966
ICHNOSUBCLASS: DIAPSIDIPIDII
NOVA
ICHNOSTEM-GROUP: NEODIAPSIDIPEDII
NOVA
ICHNOINFRACLASS: NEODIAPSIDIPEDII
NOVA
ICHNOSUBDIVISION: ARCHOSAURIPEDIDII
NOVA
ICHNOLEGION: DINOSAUROMORPHIPEDA
NOVA
ICHNOSUBLEGION: DINOSAURIFORMEIPEDIDII
NOVA
ICHNOSUPERCOHORT:
DINOSAUROPEDIDA NOVA
ICHNOCOHORT: ORNITHISCHIFORMEIPIDA
NOVA
ICHNOMAGNORDER:
ORNITHISCHIPIDA NOVA
ICHNOSUPERORDER:
NEORNITHISCHIPIDA NOVA
ICHNOGRANDORDER:
ORNITHOPODIPDIA NOVA
ICHNOMICRORDER:
EUORNITHOPODIPIDA NOVA
ICHNOORDER:
IGUANODONTIPIDIA NOVA
ICHNOSUBORDER:
ANKYLOPOLLEXIPEDIDA NOVA
ICHNOINFRAORDER:
STYRACOSTERNIPIDA NOVA
ICHNOPARVORDER:
HADROSAURIFORMIPEDIDA NOVA
ICHNOSUPERFAMILY:
HADROSAUROIDIPEDIA NOVA
Ichnofamily:
HADROSAURIOPDIDAE Vylov, 1988
Gen.
sp indet.
_____________________________________________________________________________________
NORTH SPAIN
AREN (HUESCA) PROVINCE
ELIAS SITE
ELI-1
UNIT 2
TREMP FORMATION
LATE MAASTRICHTIAN STAGE
UPPERMOST SENONIAN SUBEPOCH
UPPERMOST GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON,
1985
LEGION: ARCHOSAUROMORPHA
HUENE, 1946
SUPERCOHORT: CRUROTARSI
SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI
BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS,
1974
SUPERORDER: CROCODYLOMORPHA
WALKER, 1968
GRANDORDER: CROCODILIFORMES
HAY, 1930
MICRORDER: MESOEUROCROCODYLIA
WHETSTONE & WHYBROW, 1983
ORDER: EUSUCHIA
emend HUXLEY, 1875
INFRAORDER:
CROCODYLIA emend GMELIN, 1877
PARVORDER:
BREVIROSTRES
Superfamily:
CROCODILOIDEA Cuvier, 1807 (sensu Fitzinger,
1826)
*Arenysuchus
gascabadiolorum
_____________________________________________________________________________________
SPAIN
HUESCA PROVINCE
AREN
MONTREBEI
TREMP FORMATION
LATE CAMPANIAN STAGE-EARLY MAASTRICHTIAN
STAGE
SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL,
1866
CLASS: REPTILIA LAURENTI,
1768
SUBCLASS: DIAPSIDA OSBORN,
1904
SUPERLEGION: NEODIAPSIDA
BENTON, 1985
LEGION: ARCHOSAUROMORPHA
HUENE, 1946
SUBLEGION: DINOSAUROMORPHA
BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES
NOVAS, 1992
SUPERCOHORT:
DINOSAURIA OWEN, 1842
COHORT: THEROPODA
EMEND MARSH, 1881
Incertae
sedis
?Rhicharodoestesia sp