EAST DEVON COUNTY
PEAK HILL
PENNINGTON POINT MEMBER
HELSBY SANDSTONE FORMATION (FORMER OTTER SANDSTONE FORMATION)
ANISIAN STAGE
LOWER MID TRIASSIC EPOCH
MIDDLE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
Incertae sedis
*Feralisaurus corami
_____________________________________________________________________________________
ENGLANDEAST DEVON COUNTY
ABOUT 3 KM WEST OF THE CLIFFS NEAR PEAK HILL
20 M WEST OF THE SID RIVER OUTFALL
PENNINGTON POINT
OTTER SANDSTONE FORMATION
ANISIAN STAGE
LOWER MID TRIASSIC EPOCH
MIDDLE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: RHYNOCEPHALIA nova
ORDER: RHYNOCEPHALIA BUTHER, 1867
SUBORDER: RHYNCHOSAUROIDEA GERVAIS, 1889
Family: HYPERODAPEDONTIDAE Lydekker, 1885
*Bentonyx sidensis
_____________________________________________________________________________________
SOUTHERN UNITED KINGDOM
ENGLAND
DEVON COUNTY
JUST OVER THE BRIDGE THAT CROSSES THE RIVER SID AT THE EAST END OF TOWN
PORT ROYAL
PENNINGTON POINT
SIDMOUTH
OTTER SANDSTONE FORMATION
ANISIAN STAGE
LOWER MID TRIASSIC EPOCH
MIDDLE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
Incertae sedis
Gen. sp indet.
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
MAGNORDER: PROTEROSUCHIFORMES nova
Gen. sp indet.
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: PSEUDOSUCHIA ZITTEL 1887-1890
GRANDORDER: RAUISUCHIFORMES PARRISH, 1993
ORDER: RAUISUCHIA Bonaparte, 1984
SUBORDER: RAUISUCHIA BONAPARTE, 1982
Family: POPOSAURIDAE Nopcsa, 1928
Gen. sp indet.
_____________________________________________________________________________________
DEVON COUNTY
OTTER SANDSTONE FORMATION
ANISIAN STAGE
LOWER MID TRIASSIC EPOCH
MIDDLE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
_____________________________________________________________________________________SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
MAGNORDER: PROTOROSAURIA HUXLEY, 1871
SUPERORDER: TAPINOPLATIA PETERS EMEND NOVA
GRANDORDER: CHARACIOPODA PETERS EMEND NOVA
Family: TANYSTROPHEIDAE Geravis, 1858
Tanystropheus sp
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
SUPERORDER: ORNITHOSUCHIMORPHA NOVA
ORDER: ORNITHOSUCHIA BONAPARTE 1971
Family: ORNITHOSUCHIDAE Huene, 1908
Ornithosuchus longidens
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: PSEUDOSUCHIA ZITTEL 1887-1890
Incertae sedis
Gen. sp indet.
GRANDORDER: RAUISUCHIFORMES PARRISH, 1993
ORDER: RAUISUCHIA Bonaparte, 1984
SUBORDER: RAUISUCHIA BONAPARTE, 1982
Family: CTENOSAURISCHIDAE, Kuhn, 1964
Ctenosauriscus sp
ENGLAND
LEFT BANK OF THE OTTER RIVER
DEVON COUNTY
“NEW RED”
LATE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
_____________________________________________________________________________________SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: RHYNOCEPHALIA nova
ORDER: RHYNOCEPHALIA BUTHER, 1867
SUBORDER: RHYNCHOSAUROIDEA GERVAIS, 1889
Family: HYPERODAPEDONTIDAE Lydekker, 1885
Subfamily: HYPERODAPEDONTINAE Lydekker, 1885 (sensu Chatterjee, 1969)
Hyperodapedon sp
ENGLAND
DEVON COUNTY
OTTER RIVER
WARWICK
LOWER KEUPER
LATE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: RHYNOCEPHALIA nova
ORDER: RHYNOCEPHALIA BUTHER, 1867
SUBORDER: RHYNCHOSAUROIDEA GERVAIS, 1889
Family: HYPERODAPEDONTIDAE Lydekker, 1885
Subfamily: HYPERODAPEDONTINAE Lydekker, 1885 (sensu Chatterjee, 1969)
_____________________________________________________________________________________Hyperodapedon gordoni
ENGLAND
DEVON COUNTY
SIDMOUTH
CLIFFS OF PICKET ROCK COVE
OTTER SANDSTONE
ANISIAN STAGE
LOWER MID TRIASSIC EPOCH
MIDDLE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
_____________________________________________________________________________________SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LABYRINTHODONTIA OWEN, 1860
SUPERORDER: TEMNOSPONDYLIA ZITTEL, 1887
MICROORDER: LIMNARCHIA YATES & WARREN, 2000
ORDER: CAPITOSAURIA ROMER, 1947
Superfamily: CAPITOSAUROIDEA Watson, 1919 (sensu Romer, 1947)
Incertae sedis
Non* Mastodonsaurus lavisi
= Labyrinthodon lavisi
ENGLAND
DEVON COUNTY
SIDMOUTH
BUDLEIGH SALTERTON
HIGH PEAK
MIDDLE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
_____________________________________________________________________________________SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: RHYNOCEPHALIA nova
ORDER: RHYNOCEPHALIA BUTHER, 1867
SUBORDER: RHYNCHOSAUROIDEA GERVAIS, 1889
Family: RHYNCHOSAURIDAE Huxley, 1887
Rhynchosaurus spenceri
ENGLAND
EAST DEVON COUNTY
BUDLEIGH SALTERTON
OTTERTON POINT
OTTER SANDSTONE FORMATION
ANISIAN STAGE
LOWER MID TRIASSIC EPOCH
MIDDLE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
_____________________________________________________________________________________SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: RHYNOCEPHALIA nova
ORDER: RHYNOCEPHALIA BUTHER, 1867
SUBORDER: RHYNCHOSAUROIDEA GERVAIS, 1889
Family: RHYNCHOSAURIDAE Huxley, 1887
*Rhynchosaurus spenceri
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: PSEUDOSUCHIA ZITTEL 1887-1890
GRANDORDER: RAUISUCHIFORMES PARRISH, 1993
ORDER: RAUISUCHIA Bonaparte, 1984
SUBORDER: RAUISUCHIA BONAPARTE, 1982
Family: CTENOSAURISCHIDAE, Kuhn, 1964
Arizonasaurus-like
ENGLAND
DEVON COUNTY
SIDMOUTH
SE COAST OF DEVON FROM OTTERTON POINT NEAR BUDLEIGH SALTERTON
OTTER SANDSTONE FORMATION
ANISIAN STAGE
LOWER MID TRIASSIC EPOCH
MIDDLE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
_____________________________________________________________________________________SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LABYRINTHODONTIA OWEN, 1860
SUPERORDER: TEMNOSPONDYLIA ZITTEL, 1887
MICROORDER: LIMNARCHIA YATES & WARREN, 2000
ORDER: CAPITOSAURIA ROMER, 1947
Superfamily: CAPITOSAUROIDEA Watson, 1919 (sensu Romer, 1947)
Incertae sedis
Non Mastodonsaurus lavisi
= Labyrinthodon lavisiGen. sp indet.
Family: CYCLOTOSAURIDAE Shiskin, 1964
Subfamily: HEYLEROSAURINAE Shishkin, 1980
Eocyclotosaurus sp
ENGLAND
DEVON COUNTY
WEST OF SIDMOUTH
BELOW HIGHT PEAN
OTTER SANDSTONE FORMATION
ANISIAN STAGE
LOWER MID TRIASSIC EPOCH
MIDDLE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
_____________________________________________________________________________________SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LABYRINTHODONTIA OWEN, 1860
SUPERORDER: TEMNOSPONDYLIA ZITTEL, 1887
MICROORDER: LIMNARCHIA YATES & WARREN, 2000
ORDER: CAPITOSAURIA ROMER, 1947
Superfamily: CAPITOSAUROIDEA Watson, 1919 (sensu Romer, 1947)
Family: CYCLOTOSAURIDAE Shiskin, 1964
Subfamily: HEYLEROSAURINAE Shishkin, 1980
Eocyclotosaurus sp
ENGLAND
EAST DEVON COUNTY
SIDMOUTH
WESTERN END OF SALCOMBE HILL CLIFF
OTTER SANDSTONE FORMATION
SHERWOOD SANDSTONE GROUP
ANISIAN STAGE
LOWER MIDDLE TRIASSIC EPOCH
LATE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
_____________________________________________________________________________________SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: NEODIAPSIDA BENTON, 1985
DIVISION: LEPIDOSAUROMORPHA BENTON, 1983
Incertae sedis
*Coartaredens isaaci
ENGLAND
EAST DEVON COUNTY
SIDMOUTH
LADE FOOT
GREEN POINT
OTTER SANDSTONE FORMATION
SHERWOOD SANDSTONE GROUP
ANISIAN STAGE
LOWER MIDDLE TRIASSIC EPOCH
LATE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: PARAREPTILIA OLSON 1947
Incertae sedis
Gen sp indet.
_____________________________________________________________________________________SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: RHYNOCEPHALIA nova
ORDER: RHYNOCEPHALIA BUTHER, 1867
SUBORDER: RHYNCHOSAUROIDEA GERVAIS, 1889
Family: RHYNCHOSAURIDAE Huxley, 1887
Rhynchosaurus spenceri
ENGLAND
EAST DEVON COUNTY
SIDMOUTH
LADE FOOT
OTTER SANDSTONE FORMATION
SHERWOOD SANDSTONE GROUP
ANISIAN STAGE
LOWER MIDDLE TRIASSIC EPOCH
LATE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: PARAREPTILIA OLSON 1947
INFRACLASS: SAUROPSIDA HUXLEY, 1864
ORDER: PROCOLOPHONOMORPHA ROMER, 1964
SUBORDER: PROCOLPHOMORPHA SEELEY, 1888
Family: PROCOLOPHONIDAE, Seeley, 1888
Subfamily: PROCOLOPHONINAE Seeley, 1888 (sensu Romer, 1956)
Kapes bentoni
_____________________________________________________________________________________
SOUTHERN UNITED KINGDOM
ENGLAND
EAST DEVON COUNTY
SIDMOUTH
APPROXIMATELY 1.4 KM WEST-SOUTH-WEST OF THE BEACH ACCESS AT JACOB'S LADDER
BENEATH PEAK HILL
PENNINGTON POINT MEMBER
OTTER SANDSTONE
SHERWOOD SANDSTONE GROUP
HELSBY SANDSTONE FORMATION
ANISIAN STAGE
LOWER MIDDLE TRIASSICSTEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: PARAREPTILIA OLSON 1947
INFRACLASS: SAUROPSIDA HUXLEY, 1864
ORDER: PROCOLOPHONOMORPHA ROMER, 1964
SUBORDER: PROCOLPHOMORPHA SEELEY, 1888
Family: PROCOLOPHONIDAE, Seeley, 1888
Subfamily: PROCOLOPHONINAE Seeley, 1888 (sensu Romer, 1956)
Kapes bentoni
_____________________________________________________________________________________
ENGLAND
EAST DEVON COUNTY
SIDMOUTH
WESTERN END OF SALCOMBE HILL CLIFF
OTTER SANDSTONE FORMATION
SHERWOOD SANDSTONE GROUP
ANISIAN STAGE
LOWER MIDDLE TRIASSIC EPOCH
LATE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: PARAREPTILIA OLSON 1947
INFRACLASS: SAUROPSIDA HUXLEY, 1864
ORDER: PROCOLOPHONOMORPHA ROMER, 1964
SUBORDER: PROCOLPHOMORPHA SEELEY, 1888
Family: PROCOLOPHONIDAE, Seeley, 1888
Subfamily: PROCOLOPHONINAE Seeley, 1888 (sensu Romer, 1956)
*Kapes bentoni
_____________________________________________________________________________________
ENGLAND
DEVON COUNTY
BETWEEN LADRAM BAY AND CHILESBURY BAY
SMALLSTONES POINT
MID TRIASSIC EPOCH
MIDDLE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
_____________________________________________________________________________________SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: RHYNOCEPHALIA nova
ORDER: RHYNOCEPHALIA BUTHER, 1867
SUBORDER: RHYNCHOSAUROIDEA GERVAIS, 1889
Family: RHYNCHOSAURIDAE Huxley, 1887
Rhynchosaurus spenceri
ENGLAND
DEVON COUNTY
SIDMOUTH
PICKET ROCK COVE
OTTER SANDSTONE
ANISIAN STAGE
LOWER MID TRIASSIC EPOCH
MIDDLE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
_____________________________________________________________________________________SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: RHYNOCEPHALIA nova
ORDER: RHYNOCEPHALIA BUTHER, 1867
SUBORDER: RHYNCHOSAUROIDEA GERVAIS, 1889
Family: RHYNCHOSAURIDAE Huxley, 1887
Rhynchosaurus spenceri
ENGLAND
DEVON COUNTY
CLOSE TO THE MOUTH OF THE RIVER OTTER, LEFT BANK
MIDDLE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
_____________________________________________________________________________________SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: RHYNOCEPHALIA nova
ORDER: RHYNOCEPHALIA BUTHER, 1867
SUBORDER: RHYNCHOSAUROIDEA GERVAIS, 1889
Family: RHYNCHOSAURIDAE Huxley, 1887
Rhynchosaurus spenceri
ENGLAND
DEVON COUNTY
OTTER SANDSTONE FORMATION
SHERWOOD SANDSTONE GROUP
ANISIAN STAGE
LOWER MIDDLE TRIASSIC EPOCH
LATE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
_____________________________________________________________________________________SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: PARAREPTILIA OLSON 1947
INFRACLASS: SAUROPSIDA HUXLEY, 1864
ORDER: PROCOLOPHONOMORPHA ROMER, 1964
SUBORDER: PROCOLPHOMORPHA SEELEY, 1888
Family: PROCOLOPHONIDAE, Seeley, 1888
Incertae sedis
Gen. sp indet.
ENGLAND
DEVON COUNTY
LYME REGIS
LOWER LIAS EPOCH
EARLY JURASSIC
STEM: TETRAPODA GOODRICH, 1930
_____________________________________________________________________________________SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: NEODIAPSIDA BENTON, 1985
NEODIAPSIDA: INCERTAE SEDIS
OR AMNIOTA: INCERTAE SEDIS
MAGANORDER: ICHTHYOPTERYGIA BAUR, 1887 emend nova
SUPERORDER: EOICHTHYOSAURIA MOTANI, 1999
GRANDORDER: ICHTHYOSAURIA EMEND BLAINVILLE, 1835
MICRORDER: LONGIPINNATI HUENE, 1948
ORDER: MERRIAMOSAURIA MONTANI, 1999
SUBORDER: EUICHTHYOSAURIA MONTANI, 1999
INFRAORDER: PARVIPELVIA MONTANI, 1999
PARVORDER: NEOICHTHYOSAURIA SANDER, 2000
Superfamily: EURHINOSAUROIDEA emend Montani, 1999
Family: LEPTONECTIDAE Maisch, 1998
*Leptonectes tenuirostris
= Ichthyosaurus tenuirostris
= Leptopterygius tenuirostris
ENGLAND
DEVON COUNTY
AXMOUTH
LIAS EPOCH
EARLY JURASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: NEODIAPSIDA BENTON, 1985
NEODIAPSIDA: INCERTAE SEDIS
OR AMNIOTA: INCERTAE SEDIS
MAGANORDER: ICHTHYOPTERYGIA BAUR, 1887 emend nova
SUPERORDER: EOICHTHYOSAURIA MOTANI, 1999
GRANDORDER: ICHTHYOSAURIA EMEND BLAINVILLE, 1835
MICRORDER: LONGIPINNATI HUENE, 1948
ORDER: MERRIAMOSAURIA MONTANI, 1999
SUBORDER: EUICHTHYOSAURIA MONTANI, 1999
INFRAORDER: PARVIPELVIA MONTANI, 1999
PARVORDER: NEOICHTHYOSAURIA SANDER, 2000
Superfamily: ICHTHYOSAUROIDEA Bonaparte, 1841 (Emend McGowan & Montani, 2003)
Family: ICHTHYOSAURIDAE Bonaparte, 1841
Ichthyosaurus communis
= Ichthyosaurus intermedius
_____________________________________________________________________________________