ENGLAND
EAST SUSSEX COUNTY
HASTINGS
SPECIFIC LOCALITY UNCERTAIN
WADHURST CLAY
VALANGINIAN STAGE
LOWER NEOCOMIAN SUBEPOCH
LOWER EARLY CRETACEOUS SUBEPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
CLADE NAME TYRANNORAPTORA SERENO, 1999
Incertae sedis
Gen. sp indet
____________________________________________________________________________________
ENGLAND
EAST SUSSEX COUNTY
NEAR BEXHILL
ON TURKEY ROAD
ASHDOWN BRICKWORKS (TQ 720095)
PEVENSEY PIT
'CONGLOMERATE BED'
WADHURST CLAY
VALANGINIAN STAGE
LOWER NEOCOMIAN SUBEPOCH
LOWER EARLY CRETACEOUS SUBEPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
ORDER: SPINOSAURIA OLSHEVSKY, 1991
Family: SPINOSAURIDAE Stromer, 1915
Morphotype I
Gen. sp indet.
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA PAUL, 2001
SUBORDER: COELURIA MARSH, 1881
Superfamily: TYRANNOSAUROIDEA Osborn, 1906 (sensu Bonaparte et. al., 1990)
Morphotype IV
Gen. sp indet
____________________________________________________________________________________
ENGLAND
EAST SUSSEX COUNTY
NEAR BEXHILL
ON TURKEY ROAD
ASHDOWN BRICKWORKS (TQ 720095)
PEVENSEY PIT
'POLACANTHUS BED'
WADHURST CLAY
VALANGINIAN STAGE
LOWER NEOCOMIAN SUBEPOCH
LOWER EARLY CRETACEOUS SUBEPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA PAUL, 2001
SUBORDER: COELURIA MARSH, 1881
Superfamily: TYRANNOSAUROIDEA Osborn, 1906 (sensu Bonaparte et. al., 1990)
Morphotype II
Gen. sp indet
AVETHEROPODA PAUL, 2001
AVEPECTORA PAUL, 2001
SUBORDER: MANIRAPTORA GAUTHIER, 1986
INFRAORDER: PARVES SERENO, 1997
PARVORDER: DEINONYCHOSAURIA COLBERT & D. A. RUSSELL, 1969
Family: DROMAEOSAURIDAE Russell, 1969
Morphotype III
Gen. sp indet
____________________________________________________________________________________
ENGLAND
EAST SUSSEX COUNTY
CLIFF END
CLIFF END BONE BED
WADHURST CLAY FORMATION
VALANGINIAN STAGE
MIDDLE NEOCOMIAN SUBEPOCH
LOWER EARLY CRETACEOUS EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
SUPERORDER: TAPINOPLATIA PETERS, 2000
GRANDORDER: CHARACIOPODA PETERS, 2000
MICRORDER: FENESTRASAURIA PETERS, 2000
ORDER: PTEROSAURIA KAUP, 1834
SUBORDER: EUPTERODACTYLOIDEA BENNETT, 1994
CLADE NAME: ORNITHOCHEIROIDEA SEELEY 1870 (sensu SEELEY, 1891)
CLADE NAME: PTERANODONTOIDEA MARSH, 1876 (SENUS KELLNER, 1996)
CLADE NAME LANCEODONTIA ANDRES, CLACK & XU, 2014
CLADE NAME: ISTIODACTYLIFORMES KELLNER, CALDWELL, HOLGADO, DALLA VECCHIA, NOHRA, SAYAO & CURRIE, 2019
Family: ISTIODACTYLIDAE Howse, Milner & Martill, 2001
Gen. sp indet.
____________________________________________________________________________________
ENGLAND
EAST SUSSEX COUNTY
BEXHILL
ASHDOWN QUARRY
PEVENSEY PIT
WADHURST CLAY FORMATION
HASTINGS GROUP
VALANGINIAN STAGE
MIDDLE NEOCOMIAN SUBEPOCH
LOWER EARLY CRETACEOUS EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: EURYOPDA SERENO, 1986
GRANDORDER: ARMATOSAURIA ZHAO, 1983 (emend)
ORDER: ANKYLOSAURIA OSBORN, 1923
SUBORDER: SCELIDOSAURIA COOPER, 1985
INFRAORDER: ANKYLOSAURA OSBORN, 1923
Family: POLACANTHIDAE Weiland, 1911
Polacanthus foxii
_____________________________________________________________________________________
ENGLAND
EAST SUSSEX COUNTY
NORTHWEST OF BEXHILL
ASHDOWN BRICKWORKS
WADHURST CLAY FORMATION
HASTINGS GROUP
VALANGINIAN STAGE
MIDDLE NEOCOMIAN SUBEPOCH
LOWER EARLY CRETACEOUS EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
ORDER: MESOSUCHIA EMEND HUXLEY, 1875
PARVORDER: METASUCHIA BENTON & CLARK, 1988
INFRAORDER: METAMESOSUCHIA HULKE, 1878
NEOSUCHIA CLARK, IN BENTON & CLARK, 1988
Family: BERNISSARTIIDAE Dollo, 1883
Bernissartia sp
Family: ATOPOSAURIDAE Gervais, 1871
Theriosuchus sp
Family: GONIOPHOLIDIDAE Cope, 1875
Hulkepholissp
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
SUPERORDER: TAPINOPLATIA PETERS, 2000
GRANDORDER: CHARACIOPODA PETERS, 2000
MICRORDER: FENESTRASAURIA PETERS, 2000
ORDER: PTEROSAURIA KAUP, 1834
Incertae sedis
Gen. sp indet
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
ORDER: SPINOSAURIA OLSHEVSKY, 1991
Family: BARYONYCHIDAE Charig & Milner, 1986
Baryonyx walkeri
ORDER: NEOTETANURAE SERENO, WILSON, LARSSON, DUTHEIL &
SUBORDER: AVETHEROPODA PAUL, 1988
Superfamily: ALLOSAUROIDEA Marsh, 1878 (sensu Currie & Zhao, 1994)
Family: ALLOSAURIDAE Marsh, 1878
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
SOUTHERN UNITED KINGDOM
ENGLAND
EAST SUSSEX COUNTY
LEWES
GLAUCONITIC MARL MEMBER
WEST MELBURY MARLY CHALK FORMATION (OLD
LOWER CHALK)
CENOMANIAN STAGE
UPPER MIDDLE GALLIC SUBEPOCH
LOWER GULF EPOCH
MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: EURYAPSIDA COLBERT, 1945
SUPERORDER: SAUROPTERYGIA OWEN, 1860
GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994
ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
INFRAORDER: PLIOSAUROMORPHA NOVA
Superfamily: PLIOSAUROIDEA Seeley, 1874
Family: PLIOSAURIDAE Seeley, 1874
Incertae sedis
Polyptychodon sp
_____________________________________________________________________________________
SOUTHERN UNITED KINGDOM
ENGLAND
EAST SUSSEX COUNTY
ON THE COAST NEAR HASTINGS
HASTINGS BEDS GROUP
UPPER BERRIASIAN-VALANGININ STAGE
LOWER GALLIC SUBEPOCH
MIDDLE EARLY CRETACEOUS EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
SUPERORDER: SAUROPODOMORPHA HUENE, 1932
GRANDORDER: BRONTOSAURIA OLSHEVSKY, 1991 (emend)
MICRORDER: SAUROPODA MARSH, 1878
ORDER: EUSAUROPODA
SUBORDER: NEOSAUROPODA BONAPARTE, 1986
INFRAORDER: MACRONARIA WILSON & SERENO, 1998
Incertae sedis
*Haestasaurus becklesii
= Pelorosaurus becklesi
= Morosaurus becklesi
____________________________________________________________________________________
SOUTHERN UNITED KINGDOM
ENGLAND
EAST SUSSEX COUNTY
SUSSEX COUNTY
BEXHILL
WADHURST CLAY FORMATION
HASTINGS GROUP
VALANGINIAN STAGE
LOWER NEOCOMIAN SUBSTAGE
LOWERMOST EARLY CRETACEOUS EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGANORDER: CRYPTODIRA GAFFNEY & MEYLAN, 1988
ORDER: CRYPTODIRA COPE, 1868
Family: SOLEMYDIDAE Lapparent & Murelaga, 1997
Gen. sp indet.
_____________________________________________________________________________________
SOUTHERN UNITED KINGDOM
ENGLAND
EAST SUSSEX COUNTY
BEXHILL
TUNBRIDGE WELLS SAND FORMATION
HASTINGS GROUP
WEALDEN SUPERGROUP
VALANGINIAN STAGE
MIDDLE NEOCOMIAN SUBEPOCH
LOWER EARLY CRETACEOUS EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
SUPERORDER: TAPINOPLATIA PETERS, 2000
GRANDORDER: CHARACIOPODA PETERS, 2000
MICRORDER: FENESTRASAURIA PETERS, 2000
ORDER: PTEROSAURIA KAUP, 1834
SUBORDER: EUPTERODACTYLOIDEA BENNETT, 1994
Superfamily: ORNITHOCHEIROIDEA Seeley, 1870 (sensu Seeley, 1891)
Family: Lonchodraconidae Rodrigues & Kellner, 2013
Lonchodraco sp
_____________________________________________________________________________________
SOUTHERN UNITED KINGDOM
ENGLAND
EAST SUSSEX COUNTY
NORTHWEST OF BEXHILL
ASHDOWN BRICKWORKS
PEVENSEY PIT
WADHURST CLAY FORMATION
HASTINGS GROUP
VALANGINIAN STAGE
MIDDLE NEOCOMIAN SUBEPOCH
LOWER EARLY CRETACEOUS EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?
SUPERORDER: BATRACHIA BRONGNIART, 1800
ORDER: CAUDATA SCOPOLI, 1777
Incertae sedis
Gen. sp indet.
SUPERORDER: SALIENTIA LAURENTI, 1768
Incertae sedis
Gen. sp indet.
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUNINATA KLEIN, 1760 (sensu JOYCE et al, 2004)
CLADE NAME: MESOCHELYDIA JOYCE, 2017
CLADE NAME: PERICHELYDIA JOYCE, 2017
Family: HELOCHELYDRIDAE Nopcsa, 1928
Helochelydra bakewelli
= Compsemys bakewelli
= Trionyx bakewelli
= Helochelydra bakewelli
= Tretosternon bakewelliCOHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGANORDER: CRYPTODIRA GAFFNEY & MEYLAN, 1988
SUPERORDER: SELMACRYPTODIRA GAFFNEY, HUTCHISON, JENKINS & MEEKR, 1987
GRANDORDER: PLEUROSTERNOIDEA GAFFNEY & MEYLAN, 1988
Superfamily: PLEUROSTERNIDEA Cope, 1868 (sensu Hay, 1893)
Family: PLEUROSTERNIDAE Cope, 1868
Glyptops sp
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: EURYAPSIDA COLBERT, 1945
SUPERORDER: SAUROPTERYGIA OWEN 1860
PARVORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994
ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
Incertae sedis
Gen. sp indet.
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: LACERTILIA OWEN, 1842
GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961
MICRORDER: SCINCOMORPHA CAMP, 1923
Incertae sedis
Gen. sp indet.
ORDER: ANGUIMORPHA FURBRINGER, 1900
SUBORDER: DIPLOGLOSSA COPE, 1864
INFRAORDER: THECAGLOSSA COPE, 1890
PARVORDER: MONSTERSAURIA NORELL & GAO, 1977
Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)
Family: MOSASAURIDAE Gervais, 1853
Subfamily: AIGALOSAURIDAE Gorjanovic-Kramberger, 1892
Incertae sedis
Gen. sp indet.
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
ORDER: MESOSUCHIA EMEND HUXLEY, 1875
PARVORDER: METASUCHIA BENTON & CLARK, 1988
INFRAORDER: METAMESOSUCHIA HULKE, 1878
NEOSUCHIA CLARK, IN BENTON & CLARK, 1988
Family: BERNISSARTIIDAE Dollo, 1883
Bernissartia sp
Family: ATOPOSAURIDAE Gervais, 1871
Theriosuchus sp
Family: GONIOPHOLIDIDAE Cope, 1875
Goniopholis sp
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
SUPERORDER: TAPINOPLATIA PETERS, 2000
GRANDORDER: CHARACIOPODA PETERS, 2000
MICRORDER: FENESTRASAURIA PETERS, 2000
ORDER: PTEROSAURIA KAUP, 1834
SUBORDER: EUPTERODACTYLOIDEA BENNETT, 1994
Superfamily: ORNITHOCHEIROIDEA Seeley, 1870 (sensu Seeley, 1891)
Family: ORNITHOCHEIRIDAE Seeley, 1870
Incertae sedis
Gen. sp indet.
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
Incertae sedis
Gen. sp indet.
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
ORDER: SPINOSAURIA OLSHEVSKY, 1991
Family: BARYONYCHIDAE Charig & Milner, 1986
Baryonyx walkeri
ORDER: NEOTETANURAE SERENO, WILSON, LARSSON, DUTHEIL &
SUBORDER: AVETHEROPODA PAUL, 1988
Superfamily: ALLOSAUROIDEA Marsh, 1878 (sensu Currie & Zhao, 1994)
Family: ALLOSAURIDAE Marsh, 1878
Incertae sedis
Gen. sp indet.
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
Incertae sedis
Gen. sp indet.
AVETHEROPODA PAUL, 2001
SUBORDER: COELURIA MARSH, 1881
AVEPECTORA PAUL, 2001
SUBORDER: MANIRAPTORA GAUTHIER, 1986
INFRAORDER: PARAVES SERENO, 1997
PARVORDER: DEINONYCHOSAURIA COLBERT & D. A. RUSSELL, 1969
Family: DROMAEOSAURIDAE Russell, 1969
Gen. sp indet.
Subfamily: VELOCIRAPTORINAE Barsbold, 1978
Gen. sp indet.
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
Gen. sp indet.
SUPERORDER: EURYOPDA SERENO, 1986
GRANDORDER: ARMATOSAURIA ZHAO, 1983 (emend)
ORDER: ANKYLOSAURIA OSBORN, 1923
SUBORDER: SCELIDOSAURIA COOPER, 1985
INFRAORDER: ANKYLOSAURA OSBORN, 1923
Family: POLACANTHIDAE Weiland, 1911
Hylaeosaurus sp
Polacanthus sp
_____________________________________________________________________________________
ENGLAND
EAST SUSSEX
BEXHILL
WEALDEN
WADHURST CLAY
VALANGINIAN STAGE
LOWER NEOCOMIAN SUBEPOCH
LOWER EARLY CRETACEOUS SUBEPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
ORDER: SPINOSAURIA OLSHEVSKY, 1991
Family: BARYONYCHIDAE Charig & Milner, 1986
Baryonyx sp
ORDER: NEOTETANURAE SERENO, WILSON, LARSSON, DUTHEIL & SUES, 1994
SUBORDER: CARNOSAURIA HUENE, 1920
Superfamily: Incertae sedis
Family: MEGALOSAURIDAE Huxley, 1889
Gen. sp indet.
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA PAUL, 2001
AVEPECTORA PAUL, 2001
SUBORDER: MANIRAPTORA GAUTHIER, 1986
INFRAORDER: PARVES SERENO, 1997
PARVORDER: DEINONYCHOSAURIA COLBERT & D. A. RUSSELL, 1969
Family: DROMAEOSAURIDAE Russell, 1969
Gen. sp indet.
COHORT: ORNITHISCHIFORMES COOPER, 1985
SUPERORDER: SAUROPODOMORPHA HUENE, 1932
GRANDORDER: BRONTOSAURIA OLSHEVSKY, 1991 (emend)
MICRORDER: SAUROPODA MARSH, 1878
ORDER: EUSAUROPODA
SUBORDER: NEOSAUROPODA BONAPARTE, 1986
INFRAORDER: HOMALOSAUROPIA JANENSCH, 1929
PARVORDER: DIPLODOCIMORPHA CALVO & SALGADO, 1995
Superfamily: DIPLODOCOIDEA Marsh, 1884 (sensu Upchurch, 1995)
Family: DIPLODOCIDAE Marsh, 1884
Gen. sp indet.
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
SUBORDER: ANKYLOPOLLEXIA SERENO, 1986
PARVORDER: HADROSAURIFORMES SERENO, 1997
Superfamily: IGUANODONTIDEA Cope, 1869 (sensu Sereno, 1986)
Family: IGUANODONTIDAE Cope, 1869
Iguanodon sp
SUPERORDER: EURYOPDA SERENO, 1986
GRANDORDER: ARMATOSAURIA ZHAO, 1983 (emend)
ORDER: ANKYLOSAURIA OSBORN, 1923
SUBORDER: SCELIDOSAURIA COOPER, 1985
INFRAORDER: ANKYLOSAURA OSBORN, 1923
Family: POLACANTHIDAE Weiland, 1911
Hylaeosaurus sp
_____________________________________________________________________________________
ENGLAND
EAST SUSSEX
HASTINGS
WEALDEN
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
SUPERORDER: EURYOPDA SERENO, 1986
GRANDORDER: ARMATOSAURIA ZHAO, 1983 (emend)
ORDER: ANKYLOSAURIA OSBORN, 1923
SUBORDER: SCELIDOSAURIA COOPER, 1985
INFRAORDER: ANKYLOSAURA OSBORN, 1923
Family: NODOSAURIDAE Marsh, 1890
Gen. sp indet.
_____________________________________________________________________________________
SOUTHERN UNITED KINGDOM
ENGLAND
EAST SUSSEX COUNTY
UNCERTAIN, BUT PRESUMABLY HASTINGS
UNCERTAIN, BUT PRESUMABLY HASTINGS GROUP
BERRIASIAN-VALANGINIAN STAGE
LOWER-MIDDLE NEOCOMIAN SUBEPOCH
LOWER EARLY CRETACEOUS EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
ORDER: NEOTETANURAE SERENO, WILSON, LARSSON, DUTHEIL & SUES, 1994
SUBORDER: AVETHEROPODA PAUL, 1988
Superfamily: ALLOSAUROIDEA Marsh, 1878 (sensu Currie & Zhao, 1994)
Family: ALLOSAURIDAE Marsh, 1878
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
ENGLAND
EAST SUSSEX COUNTY
ASHDOWN SANDS
CLIFF END BONEBED
HASTINGS
WEALDEN
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: MAMMALIA LINNAEUS, 1758
SUBCLASS: THERIIFORMES (ROWE, 1988) McKENNA & BELL, 1997
INFRACLASS: ALLOTHERIA (MARSH, 1880) McKENNA & BELL, 1997
ORDER: MULTITUBERCULATA COPE, 1884
SUBORDER: PLAGIAULACOIDEA (SIMPSON, 1925) HAHN, 1969
Family: PLAGIAULACIDAE Gill, 1872
Subfamily: EOBAATARIDAE Keilan-Jaworowska, Dashzeveg & Trofimov, 1987
*Loxaulax valdensis
= Dipriodon valdensis
Loxaulax spSUBCLASS: PROTOTHERIA GILL, 1872
INFRACLASS: HOLOTHERIA (WIBEL, ROUGIER, NOVACEK, MCKENNA & DASHZEVEG, 1995) MCKENNA & BELL, 1997
SUPERLEGION: SYMMETRODONTA (SIMPSON, 1925) McKENNA & BELL, 1997
LEGION: SYMMETRODONTA SIMPSON, 1925
ORDER: SPALACOTHERIIDA PROTHERO, 1981
Family: SPALACOTHERIIDAE Marsh, 1887
Spalacotherium tricuspidens
Spalacotherium spINFRACLASS: TRECHNOTHERIA McKENNA, 1975
LEGION: CLADOTHERIA McKENNA, 1975
SUBLEGION: DRYOLESTOIDEA BUTLER, 1939
ORDER: DRYOLESTOIDA PROTHERO, 1981
Family: DRYOLESTIDAE Marsh, 1879
Laolestes hodsoni
= *Melanodon hodsoniMelanodon sp
SUBLEGION: ZATHERIA MCKENNA, 1975
INFRALEGION: TRIBOSPHENIDA (McKENNA, 1975) McKENNA & BELL, 1997
SUPERCHORT: AEGIALODONTIA (BUTLER, 1978) McKENNA & BELL, 1997
Family: AEGALODONTIDAE Kermack, Lees & Mussett, 1965
*Aegialodon dawsonii
_____________________________________________________________________________________
ENGLAND
EAST SUSSEX COUNTY
HASTINGS
WEALDEN FORMATION
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: MAMMALIA LINNAEUS, 1758
SUBCLASS: THERIIFORMES (ROWE, 1988) McKENNA & BELL, 1997
INFRACLASS: ALLOTHERIA (MARSH, 1880) McKENNA & BELL, 1997
ORDER: MULTITUBERCULATA COPE, 1884
SUBORDER: PLAGIAULACOIDEA (SIMPSON, 1925) HAHN, 1969
Family: PLAGIAULACIDAE Gill, 1872
Subfamily: PLAGIAULACINAE Gill, 1872
Gen. sp indet.
= Bolodon sp
_____________________________________________________________________________________
ENGLAND
EAST SUSSEX COUNTY
HASTINGS
WADHURST CLAY
HASTINGS BEDS
(ASHDOWN BEDS, TUNBRIDGE WELLS SAND)
BERRIASIAN-VALANGINIAN STAGE
LOWER NEOCOMIAN SUBEPOCH
LOWER EARLY CRETACEOUS SUBEPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: EURYAPSIDA COLBERT, 1945
SUPERORDER: SAUROPTERYGIA OWEN 1860
GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994
ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
Incertae sedis
Gen. sp. indet.
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
ORDER: MESOSUCHIA EMEND HUXLEY, 1875
PARVORDER: METASUCHIA BENTON & CLARK, 1988
INFRAORDER: METAMESOSUCHIA HULKE, 1878
NEOSUCHIA CLARK, IN BENTON & CLARK, 1988
Family: BERNISSARTIIDAE Dollo, 1883
Bernissartia fagesii
= *Crocodilus saulii
Bernissartia spFamily: ATOPOSAURIDAE Gervais, 1871
Theriosuchus pusillus
Theriosuchus spFamily: GONIOPHOLIDIDAE Cope, 1875
Incertae sedis
Goniopholis sp
_____________________________________________________________________________________
ENGLAND
EAST SUSSEX COUNTY
HASTINGS SAND
HASTINGS BEDS
BERRIASIAN-VALANGINIAN STAGE
LOWER-MIDDLE NEOCOMIAN SUBEPOCH
LOWER EARLY CRETACEOUS EPOCH
EARLY CRETACEOUS
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUNINATA KLEIN, 1760 (sensu JOYCE et al, 2004)
CLADE NAME: MESOCHELYDIA JOYCE, 2017
CLADE NAME: PERICHELYDIA JOYCE, 2017
Family: HELOCHELYDRIDAE Nopcsa, 1928
*Helochelydra bakewelli
_____________________________________________________________________________________
ENGLAND
EAST SUSSEX COUNTY
HASTINGS SAND
HASTINGS BEDS
(ASHDOWN BEDS, TUNBRIDGE WELLS SAND)
BERRIASIAN-VALANGINIAN STAGE
LOWER-MIDDLE NEOCOMIAN SUBEPOCH
LOWER EARLY CRETACEOUS EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
ORDER: EUSUCHIA emend HUXLEY, 1875
Family: HYLAEOCHAMPSIDAE Andrews, 1913
Hylaeochampsa vectiana
= *Heterosuchus valdensis
_____________________________________________________________________________________
ENGLAND
EAST SUSSEX COUNTY
HASTINGS
WADHURST CLAY
BERRIASIAN-VALANGINIAN STAGE
LOWER NEOCOMIAN SUBEPOCH
LOWER EARLY CRETACEOUS SUBEPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: EURYAPSIDA COLBERT, 1945
SUPERORDER: SAUROPTERYGIA OWEN, 1860
GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994
ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
INFRAORDER: PLIOSAUROMORPHA NOVA
Superfamily: PLIOSAUROIDEA Seeley, 1874
Family: PLIOSAURIDAE Seeley, 1874
*Hastanects valdensis
= Cimoliasaurus valdensis
= Plesiosaurus valdensis
_____________________________________________________________________________________
ENGLAND
EAST SUSSEX COUNTY
HASTINGS
HASTINGS BEDS
BERRIASIAN-VALANGINIAN STAGE
LOWER NEOCOMIAN SUBSTAGE
LOWERMOST EARLY CRETACEOUS EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGANORDER: CRYPTODIRA GAFFNEY & MEYLAN, 1988
SUPERORDER: SELMACRYPTODIRA GAFFNEY, HUTCHISON, JENKINS & MEEKR, 1987
GRANDORDER: DAIOCRYPTODIRA GAFFNEY & MEYLAN, 1988
MICRORDER: BAENOIDEA WILLIAMS, 1960
Family: NEURANKYLIDAE Lambe, 1902
Compsemys bakewelli
= *Helochelydra bakewelli
= Trionyx bakewelli
= Helochelydra bakewelli
= Tretosternon bakewelliMICRORDER: EUCRYPTODIRA GAFFNEY, 1975
Family: PLESIOCHELYIDAE Rutimeyer, 1873
Plesiochelys brodiei
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
SUPERORDER: TAPINOPLATIA PETERS, 2000
GRANDORDER: CHARACIOPODA PETERS, 2000
MICRORDER: FENESTRASAURIA PETERS, 2000
ORDER: PTEROSAURIA KAUP, 1834
SUBORDER: EUPTERODACTYLOIDEA BENNETT, 1994
Superfamily: ORNITHOCHEIROIDEA Seeley, 1870 (sensu Seeley, 1891)
Family: ORNITHOCHEIRIDAE Seeley, 1870
Incertae sedis
Ornithocheirus sp
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
ORDER: MESOSUCHIA EMEND HUXLEY, 1875
PARVORDER: METASUCHIA BENTON & CLARK, 1988
INFRAORDER: METAMESOSUCHIA HULKE, 1878
NEOSUCHIA CLARK, IN BENTON & CLARK, 1988
Family: GONIOPHOLIDIDAE Cope, 1875
Goniopholis crassidens
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
Incertae sedis
Gen. sp indet.
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
ORDER: NEOTETANURAE SERENO, WILSON, LARSSON, DUTHEIL & SUES, 1994
SUBORDER: CARNOSAURIA HUENE, 1920
Superfamily: Incertae sedis
Family: MEGALOSAURIDAE Huxley, 1889
Incertae sedis
Megalosaurus sp
SUBORDER: AVETHEROPODA PAUL, 1988
Superfamily: ALLOSAUROIDEA Marsh, 1878 (sensu Currie & Zhao, 1994)
Family: ALLOSAURIDAE Marsh, 1878
Incertae sedis
Becklespinax altispinax
= Acrocanthosaurus altispinaxCOHORT: ORNITHISCHIFORMES COOPER, 1985
SUPERORDER: SAUROPODOMORPHA HUENE, 1932
GRANDORDER: BRONTOSAURIA OLSHEVSKY, 1991 (emend)
MICRORDER: SAUROPODA MARSH, 1878
ORDER: EUSAUROPODA
SUBORDER: NEOSAUROPODA BONAPARTE, 1986
INFRAORDER: MACRONARIA WILSON & SERENO, 1998
PARVORDER: TITANOSAURIFORMES SALGADO ET AL, 1997
Family: BRACHIOSAURIDAE Riggs, 1904
Subfamily: BRACHIOSAURINAE Riggs, 1904 (sensu Janensch, 1929)
Ornithopsis hulkei
= Pelorosaurus hulkeiPelorosaurus conybearei
= Cetiosaurus brevis
= Morosaurus brevisMAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
SUPERORDER: EURYOPDA SERENO, 1986
GRANDORDER: STEGOSAUROMORPHA COOPER, 1985 (EMEND)
ORDER: STEGOSAURIA MARSH, 1877
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
ENGLAND
EAST SUSSEX COUNTY
HASTINGS
WADHURST CLAY
LOWER VALANGINIAN STAGE
LOWER NEOCOMIAN SUBEPOCH
LOWER EARLY CRETACEOUS SUBEPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: MAMMALIA LINNAEUS, 1758
SUBCLASS: THERIIFORMES (ROWE, 1988) McKENNA & BELL, 1997
INFRACLASS: ALLOTHERIA (MARSH, 1880) McKENNA & BELL, 1997
ORDER: MULTITUBERCULATA COPE, 1884
SUBORDER: PLAGIAULACOIDEA (SIMPSON, 1925) HAHN, 1969
Family: PLAGIAULACIDAE Gill, 1872
Subfamily: PLAGIAULACINAE Gill, 1872
?*Plagiaulax dawsoni
= Plioprion? dawsoni
= Loxaulax dawsoni
_____________________________________________________________________________________
ENGLAND
EAST SUSSEX COUNTY
CROWBOROUGH
A QUARRY NEAR WINDLESHAM HOUSE
EARLY CRETACEOUS
ICHNOSTEM: TETRAPODIPIDA NOVA
ICHNOSUPERCLASS: AMNIOTAIPIDII NOVA
ICHNOCLASS: REPTILIPEDIA VIALOV, 1966
ICHNOSUBCLASS: DIAPSIDIPIDII NOVA
ICHNOSTEM-GROUP: NEODIAPSIDIPEDII NOVA
ICHNOINFRACLASS: NEODIAPSIDIPEDII NOVA
ICHNOSUBDIVISION: ARCHOSAURIPEDIDII NOVA
ICHNOLEGION: DINOSAUROMORPHIPEDA NOVA
ICHNOSUBLEGION: DINOSAURIFORMEIPEDIDII NOVA
ICHNOSUPERCOHORT: DINOSAUROPEDIDA NOVA
ICHNOCOHORT: ORNITHISCHIFORMEIPIDA NOVA
ICHNOMAGNORDER: ORNITHISCHIPIDA NOVA
ICHNOSUPERORDER: NEORNITHISCHIPIDA NOVA
ICHNOGRANDORDER: ORNITHOPODIPDIA NOVA
ICHNOMICRORDER: EUORNITHOPODIPIDA NOVA
ICHNOORDER: IGUANODONTIPIDIA NOVA
ICHNOSUBORDER: ANKYLOPOLLEXIPEDIDA NOVA
ICHNOINFRAORDER: STYRACOSTERNIPIDA NOVA
ICHNOPARVORDER: HADROSAURIFORMIPEDIDA NOVA
Ichnosuperfamily: IGUANODONTIDEA Nova
Ichnofamily: IGUANODONTIPODIDAE emend Vylov, 1988
Gen. sp indet.
_____________________________________________________________________________________
ENGLAND
EAST SUSSEX COUNTY
EASTBOURNE-HASTINGS
ASHDOWN SAND
ASHDOWN SANDSTONE
EARLY CRETACEOUS
ICHNOSTEM: TETRAPODIPIDA NOVA
ICHNOSUPERCLASS: AMNIOTAIPIDII NOVA
ICHNOCLASS: REPTILIPEDIA VIALOV, 1966
ICHNOSUBCLASS: DIAPSIDIPIDII NOVA
ICHNOSTEM-GROUP: NEODIAPSIDIPEDII NOVA
ICHNOINFRACLASS: NEODIAPSIDIPEDII NOVA
ICHNOSUBDIVISION: ARCHOSAURIPEDIDII NOVA
ICHNOLEGION: DINOSAUROMORPHIPEDA NOVA
ICHNOSUBLEGION: DINOSAURIFORMEIPEDIDII NOVA
ICHNOSUPERCOHORT: DINOSAUROPEDIDA NOVA
ICHNOCOHORT: ORNITHISCHIFORMEIPIDA NOVA
ICHNOMAGNORDER: ORNITHISCHIPIDA NOVA
ICHNOSUPERORDER: NEORNITHISCHIPIDA NOVA
ICHNOGRANDORDER: ORNITHOPODIPDIA NOVA
ICHNOMICRORDER: EUORNITHOPODIPIDA NOVA
ICHNOORDER: IGUANODONTIPIDIA NOVA
ICHNOSUBORDER: ANKYLOPOLLEXIPEDIDA NOVA
ICHNOINFRAORDER: STYRACOSTERNIPIDA NOVA
ICHNOPARVORDER: HADROSAURIFORMIPEDIDA NOVA
Ichnosuperfamily: IGUANODONTIDEA Nova
Ichnofamily: IGUANODONTIPODIDAE emend Vylov, 1988
Gen. sp indet.
_____________________________________________________________________________________
ENGLAND
EAST SUSSEX COUNTY
PADDOCKHURST PARK
PADDOCKHURST BONE BED
WEALDEN FORMATION
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: MAMMALIA LINNAEUS, 1758
SUBCLASS: THERIIFORMES (ROWE, 1988) McKENNA & BELL, 1997
INFRACLASS: ALLOTHERIA (MARSH, 1880) McKENNA & BELL, 1997
Incertae sedis
Gen. sp indet.
SUBCLASS: PROTOTHERIA GILL, 1872
INFRACLASS: HOLOTHERIA (WIBEL, ROUGIER, NOVACEK, MCKENNA & DASHZEVEG, 1995) MCKENNA & BELL, 1997
SUPERLEGION: SYMMETRODONTA (SIMPSON, 1925) McKENNA & BELL, 1997
LEGION: SYMMETRODONTA SIMPSON, 1925
ORDER: SPALACOTHERIIDA PROTHERO, 1981
Family: SPALACOTHERIIDAE Marsh, 1887
Spalacotherium tricuspidens
*Spalacotherium tayloriINFRACLASS: TRECHNOTHERIA McKENNA, 1975
LEGION: CLADOTHERIA McKENNA, 1975
SUBLEGION: ZATHERIA MCKENNA, 1975
INFRALEGION: TRIBOSPHENIDA (McKENNA, 1975) McKENNA & BELL, 1997
SUPERCOHORT: THERIA (PARKER & HASWELL, 1987) McKENNA & BELL, 1997
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
ENGLAND
EAST SUSSEX COUNTY
HASTINGS
WEALDEN BEDS
VALANGINIAN OR HAUTERIVIAN STAGE
LOWER NEOCOMANIAN SUBEPOCH
LOWER EARLY CRETACEOUS EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
SUPERORDER: EURYOPDA SERENO, 1986
GRANDORDER: ARMATOSAURIA ZHAO, 1983 (emend)
ORDER: ANKYLOSAURIA OSBORN, 1923
SUBORDER: SCELIDOSAURIA COOPER, 1985
INFRAORDER: ANKYLOSAURA OSBORN, 1923
Family: NODOSAURIDAE Marsh, 1890
Gen. sp indet.
____________________________________________________________________________________
ENGLAND
EAST SUSSEX COUNTY
HASTINGS
UPPER WEALDEN FORMATION
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
SUPERORDER: SAUROPODOMORPHA HUENE, 1932
GRANDORDER: BRONTOSAURIA OLSHEVSKY, 1991 (emend)
MICRORDER: SAUROPODA MARSH, 1878
ORDER: EUSAUROPODA
SUBORDER: NEOSAUROPODA BONAPARTE, 1986
INFRAORDER: MACRONARIA WILSON & SERENO, 1998
PARVORDER: TITANOSAURIFORMES SALGADO ET AL, 1997
Family: BRACHIOSAURIDAE Riggs, 1904
Subfamily: BRACHIOSAURINAE Riggs, 1904 (sensu Janensch, 1929)
Pelorosaurus conybearei
_____________________________________________________________________________________
ENGLAND
EAST SUSSEX COUNTY
NEAR HASTINGS
WEALDEN FORMATION
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
SUBORDER: ANKYLOPOLLEXIA SERENO, 1986
PARVORDER: HADROSAURIFORMES SERENO, 1997
Superfamily: IGUANODONTIDEA Cope, 1869 (sensu Sereno, 1986)
Family: IGUANODONTIDAE Cope, 1869
Iguanodon sp
____________________________________________________________________________________
ENGLAND
EAST SUSSEX COUNTY
NEAR HASTINGS
NEAR THE BASE OF THE CLIFF SECTION NEAR LEE NESS, AT FAIRLIGHT
LEE NESS SANDSTONE
ASHDOWN BEDS
VALANGINIAN STAGE
UPPER NEOCOMIAN SUBEPOCH
EARLY EARLY CRETACEOUS EPOCH
EARLY CRETACEOUS
ICHNOSTEM: TETRAPODIPIDA NOVA
ICHNOSUPERCLASS: AMNIOTAIPIDII NOVA
ICHNOCLASS: REPTILIPEDIA VIALOV, 1966
ICHNOSUBCLASS: DIAPSIDIPIDII NOVA
ICHNOSTEM-GROUP: NEODIAPSIDIPEDII NOVA
ICHNOINFRACLASS: NEODIAPSIDIPEDII NOVA
ICHNOSUBDIVISION: ARCHOSAURIPEDIDII NOVA
ICHNOLEGION: DINOSAUROMORPHIPEDA NOVA
ICHNOSUBLEGION: DINOSAURIFORMEIPEDIDII NOVA
ICHNOSUPERCOHORT: DINOSAUROPEDIDA NOVA
ICHNOCOHORT: THEROPODIPEDIA NOVA
Incerate sedis
Gen. sp indet.
ICHNOCOHORT: ORNITHISCHIFORMEIPIDA NOVA
ICHNOMAGNORDER: ORNITHISCHIPIDA NOVA
ICHNOSUPERORDER: NEORNITHISCHIPIDA NOVA
ICHNOGRANDORDER: ORNITHOPODIPDIA NOVA
ICHNOMICRORDER: EUORNITHOPODIPIDA NOVA
ICHNOORDER: IGUANODONTIPIDIA NOVA
ICHNOSUBORDER: ANKYLOPOLLEXIPEDIDA NOVA
ICHNOINFRAORDER: STYRACOSTERNIPIDA NOVA
ICHNOPARVORDER: HADROSAURIFORMIPEDIDA NOVA
Ichnosuperfamily: IGUANODONTIDEA Nova
Ichnofamily: IGUANODONTIPODIDAE emend Vylov, 1988
Gen. sp indet.
_____________________________________________________________________________________
ENGLAND
EAST SUSSEX COUNTY
NEAR HASTINGS
NEAR THE BASE OF THE CLIFF SECTION NEAR LEE NESS, AT FAIRLIGHT
LEE NESS SANDSTONE
ASHDOWN BEDS
VALANGINIAN STAGE
UPPER NEOCOMIAN SUBEPOCH
EARLY EARLY CRETACEOUS EPOCH
EARLY CRETACEOUS
ICHNOSTEM: TETRAPODIPIDA NOVA
ICHNOSUPERCLASS: AMNIOTAIPIDII NOVA
ICHNOCLASS: REPTILIPEDIA VIALOV, 1966
ICHNOSUBCLASS: DIAPSIDIPIDII NOVA
ICHNOSTEM-GROUP: NEODIAPSIDIPEDII NOVA
ICHNOINFRACLASS: NEODIAPSIDIPEDII NOVA
ICHNOSUBDIVISION: ARCHOSAURIPEDIDII NOVA
ICHNOLEGION: DINOSAUROMORPHIPEDA NOVA
ICHNOSUBLEGION: DINOSAURIFORMEIPEDIDII NOVA
ICHNOSUPERCOHORT: DINOSAUROPEDIDA NOVA
ICHNOCOHORT: ORNITHISCHIFORMEIPIDA NOVA
ICHNOMAGNORDER: ORNITHISCHIPIDA NOVA
ICHNOSUPERORDER: NEORNITHISCHIPIDA NOVA
ICHNOGRANDORDER: ORNITHOPODIPDIA NOVA
ICHNOMICRORDER: EUORNITHOPODIPIDA NOVA
ICHNOORDER: IGUANODONTIPIDIA NOVA
ICHNOSUBORDER: ANKYLOPOLLEXIPEDIDA NOVA
ICHNOINFRAORDER: STYRACOSTERNIPIDA NOVA
ICHNOPARVORDER: HADROSAURIFORMIPEDIDA NOVA
Ichnosuperfamily: IGUANODONTIDEA Nova
Ichnofamily: IGUANODONTIPODIDAE emend Vylov, 1988
Gen. sp indet.
_____________________________________________________________________________________
ENGLAND
EAST SUSSEX COUNTY
NEAR HASTINGS
NEAR THE BASE OF THE CLIFF SECTION NEAR LEE NESS, AT FAIRLIGHT
DARK-RED SANDSTONE HORIZON
ASHDOWN BEDS
VALANGINIAN STAGE
UPPER NEOCOMIAN SUBEPOCH
EARLY EARLY CRETACEOUS EPOCH
EARLY CRETACEOUS
ICHNOSTEM: TETRAPODIPIDA NOVA
ICHNOSUPERCLASS: AMNIOTAIPIDII NOVA
ICHNOCLASS: REPTILIPEDIA VIALOV, 1966
ICHNOSUBCLASS: DIAPSIDIPIDII NOVA
ICHNOSTEM-GROUP: NEODIAPSIDIPEDII NOVA
ICHNOINFRACLASS: NEODIAPSIDIPEDII NOVA
ICHNOSUBDIVISION: ARCHOSAURIPEDIDII NOVA
ICHNOLEGION: DINOSAUROMORPHIPEDA NOVA
ICHNOSUBLEGION: DINOSAURIFORMEIPEDIDII NOVA
ICHNOSUPERCOHORT: DINOSAUROPEDIDA NOVA
ICHNOCOHORT: ORNITHISCHIFORMEIPIDA NOVA
ICHNOMAGNORDER: ORNITHISCHIPIDA NOVA
ICHNOSUPERORDER: NEORNITHISCHIPIDA NOVA
ICHNOGRANDORDER: ORNITHOPODIPDIA NOVA
ICHNOMICRORDER: EUORNITHOPODIPIDA NOVA
ICHNOORDER: IGUANODONTIPIDIA NOVA
ICHNOSUBORDER: ANKYLOPOLLEXIPEDIDA NOVA
ICHNOINFRAORDER: STYRACOSTERNIPIDA NOVA
ICHNOPARVORDER: HADROSAURIFORMIPEDIDA NOVA
Ichnosuperfamily: IGUANODONTIDEA Nova
Ichnofamily: IGUANODONTIPODIDAE emend Vylov, 1988
Gen. sp indet.
_____________________________________________________________________________________
ENGLAND
EAST SUSSEX COUNTY
NEAR HASTINGS
NEAR THE BASE OF THE CLIFF SECTION NEAR LEE NESS, AT FAIRLIGHT
GREY SANDSTONE HORIZON
ASHDOWN BEDS
VALANGINIAN STAGE
UPPER NEOCOMIAN SUBEPOCH
EARLY EARLY CRETACEOUS EPOCH
EARLY CRETACEOUS
ICHNOSTEM: TETRAPODIPIDA NOVA
ICHNOSUPERCLASS: AMNIOTAIPIDII NOVA
ICHNOCLASS: REPTILIPEDIA VIALOV, 1966
ICHNOSUBCLASS: DIAPSIDIPIDII NOVA
ICHNOSTEM-GROUP: NEODIAPSIDIPEDII NOVA
ICHNOINFRACLASS: NEODIAPSIDIPEDII NOVA
ICHNOSUBDIVISION: ARCHOSAURIPEDIDII NOVA
ICHNOLEGION: DINOSAUROMORPHIPEDA NOVA
ICHNOSUBLEGION: DINOSAURIFORMEIPEDIDII NOVA
ICHNOSUPERCOHORT: DINOSAUROPEDIDA NOVA
ICHNOCOHORT: THEROPODIPEDIA NOVA
Incerate sedis
Gen. sp indet.
ICHNOCOHORT: ORNITHISCHIFORMEIPIDA NOVA
ICHNOMAGNORDER: ORNITHISCHIPIDA NOVA
ICHNOSUPERORDER: NEORNITHISCHIPIDA NOVA
ICHNOGRANDORDER: ORNITHOPODIPDIA NOVA
ICHNOMICRORDER: EUORNITHOPODIPIDA NOVA
ICHNOORDER: IGUANODONTIPIDIA NOVA
ICHNOSUBORDER: ANKYLOPOLLEXIPEDIDA NOVA
ICHNOINFRAORDER: STYRACOSTERNIPIDA NOVA
ICHNOPARVORDER: HADROSAURIFORMIPEDIDA NOVA
Ichnosuperfamily: IGUANODONTIDEA Nova
Ichnofamily: IGUANODONTIPODIDAE emend Vylov, 1988
Gen. sp indet.
_____________________________________________________________________________________
ENGLAND
EAST SUSSEX COUNTY
NEAR HASTINGS
BULVERHYTHE
WEALDEN FORMATION
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
SUBORDER: ANKYLOPOLLEXIA SERENO, 1986
PARVORDER: HADROSAURIFORMES SERENO, 1997
Superfamily: IGUANODONTIDEA Cope, 1869 (sensu Sereno, 1986)
Family: IGUANODONTIDAE Cope, 1869
Iguanodon sp
____________________________________________________________________________________
ENGLAND
EAST SUSSEX COUNTY
NEAR HASTINGS
WADHURST CLAY
LOWER WEALDEN FORMATION
LOWER VALANGINIAN STAGE
MIDDLE NEOCOMIAN SUBEPOCH
LOWER EARLY CRETACEOUS EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
SUBORDER: ANKYLOPOLLEXIA SERENO, 1986
PARVORDER: HADROSAURIFORMES SERENO, 1997
Superfamily: IGUANODONTIDEA Cope, 1869 (sensu Sereno, 1986)
Family: IGUANODONTIDAE Cope, 1869
Iguanodon sp
SUPERORDER: EURYOPDA SERENO, 1986
GRANDORDER: ARMATOSAURIA ZHAO, 1983 (emend)
ORDER: ANKYLOSAURIA OSBORN, 1923
SUBORDER: SCELIDOSAURIA COOPER, 1985
INFRAORDER: ANKYLOSAURA OSBORN, 1923
Family: NODOSAURIDAE Marsh, 1890
Gen. sp indet.
____________________________________________________________________________________
ENGLAND
EAST SUSSEX COUNTY
NEAR HASTINGS
HOLLINGTON
WADHURST CLAY
LOWER WEALDEN FORMATION
LOWER VALANGINIAN STAGE
MIDDLE NEOCOMIAN SUBEPOCH
LOWER EARLY CRETACEOUS EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
SUBORDER: ANKYLOPOLLEXIA SERENO, 1986
PARVORDER: HADROSAURIFORMES SERENO, 1997
Superfamily: IGUANODONTIDEA Cope, 1869 (sensu Sereno, 1986)
Family: IGUANODONTIDAE Cope, 1869
Iguanodon anglicus
= Iguanodon mantelli
= Therosaurus mantelli
= Iguanodon mantelli
_____________________________________________________________________________________
ENGLAND
EAST SUSSEX COUNTY
NEAR HASTINGS
RIDGE QUARRY
WADHURST CLAY
LOWER WEALDEN FORMATION
LOWER VALANGINIAN STAGE
MIDDLE NEOCOMIAN SUBEPOCH
LOWER EARLY CRETACEOUS EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
SUBORDER: ANKYLOPOLLEXIA SERENO, 1986
PARVORDER: HADROSAURIFORMES SERENO, 1997
Superfamily: IGUANODONTIDEA Cope, 1869 (sensu Sereno, 1986)
Family: IGUANODONTIDAE Cope, 1869
Iguanodon sp
____________________________________________________________________________________
ENGLAND
EAST SUSSEX COUNTY
NEAR HASTINGS
SHORDEN QUARRY IN THE VILLAGE OF SHORNDEN
WADHURST CLAY
LOWER WEALDEN FORMATION
LOWER VALANGINIAN STAGE
MIDDLE NEOCOMIAN SUBEPOCH
LOWER EARLY CRETACEOUS EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
SUBORDER: ANKYLOPOLLEXIA SERENO, 1986
INFRAORDER: STYRACOSTERNA SERENO, 1986
Incertae sedis
*Darwinsaurus evolutionis
*Hypselospinus fittoni
= Iguanodon? fittoni
= Iguanodon hollingtoniensisPARVORDER: HADROSAURIFORMES SERENO, 1997
Superfamily: IGUANODONTIDEA Cope, 1869 (sensu Sereno, 1986)
Family: IGUANODONTIDAE Cope, 1869
Iguanodon sp
____________________________________________________________________________________
SOUTHERN UNITED KINGDOM
ENGLAND
EAST SUSSEX COUNTY
SAINT LEONARDS
OLD ROAR QUARRY
LOWER VALANGINIAN STAGE
MIDDLE NEOCOMIAN SUBEPOCH
LOWER EARLY CRETACEOUS EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
Incertae sedis
*Sellacoxa pauli
____________________________________________________________________________________
ENGLAND
EAST SUSSEX COUNTY
HASTINGS
HASTINGS BEDS
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
ORDER: SPINOSAURIA OLSHEVSKY, 1991
Family: BARYONYCHIDAE Charig & Milner, 1986
Suchosaurus sp
_____________________________________________________________________________________
ENGLAND
EAST SUSSEX COUNTY
HASTINGS
WEALDEN FORMATION
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
Incertae sedis
Gen. sp indet.
COHORT: ORNITHISCHIFORMES COOPER, 1985
SUPERORDER: SAUROPODOMORPHA HUENE, 1932
GRANDORDER: BRONTOSAURIA OLSHEVSKY, 1991 (emend)
MICRORDER: SAUROPODA MARSH, 1878
ORDER: EUSAUROPODA
SUBORDER: NEOSAUROPODA BONAPARTE, 1986
INFRAORDER: MACRONARIA WILSON & SERENO, 1998
PARVORDER: TITANOSAURIFORMES SALGADO ET AL, 1997
Family: BRACHIOSAURIDAE Riggs, 1904
Incertae sedis
non Pelorosaurus, P? becklesii
= Pelorosaurus becklesi
= Morosaurus becklesi
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
SUBORDER: ANKYLOPOLLEXIA SERENO, 1986
PARVORDER: HADROSAURIFORMES SERENO, 1997
Superfamily: IGUANODONTIDEA Cope, 1869 (sensu Sereno, 1986)
Family: IGUANODONTIDAE Cope, 1869
Iguanodon sp
____________________________________________________________________________________
ENGLAND
EAST SUSSEX COUNTY
HASTINGS
WADHURST CLAY
LOWER WEALDEN FORMATION
LOWER VALANGINIAN STAGE
MIDDLE NEOCOMIAN SUBEPOCH
LOWER EARLY CRETACEOUS EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
Incertae sedis
*Brailium dawsoni
= Iguanodon dawsoniSUBORDER: DRYOMORPHA SERNEO, 1986
Family: DRYOSAURIDAE Milner & Norman, 1984
Valdosaurus sp
= Hypsilophodon foxii
= Hypsilophodon foxi
= Iguanodon foxiSUBORDER: ANKYLOPOLLEXIA SERENO, 1986
INFRAORDER: STYRACOSTERNA SERENO, 1986
Hypselospinus fittoni
= Iguanodon? fittoniPARVORDER: HADROSAURIFORMES SERENO, 1997
Superfamily: IGUANODONTIDEA Cope, 1869 (sensu Sereno, 1986)
Family: IGUANODONTIDAE Cope, 1869
Iguanodon? bernissartensis
_____________________________________________________________________________________
ENGLAND
EAST SUSSEX COUNTY
FROM THE COAST OF YORKSHIRE OR HASTINGS
UNKNOWN
CORALLIAN
OXFORDIAN STAGE
LOWER MALM EPOCH
LATE JURASSIC OR WEALDEN
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
SUPERORDER: EURYOPDA SERENO, 1986
GRANDORDER: ARMATOSAURIA ZHAO, 1983 (emend)
ORDER: ANKYLOSAURIA OSBORN, 1923
SUBORDER: SCELIDOSAURIA COOPER, 1985
INFRAORDER: ANKYLOSAURA OSBORN, 1923
Family: NODOSAURIDAE Marsh, 1890
*Priodontognathus phillipsii
= Iguanodon phillipsii
= Dacentrurus phillipsii
= Omosaurus phillipsii
_____________________________________________________________________________________
ENGLAND
EAST SUSSEX COUNTY
BATTLE-NEAR-HASTINGS
WESSEX FORMATION
WEALDEN GROUP
BARREMIAN STAGE
LOWER GALLIC SUBEPOCH
EARLY CRETACEOUS EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
ORDER: NEOTETANURAE SERENO, WILSON, LARSSON, DUTHEIL & SUES, 1994
SUBORDER: AVETHEROPODA PAUL, 1988
Superfamily: ALLOSAUROIDEA Marsh, 1878 (sensu Currie & Zhao, 1994)
Family: ALLOSAURIDAE Marsh, 1878
Incertae sedis
*Becklespinax altispinax
= Acrocanthosaurus altispinax
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
SUBORDER: ANKYLOPOLLEXIA SERENO, 1986
PARVORDER: HADROSAURIFORMES SERENO, 1997
Superfamily: IGUANODONTIDEA Cope, 1869 (sensu Sereno, 1986)
Family: IGUANODONTIDAE Cope, 1869
Iguanodon sp
____________________________________________________________________________________
ENGLAND
EAST SUSSEX COUNTY
BATTLE-NEAR HASTINGS
LOWER WEALDEN FORMATION
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
SUPERORDER: EURYOPDA SERENO, 1986
GRANDORDER: ARMATOSAURIA ZHAO, 1983 (emend)
ORDER: ANKYLOSAURIA OSBORN, 1923
SUBORDER: SCELIDOSAURIA COOPER, 1985
INFRAORDER: ANKYLOSAURA OSBORN, 1923
Family: POLACANTHIDAE Weiland, 1911
Hylaeosaurus armatus
_____________________________________________________________________________________
ENGLAND
EAST SUSSEX COUNTY
NEAR BATTLE
WEALDEN FORMATION
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
ORDER: NEOTETANURAE SERENO, WILSON, LARSSON, DUTHEIL & SUES, 1994
SUBORDER: CARNOSAURIA HUENE, 1920
Superfamily: Incertae sedis
Family: MEGALOSAURIDAE Huxley, 1889
Megalosaurus bucklandii
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
SUBORDER: ANKYLOPOLLEXIA SERENO, 1986
PARVORDER: HADROSAURIFORMES SERENO, 1997
Superfamily: IGUANODONTIDEA Cope, 1869 (sensu Sereno, 1986)
Family: IGUANODONTIDAE Cope, 1869
Iguanodon anglicus
= Iguanodon mantelli
= Therosaurus mantelli
= Iguanodon mantelli
____________________________________________________________________________________
ENGLAND
EAST SUSSEX COUNTY
IN THE NEIGHBORHOOD OF BATTLE
WEALDEN FORMATION
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
ORDER: NEOTETANURAE SERENO, WILSON, LARSSON, DUTHEIL & SUES, 1994
SUBORDER: CARNOSAURIA HUENE, 1920
Superfamily: Incertae sedis
Family: MEGALOSAURIDAE Huxley, 1889
Megalosaurus bucklandii
_____________________________________________________________________________________
SOUTHERN UNITED KINGDOM
ENGLAND
EAST SUSSEX COUNTY
BATTLE
HASTINGS SUB-GROUP
BERRIASIAN-VALANGINIAN STAGE
LOWER NEOCOMIAN SUBSTAGE
LOWERMOST EARLY CRETACEOUS EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUNINATA KLEIN, 1760 (sensu JOYCE et al, 2004)
CLADE NAME: MESOCHELYDIA JOYCE, 2017
CLADE NAME: PERICHELYDIA JOYCE, 2017
Family: HELOCHELYDRIDAE Nopcsa, 1928
Helochelydra bakewelli
= Compsemys bakewelli
= Trionyx bakewelli
= Helochelydra bakewelli
= Tretosternon bakewelli
_____________________________________________________________________________________
ENGLAND
EAST SUSSEX COUNTY
BATTLE NEAR HASTINGS
HOLLINGTON QUARRY
WADHURST CLAY
LOWER WEALDEN FORMATION
LOWER VALANGINIAN STAGE
MIDDLE NEOCOMIAN SUBEPOCH
LOWER EARLY CRETACEOUS EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
ORDER: NEOTETANURAE SERENO, WILSON, LARSSON, DUTHEIL & SUES, 1994
SUBORDER: AVETHEROPODA PAUL, 1988
Superfamily: ALLOSAUROIDEA Marsh, 1878 (sensu Currie & Zhao, 1994)
Family: ALLOSAURIDAE Marsh, 1878
Incertae sedis
Becklespinax altispinax
= Acrocanthosaurus altispinaxValdoraptor oweni
= Hylaeosaurus
= Megalosaurus dunkeri
= Megalosaurus oweni
= Altispinax oweni
= Megalosaurus dunkeri
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
SUBORDER: ANKYLOPOLLEXIA SERENO, 1986
INFRAORDER: STYRACOSTERNA SERENO, 1986
*Huxleysaurus hollingtoniensis
= Iguanodon hollingtoniensisPARVORDER: HADROSAURIFORMES SERENO, 1997
Superfamily: IGUANODONTIDEA Cope, 1869 (sensu Sereno, 1986)
Family: IGUANODONTIDAE Cope, 1869
Iguanodon? bernissartensis
Iguanodon sp
____________________________________________________________________________________
ENGLAND
EAST SUSSEX COUNTY
WADHURST CLAY
BREDE BONE-BED
WEALDEN FORMATION
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: EURYAPSIDA COLBERT, 1945
SUPERORDER: SAUROPTERYGIA OWEN 1860
GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994
ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
Incertae sedis
Gen. sp. indet.
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
ORDER: SPINOSAURIA OLSHEVSKY, 1991
Family: BARYONYCHIDAE Charig & Milner, 1986
Suchosaurus cultridens
_____________________________________________________________________________________
ENGLAND
EAST SUSSEX COUNTY
BREDE
HARE FARM LANE
BASAL WADHURST CLAY
LOWER WEALDEN FORMATION
LOWER VALANGINIAN STAGE
MIDDLE NEOCOMIAN SUBEPOCH
LOWER EARLY CRETACEOUS EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
ORDER: SPINOSAURIA OLSHEVSKY, 1991
Family: BARYONYCHIDAE Charig & Milner, 1986
Suchosaurus sp
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
SUBORDER: ANKYLOPOLLEXIA SERENO, 1986
INFRAORDER: STYRACOSTERNA SERENO, 1986
Hypselospinus fittoni
= Iguanodon? fittoni
_____________________________________________________________________________________
ENGLAND
EAST SUSSEX COUNTY
BREDE
WADHURST CLAY
LOWER WEALDEN FORMATION
LOWER VALANGINIAN STAGE
MIDDLE NEOCOMIAN SUBEPOCH
LOWER EARLY CRETACEOUS EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
SUBORDER: ANKYLOPOLLEXIA SERENO, 1986
INFRAORDER: STYRACOSTERNA SERENO, 1986
Incertae sedis
Brailium dawsoni
= Iguanodon dawsoniSUBORDER: ANKYLOPOLLEXIA SERENO, 1986
PARVORDER: HADROSAURIFORMES SERENO, 1997
Superfamily: IGUANODONTIDEA Cope, 1869 (sensu Sereno, 1986)
Family: IGUANODONTIDAE Cope, 1869
Iguanodon sp
_____________________________________________________________________________________
SOUTHERN UNITED KINGDOM
ENGLAND
EAST SUSSEX COUNTY
JUST TO THE WEST OF HASTINGS
WEST MARINA, THE QUARRY MAY HAVE BEEN LINKED TO THE CONSTRUCTION OF EITHER THE MARINA PARADE OR ST LEONARDS RAILWAY STATION
WADHURST CLAY
LOWER WEALDEN FORMATION
HASTINGS BEDS
VALANGINIAN STAGE
MIDDLE NEOCOMIAN SUBEPOCH
LOWER EARLY CRETACEOUS EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
SUBORDER: ANKYLOPOLLEXIA SERENO, 1986
INFRAORDER: STYRACOSTERNA SERENO, 1986
Incertae sedis
Brailium dawsoni
= Iguanodon dawsoni
_____________________________________________________________________________________
SOUTHERN UNITED KINGDOM
ENGLAND
EAST SUSSEX COUNTY
HASTINGS
SHORNDEN LOCALITY
WADHURST CLAY
LOWER WEALDEN FORMATION
HASTINGS BEDS
VALANGINIAN STAGE
MIDDLE NEOCOMIAN SUBEPOCH
LOWER EARLY CRETACEOUS EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
SUBORDER: ANKYLOPOLLEXIA SERENO, 1986
INFRAORDER: STYRACOSTERNA SERENO, 1986
Incertae sedis
Brailium dawsoni
= Iguanodon dawsoni
_____________________________________________________________________________________
SOUTHERN UNITED KINGDOM
ENGLAND
EAST SUSSEX COUNTY
HASTINGS
BUCKSHOLE QUARRY
WADHURST CLAY
LOWER WEALDEN FORMATION
HASTINGS BEDS
VALANGINIAN STAGE
MIDDLE NEOCOMIAN SUBEPOCH
LOWER EARLY CRETACEOUS EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
SUBORDER: ANKYLOPOLLEXIA SERENO, 1986
INFRAORDER: STYRACOSTERNA SERENO, 1986
Incertae sedis
Brailium dawsoni
= Iguanodon dawsoni
_____________________________________________________________________________________
ENGLAND
EAST SUSSEX COUNTY
HASTINGS
BREDE-BONE BED
WEALDEN FORMATION
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUNINATA KLEIN, 1760 (sensu JOYCE et al, 2004)
Incertae sedis
Genus sp indet.
CLADE NAME: MESOCHELYDIA JOYCE, 2017
CLADE NAME: PERICHELYDIA JOYCE, 2017
Family: HELOCHELYDRIDAE Nopcsa, 1928
Helochelydra bakewelli
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
Incerate sedis
Gen. sp. indet.
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
ORDER: MESOSUCHIA EMEND HUXLEY, 1875
PARVORDER: METASUCHIA BENTON & CLARK, 1988
INFRAORDER: METAMESOSUCHIA HULKE, 1878
NEOSUCHIA CLARK, IN BENTON & CLARK, 1988
Family: GONIOPHOLIDIDAE Cope, 1875
Goniopholis crassidens
_____________________________________________________________________________________
ENGLAND
EAST SUSSEX COUNTY
NEAR HASTINGS
SILVER HILL
WADHURST CLAY
LOWER WEALDEN FORMATION
LOWER VALANGINIAN STAGE
MIDDLE NEOCOMIAN SUBEPOCH
LOWER EARLY CRETACEOUS EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
SUPERORDER: EURYOPDA SERENO, 1986
GRANDORDER: ARMATOSAURIA ZHAO, 1983 (emend)
ORDER: ANKYLOSAURIA OSBORN, 1923
SUBORDER: SCELIDOSAURIA COOPER, 1985
INFRAORDER: ANKYLOSAURA OSBORN, 1923
Family: POLACANTHIDAE Weiland, 1911
Hylaeosaurus armatus
_____________________________________________________________________________________
ENGLAND
EAST SUSSEX COUNTY
NEAR HASTINGS
SILVERHILL
WEALDEN FORMATION
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
SUBORDER: ANKYLOPOLLEXIA SERENO, 1986
PARVORDER: HADROSAURIFORMES SERENO, 1997
Superfamily: IGUANODONTIDEA Cope, 1869 (sensu Sereno, 1986)
Family: IGUANODONTIDAE Cope, 1869
Iguanodon sp
____________________________________________________________________________________
ENGLAND
EAST SUSSEX COUNTY
7 KM NE OF HASTINGS
CLIFF END BONE BED
WEALDEN GROUP
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
Incerate sedis
Gen. sp. indet.
_____________________________________________________________________________________
ENGLAND
EAST SUSSEX COUNTY
TELHAM
BLACK HORSE QUARRY
HASTINGS BEDS
VALANGINIAN STAGE
MIDDLE NEOCOMIAN SUBEPOCH
LOWER EARLY CRETACEOUS EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUNINATA KLEIN, 1760 (sensu JOYCE et al, 2004)
CLADE NAME: MESOCHELYDIA JOYCE, 2017
CLADE NAME: PERICHELYDIA JOYCE, 2017
Family: HELOCHELYDRIDAE Nopcsa, 1928
Helochelydra bakewelli
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGANORDER: CRYPTODIRA GAFFNEY & MEYLAN, 1988
SUPERORDER: SELMACRYPTODIRA GAFFNEY, HUTCHISON, JENKINS & MEEKR, 1987
GRANDORDER: DAIOCRYPTODIRA GAFFNEY & MEYLAN, 1988
MICRORDER: EUCRYPTODIRA GAFFNEY, 1975
Family: PLESIOCHELYIDAE Rutimeyer, 1873
Plesiochelys sp
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: EURYAPSIDA COLBERT, 1945
SUPERORDER: SAUROPTERYGIA OWEN 1860
GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994
ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
Incertae sedis
Gen. sp. indet.
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
SUPERORDER: TAPINOPLATIA PETERS, 2000
GRANDORDER: CHARACIOPODA PETERS, 2000
MICRORDER: FENESTRASAURIA PETERS, 2000
ORDER: PTEROSAURIA KAUP, 1834
SUBORDER: EUPTERODACTYLOIDEA BENNETT, 1994
Superfamily: ORNITHOCHEIROIDEA Seeley, 1870 (sensu Seeley, 1891)
Family: ORNITHOCHEIRIDAE Seeley, 1870
Ornithocheirus? clifti
= Palaeornis clifti
= Ornithochirus clifti
= Pterodactylus cliftiSUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
ORDER: MESOSUCHIA EMEND HUXLEY, 1875
PARVORDER: METASUCHIA BENTON & CLARK, 1988
INFRAORDER: METAMESOSUCHIA HULKE, 1878
NEOSUCHIA CLARK, IN BENTON & CLARK, 1988
Family: PHOLIDOSAURIDAE Williston, 1902
Suchosaurus cultridens (referred material)
Family: GONIOPHOLIDIDAE Cope, 1875
Goniopholis crassidens
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
ORDER: NEOTETANURAE SERENO, WILSON, LARSSON, DUTHEIL & SUES, 1994
SUBORDER: CARNOSAURIA HUENE, 1920
Superfamily: Incertae sedis
Family: MEGALOSAURIDAE Huxley, 1889
Incertae sedi
Megalosaurus sp
SUBORDER: AVETHEROPODA PAUL, 1988
Superfamily: ALLOSAUROIDEA Marsh, 1878 (sensu Currie & Zhao, 1994)
Family: ALLOSAURIDAE Marsh, 1878
Incertae sedis
Becklespinax altispinax
= Acrocanthosaurus altispinaxCOHORT: ORNITHISCHIFORMES COOPER, 1985
SUPERORDER: SAUROPODOMORPHA HUENE, 1932
GRANDORDER: BRONTOSAURIA OLSHEVSKY, 1991 (emend)
MICRORDER: SAUROPODA MARSH, 1878
ORDER: EUSAUROPODA
SUBORDER: NEOSAUROPODA BONAPARTE, 1986
Family: CETIOSAURIDAE Lydekker, 1888
Subfamily: CETIOSAURINAE Lydekker, 1888 (sensu Janensch, 1929)
Cetiosaurus sp
INFRAORDER: MACRONARIA WILSON & SERENO, 1998
PARVORDER: TITANOSAURIFORMES SALGADO ET AL, 1997
Family: BRACHIOSAURIDAE Riggs, 1904
Subfamily: BRACHIOSAURINAE Riggs, 1904 (sensu Janensch, 1929)
Pelorosaurus conybearei
= Pleurocelus valdensis
= Astrodon valdensis
= Astrodon (Pleurocelus) valdensis
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
SUBORDER: ANKYLOPOLLEXIA SERENO, 1986
PARVORDER: HADROSAURIFORMES SERENO, 1997
Superfamily: IGUANODONTIDEA Cope, 1869 (sensu Sereno, 1986)
Family: IGUANODONTIDAE Cope, 1869
Iguanodon sp
SUPERORDER: EURYOPDA SERENO, 1986
GRANDORDER: ARMATOSAURIA ZHAO, 1983 (emend)
ORDER: ANKYLOSAURIA OSBORN, 1923
SUBORDER: SCELIDOSAURIA COOPER, 1985
INFRAORDER: ANKYLOSAURA OSBORN, 1923
Family: POLACANTHIDAE Weiland, 1911
Hylaeosaurus armatus
_____________________________________________________________________________________
ENGLAND
EAST SUSSEX COUNTY
NEAR LEWES AND BRIGHTON
LEWES NODULAR CHALK MEMBER
UPPER CHALK FORMATION
UPPER TURONIAN STAGE - LOWER CONIACIAN STAGE
LOWEST SENONIAN SUBEPOCH
LOWER GULF EPOCH
EARLY LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: LACERTILIA OWEN, 1842
GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961
MICRORDER: SCINCOMORPHA CAMP, 1923
ORDER: ANGUIMORPHA FURBRINGER, 1900
SUBORDER: DIPLOGLOSSA COPE, 1864
INFRAORDER: THECAGLOSSA COPE, 1890
PARVORDER: MONSTERSAURIA NORELL & GAO, 1977
Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)
Family: MOSASAURIDAE Gervais, 1853
Infrafamily: NATANIA
Subfamily: MOSASAURINAE Gervais, 1853 (sensu Williston, 1897)
Tribe: MOSASAURINI Gervais, 1853 (sensu Russell, 1967)
Mosasaurus gracilis
_____________________________________________________________________________________
ENGLAND
EAST SUSSEX COUNTY
BRIGHTON AND LEWES RAILWAY
CHALK
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: LACERTILIA OWEN, 1842
GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961
MICRORDER: SCINCOMORPHA CAMP, 1923
ORDER: ANGUIMORPHA FURBRINGER, 1900
SUBORDER: DIPLOGLOSSA COPE, 1864
INFRAORDER: THECAGLOSSA COPE, 1890
PARVORDER: MONSTERSAURIA NORELL & GAO, 1977
Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)
Family: MOSASAURIDAE Gervais, 1853
Infrafamily: NATANIA
Subfamily: MOSASAURINAE Gervais, 1853 (sensu Williston, 1897)
Tribe: LEIODONTINI Lingham-Soliar, 1993
Leiodon anceps
_____________________________________________________________________________________
ENGLAND
EAST SUSSEX COUNTY
BRIGHTON
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: LACERTILIA OWEN, 1842
GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961
MICRORDER: SCINCOMORPHA CAMP, 1923
ORDER: ANGUIMORPHA FURBRINGER, 1900
SUBORDER: DIPLOGLOSSA COPE, 1864
INFRAORDER: THECAGLOSSA COPE, 1890
PARVORDER: MONSTERSAURIA NORELL & GAO, 1977
Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)
Family: MOSASAURIDAE Gervais, 1853
Infrafamily: NATANIA
Subfamily: MOSASAURINAE Gervais, 1853 (sensu Williston, 1897)
Tribe: LEIODONTINI Lingham-Soliar, 1993
Leiodon anceps
_____________________________________________________________________________________
ENGLAND
EAST SUSSEX COUNTY
NEAR BRIGHTON
CLAYTON CHALK PIT
CHALK MARL FORMATION (LOWER CHALK)
LOWER TO MIDDLE CENOMANIAN STAGE
UPPER GALLIC SUBEPOCH
LOWER GULF EPOCH
MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: LACERTILIA OWEN, 1842
GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961
MICRORDER: SCINCOMORPHA CAMP, 1923
ORDER: ANGUIMORPHA FURBRINGER, 1900
SUBORDER: DIPLOGLOSSA COPE, 1864
INFRAORDER: THECAGLOSSA COPE, 1890
PARVORDER: MONSTERSAURIA NORELL & GAO, 1977
Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)
Family: MOSASAURIDAE Gervais, 1853
Subfamily: DOLICHOSAURIDAE Gervais, 1852
Tribe: DOLICHOSAURINI Gervais, 1852
*Coniasaurus crassidens
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ENGLAND
EAST SUSSEX COUNTY
ECCLESBOURNE
HASTINGS BEDS
BERRIASIAN-VALANGINIAN STAGE
LOWER-MIDDLE NEOCOMIAN SUBEPOCH
LOWER EARLY CRETACEOUS EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
ORDER: MESOSUCHIA EMEND HUXLEY, 1875
PARVORDER: METASUCHIA BENTON & CLARK, 1988
INFRAORDER: METAMESOSUCHIA HULKE, 1878
NEOSUCHIA CLARK, IN BENTON & CLARK, 1988
Family: GONIOPHOLIDIDAE Cope, 1875
Goniopholis crassidens
Incertae sedis
Gen. sp. Indet.
_____________________________________________________________________________________
SOUTHERN UNITED KINGDOM
ENGLAND
EAST SUSSEX COUNTY
ECCLESBOURNE GLEN
HASTINGS BEDS
BERRIASIAN-VALANGINIAN STAGE
LOWER-MIDDLE NEOCOMIAN SUBEPOCH
LOWER EARLY CRETACEOUS EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
SUPERORDER: SAUROPODOMORPHA HUENE, 1932
GRANDORDER: BRONTOSAURIA OLSHEVSKY, 1991 (emend)
MICRORDER: SAUROPODA MARSH, 1878
ORDER: EUSAUROPODA
SUBORDER: NEOSAUROPODA BONAPARTE, 1986
INFRAORDER: HOMALOSAUROPIA JANENSCH, 1929
PARVORDER: DIPLODOCIMORPHA CALVO & SALGADO, 1995
Superfamily: DIPLODOCOIDEA Marsh, 1884 (sensu Upchurch, 1995)
Family: REBBACHISAURIDAE Bonaparte, 1996
*Xenoposeidon proneneukos
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ENGLAND
EAST SUSSEX COUNTY
UPPER PART OF THE CLIFF A LITTLE WEST OF ECCLESBOURNE GLEN
WEALDEN FORMATION
EARLY CRETACEOUS
ICHNOSTEM: TETRAPODIPIDA NOVA
ICHNOSUPERCLASS: AMNIOTAIPIDII NOVA
ICHNOCLASS: REPTILIPEDIA VIALOV, 1966
ICHNOSUBCLASS: DIAPSIDIPIDII NOVA
ICHNOSTEM-GROUP: NEODIAPSIDIPEDII NOVA
ICHNOINFRACLASS: NEODIAPSIDIPEDII NOVA
ICHNOSUBDIVISION: ARCHOSAURIPEDIDII NOVA
ICHNOLEGION: DINOSAUROMORPHIPEDA NOVA
ICHNOSUBLEGION: DINOSAURIFORMEIPEDIDII NOVA
ICHNOSUPERCOHORT: DINOSAUROPEDIDA NOVA
ICHNOCOHORT: ORNITHISCHIFORMEIPIDA NOVA
ICHNOMAGNORDER: ORNITHISCHIPIDA NOVA
ICHNOSUPERORDER: NEORNITHISCHIPIDA NOVA
ICHNOGRANDORDER: ORNITHOPODIPDIA NOVA
ICHNOMICRORDER: EUORNITHOPODIPIDA NOVA
ICHNOORDER: IGUANODONTIPIDIA NOVA
ICHNOSUBORDER: ANKYLOPOLLEXIPEDIDA NOVA
ICHNOINFRAORDER: STYRACOSTERNIPIDA NOVA
ICHNOPARVORDER: HADROSAURIFORMIPEDIDA NOVA
Ichnosuperfamily: IGUANODONTIDEA Nova
Ichnofamily: IGUANODONTIPODIDAE emend Vylov, 1988
Gen. sp indet.
_____________________________________________________________________________________
ENGLAND
EAST SUSSEX COUNTY
ECCLESBOURNE
WADHURST CLAY
WEALDEN FORMATION
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
SUBORDER: ANKYLOPOLLEXIA SERENO, 1986
PARVORDER: HADROSAURIFORMES SERENO, 1997
Superfamily: IGUANODONTIDEA Cope, 1869 (sensu Sereno, 1986)
Family: IGUANODONTIDAE Cope, 1869
Iguanodon sp
____________________________________________________________________________________
ENGLAND
EAST SUSSEX COUNTY
HARE FARM LANE
BASAL WADHURST CLAY
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
ORDER: MESOSUCHIA EMEND HUXLEY, 1875
PARVORDER: METASUCHIA BENTON & CLARK, 1988
INFRAORDER: METAMESOSUCHIA HULKE, 1878
NEOSUCHIA CLARK, IN BENTON & CLARK, 1988
Family: GONIOPHOLIDIDAE Cope, 1875
Goniopholis crassidens
_____________________________________________________________________________________
ENGLAND
EAST SUSSEX COUNTY
ST. LEONARDS
LOWER WEALDEN FORMATION
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
SUPERORDER: EURYOPDA SERENO, 1986
GRANDORDER: ARMATOSAURIA ZHAO, 1983 (emend)
ORDER: ANKYLOSAURIA OSBORN, 1923
SUBORDER: SCELIDOSAURIA COOPER, 1985
INFRAORDER: ANKYLOSAURA OSBORN, 1923
Family: POLACANTHIDAE Weiland, 1911
Hylaeosaurus armatus
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ENGLAND
EAST SUSSEX COUNTY
NEAR LEWES
OFFHAM CHALK PIT
LEWES NODULAR CHALK MEMBER
UPPER CHALK FORMATION
UPPER TURONIAN STAGE - LOWER CONIACIAN STAGE
LOWEST SENONIAN SUBEPOCH
LOWER GULF EPOCH
EARLY LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: LACERTILIA OWEN, 1842
GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961
MICRORDER: SCINCOMORPHA CAMP, 1923
ORDER: ANGUIMORPHA FURBRINGER, 1900
SUBORDER: DIPLOGLOSSA COPE, 1864
INFRAORDER: THECAGLOSSA COPE, 1890
PARVORDER: MONSTERSAURIA NORELL & GAO, 1977
Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)
Family: MOSASAURIDAE Gervais, 1853
Infrafamily: NATANIA
Subfamily: MOSASAURINAE Gervais, 1853 (sensu Williston, 1897)
Tribe: MOSASAURINI Gervais, 1853 (sensu Russell, 1967)
*Mosasaurus gracilis
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ENGLAND
EAST SUSSEX COUNTY
NEAR LEWES
SOUTHERAM PIT
MIDDLE LOWER CHALK
ACANTHOSCERAS RHOTOMANGENSE ZONE
MIDDLE CENOMANIAN STAGE
UPPERMOST GALLIC SUBEPOCH
LOWER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: LACERTILIA OWEN, 1842
GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961
MICRORDER: SCINCOMORPHA CAMP, 1923
ORDER: ANGUIMORPHA FURBRINGER, 1900
SUBORDER: DIPLOGLOSSA COPE, 1864
INFRAORDER: THECAGLOSSA COPE, 1890
PARVORDER: MONSTERSAURIA NORELL & GAO, 1977
Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)
Family: MOSASAURIDAE Gervais, 1853
Subfamily: DOLICHOSAURIDAE Gervais, 1852
Tribe: DOLICHOSAURINI Gervais, 1852
Dolichosaurus longicollis
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ENGLAND
EAST SUSSEX COUNTY
ST.-LEONARD'S-ON-SEA
HASTINGS SANDS
UPPER TUNBRIDGE WELLS SAND FORMATION (OLD WEALDEN FORMATION)
UPPER VALANGINIAN OR LOWER HAUTERIVIAN STAGE
MIDDLE NEOCOMIAN SUBEPOCH
LOWER EARLY CRETACEOUS EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
SUPERORDER: TAPINOPLATIA PETERS, 2000
GRANDORDER: CHARACIOPODA PETERS, 2000
MICRORDER: FENESTRASAURIA PETERS, 2000
ORDER: PTEROSAURIA KAUP, 1834
SUBORDER: EUPTERODACTYLOIDEA BENNETT, 1994
Superfamily: ORNITHOCHEIROIDEA Seeley, 1870 (sensu Seeley, 1891)
Family: ORNITHOCHEIRIDAE Seeley, 1870
*Coloborhynchus clavirostris
= Criorhynchus clavirostris
= Ornithocheirus clavirostrisFamily: LonchodraconidaeRodrigues & Kellner, 2013
*Serradraco sagittirostris= Ornithocheirus, sagittirostris
= Pterodactylus sagittirostris
= Lonchodectes sagittirostris
= Ornithodesmus sagittirostris
_____________________________________________________________________________________
ENGLAND
EAST SUSSEX COUNTY
LOWER CHALK
CENOMANIAN STAGE
UPPER GALLIC SUBEPOCH
LOWER GULF EPOCH
MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: LACERTILIA OWEN, 1842
GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961
MICRORDER: SCINCOMORPHA CAMP, 1923
ORDER: ANGUIMORPHA FURBRINGER, 1900
SUBORDER: DIPLOGLOSSA COPE, 1864
INFRAORDER: THECAGLOSSA COPE, 1890
PARVORDER: MONSTERSAURIA NORELL & GAO, 1977
Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)
Family: MOSASAURIDAE Gervais, 1853
Subfamily: DOLICHOSAURIDAE Gervais, 1852
Tribe: DOLICHOSAURINI Gervais, 1852
Coniasaurus crassidens
_____________________________________________________________________________________
ENGLAND
EAST SUSSEX COUNTY
NEAR FOLKSTONE
LIDDNE SPOUT
LOWER CHALK
CENOMANIAN STAGE
UPPER GALLIC SUBEPOCH
LOWER GULF EPOCH
MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: LACERTILIA OWEN, 1842
GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961
MICRORDER: SCINCOMORPHA CAMP, 1923
ORDER: ANGUIMORPHA FURBRINGER, 1900
SUBORDER: DIPLOGLOSSA COPE, 1864
INFRAORDER: THECAGLOSSA COPE, 1890
PARVORDER: MONSTERSAURIA NORELL & GAO, 1977
Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)
Family: MOSASAURIDAE Gervais, 1853
Subfamily: DOLICHOSAURIDAE Gervais, 1852
Tribe: DOLICHOSAURINI Gervais, 1852
*Coniasaurus gracilodens
_____________________________________________________________________________________
ENGLAND
EAST SUSSEX COUNTY
NEAR BRIGHTON
FALMER
LOWER CHALK
CENOMANIAN STAGE
UPPER GALLIC SUBEPOCH
LOWER GULF EPOCH
MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: LACERTILIA OWEN, 1842
GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961
MICRORDER: SCINCOMORPHA CAMP, 1923
ORDER: ANGUIMORPHA FURBRINGER, 1900
SUBORDER: DIPLOGLOSSA COPE, 1864
INFRAORDER: THECAGLOSSA COPE, 1890
PARVORDER: MONSTERSAURIA NORELL & GAO, 1977
Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)
Family: MOSASAURIDAE Gervais, 1853
Subfamily: DOLICHOSAURIDAE Gervais, 1852
Tribe: DOLICHOSAURINI Gervais, 1852
Coniasaurus crassidens
_____________________________________________________________________________________