ENGLAND
KENT COUNTY
FOLKESTONE WARREN
LOWER GREENSAND
MAMMILATUM
EARLY CRETACEOUS
ICHNOSTEM: TETRAPODIPIDA NOVA
ICHNOSUPERCLASS: AMNIOTAIPIDII NOVA
ICHNOCLASS: REPTILIPEDIA VIALOV, 1966
ICHNOSUBCLASS: DIAPSIDIPIDII NOVA
ICHNOSTEM-GROUP: NEODIAPSIDIPEDII NOVA
ICHNOINFRACLASS: NEODIAPSIDIPEDII NOVA
ICHNOSUBDIVISION: ARCHOSAURIPEDIDII NOVA
ICHNOSUPERCOHORT: AVEMETATARSALIPEDII NOVA
ICHNOLEGION: DINOSAUROMORPHIPEDA NOVA
ICHNOSUBLEGION: DINOSAURIFORMEIPEDIDII NOVA
ICHNOSUPERCOHORT: DINOSAUROPEDIDA NOVA
ICHNOCOHORT: THEROPODIPEDIA NOVA
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
SOUTHERN UNITED KINGDOM
ENGLAND
KENT COUNTY
1.5 KM NNE OF ST. MARGARETS AT CLIFFE, BETWEEN DOVER AND DEAL ON THE EAST KENT COAST
HOPE POINT
HOPE POINT MARLS
BASAL BROADSTAIRS MEMBER
WHITE CHALK FORMATION
LOWER MICRASTER CORANGUINUM ZONE
LOWER PART OF THE RANGE OF VOLVICERAMUS INVOLUTUS
CONIACIAN STAGE
LOWER SENONIAN SUBEPOCH
LOWER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
SUPERORDER: TAPINOPLATIA PETERS, 2000
GRANDORDER: CHARACIOPODA PETERS, 2000
MICRORDER: FENESTRASAURIA PETERS, 2000
ORDER: PTEROSAURIA KAUP, 1834
SUBORDER: EUPTERODACTYLOIDEA BENNETT, 1994
Superfamily: AZHDARCHIDEA Nesov, 1984
Family: TAPEJARIDAE Kellner, 1989
Gen. sp indet.
_____________________________________________________________________________________
ENGLANDKENT COUNTY
LOWER WEALDEN FORMATION
UPPER BARREMIAN STAGE
LOWER NEOCOMIAN SUBEPOCH
MIDDLE EARLY CRETACEOUS EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
SUBORDER: ANKYLOPOLLEXIA SERENO, 1986
PARVORDER: HADROSAURIFORMES SERENO, 1997
Superfamily: IGUANODONTIDEA Cope, 1869 (sensu Sereno, 1986)
Family: IGUANODONTIDAE Cope, 1869
*Iguanodon anglicus
_____________________________________________________________________________________
ENGLAND
KENT COUNTY
WEALDEN FORMATION
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
SUPERORDER: SAUROPODOMORPHA HUENE, 1932
GRANDORDER: BRONTOSAURIA OLSHEVSKY, 1991 (emend)
MICRORDER: SAUROPODA MARSH, 1878
ORDER: EUSAUROPODA
SUBORDER: NEOSAUROPODA BONAPARTE, 1986
INFRAORDER: MACRONARIA WILSON & SERENO, 1998
PARVORDER: TITANOSAURIFORMES SALGADO ET AL, 1997
Family: BRACHIOSAURIDAE Riggs, 1904
Subfamily: BRACHIOSAURINAE Riggs, 1904 (sensu Janensch, 1929)
Pelorosaurus conybearei
Pelorosaurus sp
= Ornithopsis sp
_____________________________________________________________________________________
ENGLAND
KENT COUNTY
AN ABANDONED QUARRY
MEMBER OF THE HASTINGS BEDS
NEOCOMIAN SUBEPOCH
LOWER EARLY CRETACEOUS EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
_____________________________________________________________________________________SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: MAMMALIA LINNAEUS, 1758
SUBCLASS: PROTOTHERIA GILL, 1872
INFRACLASS: TRECHNOTHERIA McKENNA, 1975
LEGION: CLADOTHERIA McKENNA, 1975
SUBLEGION: DRYOLESTOIDEA BUTLER, 1939
ORDER: DRYOLESTOIDA PROTHERO, 1981
Family: DRYOLESTIDAE Marsh, 1879
Laolestes hodsoni
= *Melanodon hodsoni
ENGLAND
KENT COUNTY
NEAR FOLKESTONE
CHALK MARL
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
SUPERORDER: EURYOPDA SERENO, 1986
GRANDORDER: ARMATOSAURIA ZHAO, 1983 (emend)
ORDER: ANKYLOSAURIA OSBORN, 1923
SUBORDER: SCELIDOSAURIA COOPER, 1985
INFRAORDER: ANKYLOSAURA OSBORN, 1923
Family: NODOSAURIDAE Marsh, 1890
Acanthopholis sp
_____________________________________________________________________________________
ENGLAND
KENT COUNTY
TURON
MIDDLE CHALK
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
SUPERORDER: TAPINOPLATIA PETERS, 2000
GRANDORDER: CHARACIOPODA PETERS, 2000
MICRORDER: FENESTRASAURIA PETERS, 2000
ORDER: PTEROSAURIA KAUP, 1834
SUBORDER: EUPTERODACTYLOIDEA BENNETT, 1994
Superfamily: ORNITHOCHEIROIDEA Seeley, 1870 (sensu Seeley, 1891)
Family: ORNITHOCHEIRIDAE Seeley, 1870
*Lonchodectes compressirostris
= Pterodactylus compressirostris
= Lonchodectes compressirostris
= Ornithocheirus compressirostrisIncertae sedis
*Ornithocheirus? diomedius
= Osteornis diomedius
= Cimoliornis diomedius
= Ornithocheirus diomedius
= Ornithochirus diomedius
= Pterodactylus diomedius
= Pterodactylus conirostris
Ornithocheirus spSuperfamily: PTERANODONTIDEA Marsh, 1876
Family: PTERANODONTIDAE Marsh, 1876
Incertae sedis
non Ornithocheirus, O. hlavatschi
= Cretornis hlavatschi
= Ornithochirus hlavatschi
_____________________________________________________________________________________
ENGLAND
KENT COUNTY
NEAR DOVER
SHAKESPEARE’S CLIFF
CHALK
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
_____________________________________________________________________________________SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: EURYAPSIDA COLBERT, 1945
SUPERORDER: SAUROPTERYGIA OWEN, 1860
GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994
ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
INFRAORDER: PLIOSAUROMORPHA NOVA
Superfamily: PLIOSAUROIDEA Seeley, 1874
Family: PLIOSAURIDAE Seeley, 1874
Incertae sedis
Crymocetus bernardi
= Plesiosaurus bernardi
= Plesiosaurus bernardi
= Cimoliasaurus bernardi
ENGLAND
SOUTHERN UNITED KINGDOM
ENGLAND
KENT COUNTY?
UNKNOWN LOCALITY
WEST MELBURY MARLY CHALK OR THE ZIG ZAG CHALK FORMATION? (OLD LOWER CHALK)
CENOMANIAN STAGE
UPPER MIDDLE GALLIC SUBEPOCH
LOWER GULF EPOCH
MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: EURYAPSIDA COLBERT, 1945
SUPERORDER: SAUROPTERYGIA OWEN, 1860
GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994
ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
INFRAORDER: PLIOSAUROMORPHA NOVA
Superfamily: PLIOSAUROIDEA Seeley, 1874
Family: PLIOSAURIDAE Seeley, 1874
Incertae sedis
Polyptychodon sp
_____________________________________________________________________________________
SOUTHERN UNITED KINGDOM
ENGLAND
KENT COUNTY
FLOKESTONE
GAULT FORMATION
MIDDLE TO UPPER ALBIAN STAGE
MIDDLE MALM EPOCH
UPPER LOWER CRETACEOUS EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: EURYAPSIDA COLBERT, 1945
SUPERORDER: SAUROPTERYGIA OWEN, 1860
GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994
ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
INFRAORDER: PLIOSAUROMORPHA NOVA
Superfamily: PLIOSAUROIDEA Seeley, 1874
Family: PLIOSAURIDAE Seeley, 1874
Incertae sedis
Polyptychodon sp
_____________________________________________________________________________________
SOUTHERN UNITED KINGDOM
ENGLAND
KENT COUNTY
GRAVESEND
SEAFORD CHALK FORMATION
LOWER CONIACIAN TO MIDDLE SANTONIAN STAGE
LOWER GULF EPOCH
EARLY LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: EURYAPSIDA COLBERT, 1945
SUPERORDER: SAUROPTERYGIA OWEN, 1860
GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994
ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
INFRAORDER: PLIOSAUROMORPHA NOVA
Superfamily: PLIOSAUROIDEA Seeley, 1874
Family: PLIOSAURIDAE Seeley, 1874
Incertae sedis
Polyptychodon sp
_____________________________________________________________________________________
SOUTHERN UNITED KINGDOM
ENGLAND
KENT COUNTY?
HALLING
PLENUS MARLS MEMBER
HOLYWELL NODULAR CHALK FORMATION? (OLD CHALK)
UPPER CENOMANIAN STAGE
UPPER MIDDLE GALLIC SUBEPOCH
LOWERMOST GULF EPOCH
MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: EURYAPSIDA COLBERT, 1945
SUPERORDER: SAUROPTERYGIA OWEN, 1860
GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994
ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
INFRAORDER: PLIOSAUROMORPHA NOVA
Superfamily: PLIOSAUROIDEA Seeley, 1874
Family: PLIOSAURIDAE Seeley, 1874
Incertae sedis
Polyptychodon sp
_____________________________________________________________________________________
SOUTHERN UNITED KINGDOM
ENGLAND
KENT COUNTY?
OFFHAM
CHALK GROUP
?UPPER CRETACEOUS.
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: EURYAPSIDA COLBERT, 1945
SUPERORDER: SAUROPTERYGIA OWEN, 1860
GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994
ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
INFRAORDER: PLIOSAUROMORPHA NOVA
Superfamily: PLIOSAUROIDEA Seeley, 1874
Family: PLIOSAURIDAE Seeley, 1874
Incertae sedis
Polyptychodon sp
_____________________________________________________________________________________
SOUTHERN UNITED KINGDOM
ENGLAND
KENT COUNTY
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: EURYAPSIDA COLBERT, 1945
SUPERORDER: SAUROPTERYGIA OWEN, 1860
GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994
ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
INFRAORDER: PLIOSAUROMORPHA NOVA
Superfamily: PLIOSAUROIDEA Seeley, 1874
Family: PLIOSAURIDAE Seeley, 1874
Incertae sedis
Polyptychodon sp
_____________________________________________________________________________________
ENGLAND
KENT COUNTY
FOLKESTONE
GAULT CLAY
ALBIAN STAGE
MIDDLE GALLIC SUBEPOCH
UPPER EARLY CRETACEOUS EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
_____________________________________________________________________________________SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
Incertae sedis
Gen. sp indet.
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
Incertae sedis
Genus sp indet.
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGANORDER: CRYPTODIRA GAFFNEY & MEYLAN, 1988
SUPERORDER: SELMACRYPTODIRA GAFFNEY, HUTCHISON, JENKINS & MEEKR, 1987
GRANDORDER: DAIOCRYPTODIRA GAFFNEY & MEYLAN, 1988
MICRORDER: CENTROCRYPTODIRA GAFFNEY & MEYLAN, 1988
ORDER: POLYCRPTODIRA GAFFNEY & MEYLAN, 1988
SUBORDER: PROCOELOCRYPTODIRA GAFFNEY & MEYLAN, 1988
INFRAORDER: CHELONOIDEA GAFFNEY & MEYLAN, 1988
Superfamily: CHELONIOIDEA Oppel, 1811
Family: DESMATOCHELIDAE Williston, 1894
Rhinochelys sp
SUBCLASS: DIAPSIDA OSBORN, 1904 OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
NEODIAPSIDA: INCERTAE SEDIS
OR AMNIOTA: INCERTAE SEDIS
MAGANORDER: ICHTHYOPTERYGIA BAUR, 1887 emend nova
SUPERORDER: EOICHTHYOSAURIA MOTANI, 1999
GRANDORDER: ICHTHYOSAURIA EMEND BLAINVILLE, 1835
Incertae sedis
Gen. sp indet.
INFRACLASS: EURYAPSIDA COLBERT, 1945
SUPERORDER: SAUROPTERYGIA OWEN 1860
GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994
ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
Incertae sedis
Gen. sp. indet.
ENGLAND
KENT COUNTY
FOLKESTONE
GAULT
CAMBRIDGE GREENSAND
CENOMANIAN STAGE
UPPERMIDDLE GALLIC SUBEPOCH
LOWERMOST GULF EPOCH
MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
_____________________________________________________________________________________SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
SUPERORDER: TAPINOPLATIA PETERS, 2000
GRANDORDER: CHARACIOPODA PETERS, 2000
MICRORDER: FENESTRASAURIA PETERS, 2000
ORDER: PTEROSAURIA KAUP, 1834
Incertae sedis
Gen. sp indet.
SUBORDER: EUPTERODACTYLOIDEA BENNETT, 1994
Superfamily: ORNITHOCHEIROIDEA Seeley, 1870 (sensu Seeley, 1891)
Family: ORNITHOCHEIRIDAE Seeley, 1870
Incertae sedis
*Ornithocheirus daviesii
= Pterodactylus daviesii
= Lonchodectes daviesii
= Ornithochirus daviesii
Ornithocheirus sp
ENGLAND
KENT COUNTY
FOLKESTONE
EAST WEAR BAY
LOWER UPPER GAULT
ALBIAN STAGE
MIDDLE GALLIC SUBEPOCH
UPPER EARLY CRETACEOUS EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
_____________________________________________________________________________________SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: EURYAPSIDA COLBERT, 1945
SUPERORDER: SAUROPTERYGIA OWEN 1860
GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994
ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
INFRAORDER: PLESIOSAUROMORPHA NOVA
EUPLESIOSAURIA O’KEEFE, 2001
Superfamily: CRYPTOCLEIDOIDEA Williston, 1925
TRICLEDIA O’KEEFE , 2001
Family: CIMOLIASAURIDAE Delair, 1959
Cimoliasaurus australis
= Cimoliasaurus cantabrigiensisSuperfamily: ELASMOSAUROIDEA Cope, 1869
Family: ELASMOSAURIDAE Cope, 1869
*Mauisaurus gardneri
INFRAORDER: PLIOSAUROMORPHA NOVA
Superfamily: PLIOSAUROIDEA Seeley, 1874
Family: PLIOSAURIDAE Seeley, 1874
Polyptychodon interruptus
ENGLAND
KENT COUNTY
LOWER GREENSAND FORMATION
APTIAN STAGE
MIDDLE GALLIC SUBEPOCH
UPPER EARLY CRETACEOUS EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
_____________________________________________________________________________________SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: EURYAPSIDA COLBERT, 1945
SUPERORDER: SAUROPTERYGIA OWEN 1860
GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994
ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
INFRAORDER: PLESIOSAUROMORPHA NOVA
EUPLESIOSAURIA O’KEEFE, 2001
Family: PLESIOSAURIDAE Gray, 1825
Incertae sedis
non Plesiosaurus, *P. latispinus
ENGLAND
KENT COUNTY
CHALK
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGANORDER: CRYPTODIRA GAFFNEY & MEYLAN, 1988
SUPERORDER: SELMACRYPTODIRA GAFFNEY, HUTCHISON, JENKINS & MEEKR, 1987
GRANDORDER: DAIOCRYPTODIRA GAFFNEY & MEYLAN, 1988
MICRORDER: CENTROCRYPTODIRA GAFFNEY & MEYLAN, 1988
ORDER: POLYCRPTODIRA GAFFNEY & MEYLAN, 1988
SUBORDER: PROCOELOCRYPTODIRA GAFFNEY & MEYLAN, 1988
INFRAORDER: CHELONOIDEA GAFFNEY & MEYLAN, 1988
Superfamily: CHELONIOIDEA Oppel, 1811
Family: CHELONIIDAE Oppel, 1811
Subfamily: CHELONINAE Oppel, 1811
Chelone sp
SUBCLASS: DIAPSIDA OSBORN, 1904 OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
NEODIAPSIDA: INCERTAE SEDIS
OR AMNIOTA: INCERTAE SEDIS
MAGANORDER: ICHTHYOPTERYGIA BAUR, 1887 emend nova
SUPERORDER: EOICHTHYOSAURIA MOTANI, 1999
GRANDORDER: ICHTHYOSAURIA EMEND BLAINVILLE, 1835
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
ENGLAND
KENT COUNTY
LOWER CHALK
CENOMANIAN STAGE
UPPER MIDDLE GALLIC SUBEPOCH
LOWERMOST GULF EPOCH
MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: EURYAPSIDA COLBERT, 1945
SUPERORDER: SAUROPTERYGIA OWEN, 1860
GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994
ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
INFRAORDER: PLIOSAUROMORPHA NOVA
Superfamily: PLIOSAUROIDEA Seeley, 1874
Family: PLIOSAURIDAE Seeley, 1874
Polyptychodon interruptus
_____________________________________________________________________________________
ENGLAND
KENT COUNTY
MIDDLE CHALK
TURONIAN STAGE
UPPER GALLIC SUBEPOCH
EARLY GULF EPOCH
EARLY LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGANORDER: CRYPTODIRA GAFFNEY & MEYLAN, 1988
SUPERORDER: SELMACRYPTODIRA GAFFNEY, HUTCHISON, JENKINS & MEEKR, 1987
GRANDORDER: DAIOCRYPTODIRA GAFFNEY & MEYLAN, 1988
MICRORDER: CENTROCRYPTODIRA GAFFNEY & MEYLAN, 1988
ORDER: POLYCRPTODIRA GAFFNEY & MEYLAN, 1988
SUBORDER: PROCOELOCRYPTODIRA GAFFNEY & MEYLAN, 1988
INFRAORDER: CHELONOIDEA GAFFNEY & MEYLAN, 1988
Superfamily: CHELONIOIDEA Oppel, 1811
Family: CHELONIIDAE Oppel, 1811
Subfamily: CHELONINAE Oppel, 1811
*Chelone camperi
Superfamily: DERMOCHELYOIDEA Willams, 1950
Family: PROTOSTEGIDAE Cope, 1889
Subfamily: CHELOSPHARGINAE Zangerl, 1953
Cimochelys benstedi
= Chelone (Cimochelys) benstedi
= Rhinochelys benstedi
= Emys benstedi
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: EURYAPSIDA COLBERT, 1945
SUPERORDER: SAUROPTERYGIA OWEN, 1860
GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994
ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
INFRAORDER: PLIOSAUROMORPHA NOVA
Superfamily: PLIOSAUROIDEA Seeley, 1874
Family: PLIOSAURIDAE Seeley, 1874
Polyptychodon interruptus
_____________________________________________________________________________________
ENGLAND
KENT COUNTY
CHALK MARL
CENOMANIAN STAGE
UPPER MIDDLE GALLIC SUBEPOCH
LOWERMOST GULF EPOCH
MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: EURYAPSIDA COLBERT, 1945
SUPERORDER: SAUROPTERYGIA OWEN, 1860
GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994
ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
INFRAORDER: PLIOSAUROMORPHA NOVA
Superfamily: PLIOSAUROIDEA Seeley, 1874
Family: PLIOSAURIDAE Seeley, 1874
Polyptychodon interruptus
Incertae sedis
non Polyptychodon, P. continus
= Polyptychodon mackesoni
Polyptychodon sp
_____________________________________________________________________________________
ENGLAND
KENT COUNTY
UPPER CHALK
CENOMANIAN STAGE
MIDDLE GALLIC SUBEPOCH
LOWERMOST GULF EPOCH
MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGANORDER: CRYPTODIRA GAFFNEY & MEYLAN, 1988
SUPERORDER: SELMACRYPTODIRA GAFFNEY, HUTCHISON, JENKINS & MEEKR, 1987
GRANDORDER: DAIOCRYPTODIRA GAFFNEY & MEYLAN, 1988
MICRORDER: CENTROCRYPTODIRA GAFFNEY & MEYLAN, 1988
ORDER: POLYCRPTODIRA GAFFNEY & MEYLAN, 1988
SUBORDER: PROCOELOCRYPTODIRA GAFFNEY & MEYLAN, 1988
INFRAORDER: CHELONOIDEA GAFFNEY & MEYLAN, 1988
Superfamily: CHELONIOIDEA Oppel, 1811
Family: CHELONIIDAE Oppel, 1811
Subfamily: CHELONINAE Oppel, 1811
Chelone camperi
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: EURYAPSIDA COLBERT, 1945
SUPERORDER: SAUROPTERYGIA OWEN, 1860
GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994
ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
INFRAORDER: PLIOSAUROMORPHA NOVA
Superfamily: PLIOSAUROIDEA Seeley, 1874
Family: PLIOSAURIDAE Seeley, 1874
Polyptychodon interruptus
_____________________________________________________________________________________
ENGLAND
KENT COUNTY
UPPER CHALK
TURONIAN STAGE
UPPERMOST GALLIC SUBEPOCH
LOWER GULF EPOCH
MIDDLE CRETACEOUS -CAMPANIAN STAGE
MIDDLE SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: EURYAPSIDA COLBERT, 1945
SUPERORDER: SAUROPTERYGIA OWEN, 1860
GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994
ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
INFRAORDER: PLESIOSAUROMORPHA NOVA
EUPLESIOSAURIA O’KEEFE, 2001
Superfamily: CRYPTOCLEIDOIDEA Williston, 1925
TRICLEDIA O’KEEFE , 2001
Family: CIMOLIASAURIDAE Delair, 1959
*Cimoliasaurus smithi
_____________________________________________________________________________________
ENGLAND
KENT COUNTY
BURHAM CHALKPIT
CHALK
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGANORDER: CRYPTODIRA GAFFNEY & MEYLAN, 1988
SUPERORDER: SELMACRYPTODIRA GAFFNEY, HUTCHISON, JENKINS & MEEKR, 1987
GRANDORDER: DAIOCRYPTODIRA GAFFNEY & MEYLAN, 1988
MICRORDER: CENTROCRYPTODIRA GAFFNEY & MEYLAN, 1988
ORDER: POLYCRPTODIRA GAFFNEY & MEYLAN, 1988
SUBORDER: PROCOELOCRYPTODIRA GAFFNEY & MEYLAN, 1988
INFRAORDER: CHELONOIDEA GAFFNEY & MEYLAN, 1988
Superfamily: CHELONIOIDEA Oppel, 1811
Family: CHELONIIDAE Oppel, 1811
Subfamily: CHELONINAE Oppel, 1811
Chelone sp
_____________________________________________________________________________________
ENGLAND
KENT COUNTY
CHALK PIT AT BURHAM
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: EURYAPSIDA COLBERT, 1945
SUPERORDER: SAUROPTERYGIA OWEN 1860
GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994
ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
INFRAORDER: PLESIOSAUROMORPHA NOVA
EUPLESIOSAURIA O’KEEFE, 2001
Family: PLESIOSAURIDAE Gray, 1825
Incertae sedis
non Plesiosaurus, *P. simthi
Plesiosaurus sp
_____________________________________________________________________________________
ENGLAND
KENT COUNTY
PLENUS MARLS
CENOMANIAN STAGE
UPPER MIDDLE GALLIC SUBEPOCH
LOWERMOST GULF EPOCH
MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: EURYAPSIDA COLBERT, 1945
SUPERORDER: SAUROPTERYGIA OWEN, 1860
GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994
ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
INFRAORDER: PLIOSAUROMORPHA NOVA
Superfamily: PLIOSAUROIDEA Seeley, 1874
Family: PLIOSAURIDAE Seeley, 1874
Polyptychodon interruptus
_____________________________________________________________________________________
ENGLAND
KENT COUNTY
BURHAM
CULAND PITS
LOWER CHALK-UPPER CHALK
CENOMANIAN-TURONIAN STAGE
UPPER GALLIC SUBEPOCH
LOWER GULF EPOCH
MIDDLE-LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
Incertae sedis
Genus sp indet.
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGANORDER: CRYPTODIRA GAFFNEY & MEYLAN, 1988
SUPERORDER: SELMACRYPTODIRA GAFFNEY, HUTCHISON, JENKINS & MEEKR, 1987
GRANDORDER: DAIOCRYPTODIRA GAFFNEY & MEYLAN, 1988
MICRORDER: CENTROCRYPTODIRA GAFFNEY & MEYLAN, 1988
ORDER: POLYCRPTODIRA GAFFNEY & MEYLAN, 1988
SUBORDER: PROCOELOCRYPTODIRA GAFFNEY & MEYLAN, 1988
INFRAORDER: CHELONOIDEA GAFFNEY & MEYLAN, 1988
Superfamily: CHELONIOIDEA Oppel, 1811
Family: CHELONIIDAE Oppel, 1811
Incertae sedis
Gen sp indet.
Subfamily: CHELONINAE Oppel, 1811
Chelone camperi
Chelone sp
= Chelone/Lytolma spSuperfamily: DERMOCHELYOIDEA Willams, 1950
Family: PROTOSTEGIDAE Cope, 1889
Subfamily: CHELOSPHARGINAE Zangerl, 1953
*Cimochelys benstedi
Subfamily: PROTOSTEGINAE Cope, 1889
Tribe: PROTOSTEGINI Cope, 1889 (sensu Hooks, 1998)
Subtribe: PROTOSTEGINA Cope, 1889 (sensu Hooks, 1998)
Protostega sp
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: EURYAPSIDA COLBERT, 1945
SUPERORDER: SAUROPTERYGIA OWEN, 1860
GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994
ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
INFRAORDER: PLESIOSAUROMORPHA NOVA
EUPLESIOSAURIA O’KEEFE, 2001
Superfamily: CRYPTOCLEIDOIDEA Williston, 1925
TRICLEDIA O’KEEFE , 2001
Family: CIMOLIASAURIDAE Delair, 1959
Cimoliasaurus smithi
INFRAORDER: PLIOSAUROMORPHA NOVA
Superfamily: PLIOSAUROIDEA Seeley, 1874
Family: PLIOSAURIDAE Seeley, 1874
Polyptychodon interruptus
SUPERLEGION: NEODIAPSIDA BENTON, 1985LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
SUPERORDER: TAPINOPLATIA PETERS, 2000
GRANDORDER: CHARACIOPODA PETERS, 2000
MICRORDER: FENESTRASAURIA PETERS, 2000
ORDER: PTEROSAURIA KAUP, 1834
SUBORDER: EUPTERODACTYLOIDEA BENNETT, 1994
Superfamily: PTERANODONTIDEA Marsh, 1876
Incertae sedis
*Cimoliopterus cuvieri = Anhanguera cuvieri
= Pterodactylus cuvieri
= Coloborhynchus cuvieri
= Ornithochirus cuvieri
= Ptenodactylus cuvieriSuperfamily: ORNITHOCHEIROIDEA Seeley, 1870 (sensu Seeley, 1891)
Family: ORNITHOCHEIRIDAE Seeley, 1870
Incertae sedis
non Ornithocheirus, *O. giganteus
= Pterodactylus giganteus
= Lonchodectes giganteus
= Ornithochirus giganetus
= Ornithodesmus giganteus
_____________________________________________________________________________________
ENGLAND
KENT COUNTY
BURHAM
CULAND PITS
ZIG-ZAG CHALK FORMATION [SOUTH DOWNS NOMENCLATURE] (LOWER CHALK, ABBOTTS CLIFF CHALK FORMATION [NORTH DOWNS NOMENCLATURE]), LOWER CHALK-UPPER CHALK.
CALYCOCERAS GUERANGERI ZONE
UPPER CENOMANIAN STAGE
UPPERMOST GALLIC SUBEPOCH
LOWER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: LACERTILIA OWEN, 1842
GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961
MICRORDER: SCINCOMORPHA CAMP, 1923
ORDER: ANGUIMORPHA FURBRINGER, 1900
SUBORDER: DIPLOGLOSSA COPE, 1864
INFRAORDER: THECAGLOSSA COPE, 1890
PARVORDER: MONSTERSAURIA NORELL & GAO, 1977
Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)
Family: MOSASAURIDAE Gervais, 1853
Subfamily: DOLICHOSAURIDAE Gervais, 1852
Tribe: DOLICHOSAURINI Gervais, 1852
*Dolichosaurus longicollis
Infrafamily: NATANIA
Subfamily: MOSASAURINAE Gervais, 1853 (sensu Williston, 1897)
Tribe: MOSASAURINI Gervais, 1853 (sensu Russell, 1967)
Mosasaurus gracilis
_____________________________________________________________________________________
ENGLAND
KENT COUNTY
BURHAM
LOWER CHALK
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
SUPERORDER: TAPINOPLATIA PETERS, 2000
GRANDORDER: CHARACIOPODA PETERS, 2000
MICRORDER: FENESTRASAURIA PETERS, 2000
ORDER: PTEROSAURIA KAUP, 1834
SUBORDER: EUPTERODACTYLOIDEA BENNETT, 1994
Superfamily: PTERANODONTIDEA Marsh, 1876
Family: PTERANODONTIDAE Marsh, 1876
Incertae sedis
non Ornithocheirus, O. hlavatschi
= Cretornis hlavatschi
= Ornithochirus hlavatschi
_____________________________________________________________________________________
ENGLAND
KENT COUNTY
SNODLAND
CHALK
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
SUPERORDER: TAPINOPLATIA PETERS, 2000
GRANDORDER: CHARACIOPODA PETERS, 2000
MICRORDER: FENESTRASAURIA PETERS, 2000
ORDER: PTEROSAURIA KAUP, 1834
SUBORDER: EUPTERODACTYLOIDEA BENNETT, 1994
Superfamily: PTERANODONTIDEA Marsh, 1876
Family: PTERANODONTIDAE Marsh, 1876
Incertae sedis
non Ornithocheirus, O. hlavatschi
= Cretornis hlavatschi
= Ornithochirus hlavatschi
_____________________________________________________________________________________
ENGLAND
KENT COUNTY
WHITE CHALK
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
SUPERORDER: TAPINOPLATIA PETERS, 2000
GRANDORDER: CHARACIOPODA PETERS, 2000
MICRORDER: FENESTRASAURIA PETERS, 2000
ORDER: PTEROSAURIA KAUP, 1834
SUBORDER: EUPTERODACTYLOIDEA BENNETT, 1994
Superfamily: ORNITHOCHEIROIDEA Seeley, 1870 (sensu Seeley, 1891)
Family: ANHANGUERIDAE Campos & Kellner 1985
Anhanguera cuvieri
= Pterodactylus cuvieri
= Coloborhynchus cuvieri
= Ornithochirus cuvieri
= Ptenodactylus cuvieri
_____________________________________________________________________________________
ENGLAND
KENT COUNTY
HALLING
LOWER CHALK
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
SUPERORDER: TAPINOPLATIA PETERS, 2000
GRANDORDER: CHARACIOPODA PETERS, 2000
MICRORDER: FENESTRASAURIA PETERS, 2000
ORDER: PTEROSAURIA KAUP, 1834
SUBORDER: EUPTERODACTYLOIDEA BENNETT, 1994
Superfamily: ORNITHOCHEIROIDEA Seeley, 1870 (sensu Seeley, 1891)
Family: ANHANGUERIDAE Campos & Kellner 1985
Anhanguera cuvieri
= Pterodactylus cuvieri
= Coloborhynchus cuvieri
= Ornithochirus cuvieri
= Ptenodactylus cuvieri
_____________________________________________________________________________________
ENGLAND
KENT COUNTY
CUXTON
MIDDLE CHALK
MIDDLE TO LATE TURONIAN STAGE
LOWER SENONIAN SUBEPOCH
MIDDLE GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: LACERTILIA OWEN, 1842
GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961
MICRORDER: SCINCOMORPHA CAMP, 1923
ORDER: ANGUIMORPHA FURBRINGER, 1900
SUBORDER: DIPLOGLOSSA COPE, 1864
INFRAORDER: THECAGLOSSA COPE, 1890
PARVORDER: MONSTERSAURIA NORELL & GAO, 1977
Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)
Family: MOSASAURIDAE Gervais, 1853
Infrafamily: NATANIA
Subfamily: MOSASAURINAE Gervais, 1853 (sensu Williston, 1897)
Tribe: MOSASAURINI Gervais, 1853 (sensu Russell, 1967)
Mosasaurus gracilis
_____________________________________________________________________________________
ENGLAND
KENT COUNTY
EAST WEAR BAY
LOWER-UPPER GAULT
ALBIAN STAGE
LOWER GALLIC SUBEPOCH
UPPER EARLY CRETACEOUS EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: NEODIAPSIDA BENTON, 1985
NEODIAPSIDA: INCERTAE SEDIS
OR AMNIOTA: INCERTAE SEDIS
MAGANORDER: ICHTHYOPTERYGIA BAUR, 1887 emend nova
SUPERORDER: EOICHTHYOSAURIA MOTANI, 1999
GRANDORDER: ICHTHYOSAURIA EMEND BLAINVILLE, 1835
MICRORDER: LONGIPINNATI HUENE, 1948
ORDER: MERRIAMOSAURIA MONTANI, 1999
SUBORDER: EUICHTHYOSAURIA MONTANI, 1999
INFRAORDER: PARVIPELVIA MONTANI, 1999
PARVORDER: NEOICHTHYOSAURIA SANDER, 2000
SUPERFAMILY: OPHTHALMOSAURIDEA SEELEY, 1874
Family: OPTHALMOSAURIDAE Seeley, 1874
Subfamily: PLATYPTERYGIINAE Arkhangelsky 2011
Pervushovisaurus campylodon = Platypterygius campylodon
= Ichthyosaurus campylodon
= Myopterygius campylodonPlatypterygius sp
_____________________________________________________________________________________
ENGLAND
KENT COUNTY
TOWARDS FOLKESTONE
END OF THE ROUND DOWN TUNNEL
GREY LOWER CHALK
CENOMANIAN STAGE
GALLIC SUBEPOCH
LOWERMOST GULF EPOCH
MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: NEODIAPSIDA BENTON, 1985
NEODIAPSIDA: INCERTAE SEDIS
OR AMNIOTA: INCERTAE SEDIS
MAGANORDER: ICHTHYOPTERYGIA BAUR, 1887 emend nova
SUPERORDER: EOICHTHYOSAURIA MOTANI, 1999
GRANDORDER: ICHTHYOSAURIA EMEND BLAINVILLE, 1835
MICRORDER: LONGIPINNATI HUENE, 1948
ORDER: MERRIAMOSAURIA MONTANI, 1999
SUBORDER: EUICHTHYOSAURIA MONTANI, 1999
INFRAORDER: PARVIPELVIA MONTANI, 1999
PARVORDER: NEOICHTHYOSAURIA SANDER, 2000
SUPERFAMILY: OPHTHALMOSAURIDEA SEELEY, 1874
Family: OPTHALMOSAURIDAE Seeley, 1874
Subfamily: PLATYPTERYGIINAE Arkhangelsky 2011
Pervushovisaurus campylodon = Platypterygius campylodon
= Ichthyosaurus campylodon
= Myopterygius campylodon
_____________________________________________________________________________________
ENGLAND
KENT COUNTY
NEAR HYTHE
LOWER GREENSAND
APTIAN STAGE
GALLIC SUBEPOCH
EARLY CRETACEOUS EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
SUPERORDER: SAUROPODOMORPHA HUENE, 1932
GRANDORDER: BRONTOSAURIA OLSHEVSKY, 1991 (emend)
MICRORDER: SAUROPODA MARSH, 1878
ORDER: EUSAUROPODA
SUBORDER: NEOSAUROPODA BONAPARTE, 1986
INFRAORDER: MACRONARIA WILSON & SERENO, 1998
PARVORDER: TITANOSAURIFORMES SALGADO ET AL, 1997
Family: BRACHIOSAURIDAE Riggs, 1904
Incertae sedis
*Dinodocus mackesoni
= Pelorosaurus mackesoni
_____________________________________________________________________________________
ENGLAND
KENT COUNTY
NEAR HYTHE
CHALK
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: EURYAPSIDA COLBERT, 1945
SUPERORDER: SAUROPTERYGIA OWEN, 1860
GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994
ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
INFRAORDER: PLIOSAUROMORPHA NOVA
Superfamily: PLIOSAUROIDEA Seeley, 1874
Family: PLIOSAURIDAE Seeley, 1874
Polyptychodon interruptus
_____________________________________________________________________________________
ENGLAND
KENT COUNTY
NEAR HYTHE
UPPER GREENSAND
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: EURYAPSIDA COLBERT, 1945
SUPERORDER: SAUROPTERYGIA OWEN, 1860
GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994
ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
INFRAORDER: PLIOSAUROMORPHA NOVA
Superfamily: PLIOSAUROIDEA Seeley, 1874
Family: PLIOSAURIDAE Seeley, 1874
Polyptychodon interruptus
Incertae sedis
non Polyptychodon, P. continus
= *Polyptychodon mackesoni
_____________________________________________________________________________________
ENGLAND
KENT COUNTY
HYTHE
LOWER GREENSAND
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: EURYAPSIDA COLBERT, 1945
SUPERORDER: SAUROPTERYGIA OWEN, 1860
GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994
ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
INFRAORDER: PLIOSAUROMORPHA NOVA
Superfamily: PLIOSAUROIDEA Seeley, 1874
Family: PLIOSAURIDAE Seeley, 1874
Incertae sedis
Polyptychodon sp
___________________________________________________________________________________
ENGLAND
KENT COUNTY
LIDDEN SPUT
CHALK
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: LACERTILIA OWEN, 1842
GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961
MICRORDER: SCINCOMORPHA CAMP, 1923
ORDER: ANGUIMORPHA FURBRINGER, 1900
SUBORDER: DIPLOGLOSSA COPE, 1864
INFRAORDER: THECAGLOSSA COPE, 1890
PARVORDER: MONSTERSAURIA NORELL & GAO, 1977
Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)
Family: MOSASAURIDAE Gervais, 1853
Subfamily: DOLICHOSAURIDAE Gervais, 1852
Tribe: DOLICHOSAURINI Gervais, 1852
Dolichosaurus longicollis
_____________________________________________________________________________________
ENGLAND
KENT COUNTY
GRAVESAND
UPPER CHALK
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: LACERTILIA OWEN, 1842
GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961
MICRORDER: SCINCOMORPHA CAMP, 1923
ORDER: ANGUIMORPHA FURBRINGER, 1900
SUBORDER: DIPLOGLOSSA COPE, 1864
INFRAORDER: THECAGLOSSA COPE, 1890
PARVORDER: MONSTERSAURIA NORELL & GAO, 1977
Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)
Family: MOSASAURIDAE Gervais, 1853
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
ENGLAND
KENT COUNTY
BURHAM
WOULDHAM
PETER’S PIT
LOWER CHALK
HOLBASTER SUBGLOBOSUS ZONE
CENOMANIAN STAGE
UPPER GALLIC SUBEPOCH
LOWER GULF EPOCH
MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: LACERTILIA OWEN, 1842
GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961
MICRORDER: SCINCOMORPHA CAMP, 1923
ORDER: ANGUIMORPHA FURBRINGER, 1900
SUBORDER: DIPLOGLOSSA COPE, 1864
INFRAORDER: THECAGLOSSA COPE, 1890
PARVORDER: MONSTERSAURIA NORELL & GAO, 1977
Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)
Family: MOSASAURIDAE Gervais, 1853
Subfamily: DOLICHOSAURIDAE Gervais, 1852
Tribe: DOLICHOSAURINI Gervais, 1852
Coniasaurus crassidens
_____________________________________________________________________________________
ENGLAND
KENT COUNTY
BURHAM
LOWER CHALK
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGANORDER: CRYPTODIRA GAFFNEY & MEYLAN, 1988
SUPERORDER: SELMACRYPTODIRA GAFFNEY, HUTCHISON, JENKINS & MEEKR, 1987
GRANDORDER: DAIOCRYPTODIRA GAFFNEY & MEYLAN, 1988
MICRORDER: CENTROCRYPTODIRA GAFFNEY & MEYLAN, 1988
ORDER: POLYCRPTODIRA GAFFNEY & MEYLAN, 1988
SUBORDER: PROCOELOCRYPTODIRA GAFFNEY & MEYLAN, 1988
INFRAORDER: CHELONOIDEA GAFFNEY & MEYLAN, 1988
Superfamily: DERMOCHELYOIDEA Willams, 1950
Family: PROTOSTEGIDAE Cope, 1889
Subfamily: CHELOSPHARGINAE Zangerl, 1953
Cimochelys benstedi
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ENGLAND
KENT COUNTY
EAST WEAR BAY
LOWER-UPPER GAULT
ALBIAN STAGE
MIDDLE GALLIC SUBEPOCH
UPPERMOST EARLY CRETACEOUS EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGANORDER: CRYPTODIRA GAFFNEY & MEYLAN, 1988
SUPERORDER: SELMACRYPTODIRA GAFFNEY, HUTCHISON, JENKINS & MEEKR, 1987
GRANDORDER: DAIOCRYPTODIRA GAFFNEY & MEYLAN, 1988
MICRORDER: CENTROCRYPTODIRA GAFFNEY & MEYLAN, 1988
ORDER: POLYCRPTODIRA GAFFNEY & MEYLAN, 1988
SUBORDER: PROCOELOCRYPTODIRA GAFFNEY & MEYLAN, 1988
INFRAORDER: CHELONOIDEA GAFFNEY & MEYLAN, 1988
Superfamily: CHELONIOIDEA Oppel, 1811
Family: DESMATOCHELIDAE Williston, 1894
Rhinochelys pulchriceps
= Chelone pulchriceps
Rhinochelys spSuperfamily: DERMOCHELYOIDEA Willams, 1950
Family: PROTOSTEGIDAE Cope, 1889
Subfamily: CHELOSPHARGINAE Zangerl, 1953
Cimochelys benstedi
_____________________________________________________________________________________
ENGLAND
KENT COUNTY
CHARING
HART HILL
CENOMANIAN STAGE
UPPER GALLIC SUBEPOCH
LOWER GULF EPOCH
MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: LACERTILIA OWEN, 1842
GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961
MICRORDER: SCINCOMORPHA CAMP, 1923
ORDER: ANGUIMORPHA FURBRINGER, 1900
SUBORDER: DIPLOGLOSSA COPE, 1864
INFRAORDER: THECAGLOSSA COPE, 1890
PARVORDER: MONSTERSAURIA NORELL & GAO, 1977
Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)
Family: MOSASAURIDAE Gervais, 1853
Subfamily: DOLICHOSAURIDAE Gervais, 1852
Tribe: DOLICHOSAURINI Gervais, 1852
Coniasaurus crassidens
_____________________________________________________________________________________
ENGLAND
KENT COUNTY
CHALK
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA DUMERIL & BIBRON, 1839
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA OPPEL, 1811
SUPERORDER: LACERTILIA OWEN, 1842
Incertae sedis
*Raphiosaurus no species given
_____________________________________________________________________________________
ENGLAND
KENT COUNTY
UPPER CHALK
SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: LACERTILIA OWEN, 1842
GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961
MICRORDER: SCINCOMORPHA CAMP, 1923
ORDER: ANGUIMORPHA FURBRINGER, 1900
SUBORDER: DIPLOGLOSSA COPE, 1864
INFRAORDER: THECAGLOSSA COPE, 1890
PARVORDER: MONSTERSAURIA NORELL & GAO, 1977
Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)
Family: MOSASAURIDAE Gervais, 1853
Infrafamily: NATANIA
Subfamily: TYLOSAURINAE Williston, 1897
Tylosaurus sp
_____________________________________________________________________________________
ENGLAND
KENT COUNTY
HART HILL
CHALK
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: LACERTILIA OWEN, 1842
GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961
MICRORDER: SCINCOMORPHA CAMP, 1923
ORDER: ANGUIMORPHA FURBRINGER, 1900
SUBORDER: DIPLOGLOSSA COPE, 1864
INFRAORDER: THECAGLOSSA COPE, 1890
PARVORDER: MONSTERSAURIA NORELL & GAO, 1977
Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)
Family: MOSASAURIDAE Gervais, 1853
Subfamily: DOLICHOSAURIDAE Gervais, 1852
Tribe: DOLICHOSAURINI Gervais, 1852
Coniasaurus crassidens
_____________________________________________________________________________________
ENGLAND
KENT COUNTY
WOULDHAM
PETER’S QUARRIES
LOWER PART OF THE GREY CHALK FORMATION
CENOMANIAN-TURONIAN STAGE
UPPER GALLIC SUBEPOCH
LOWER GULF EPOCH
MIDDLE-LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
Incertae sedis
Genus sp indet.
_____________________________________________________________________________________
ENGLAND
KENT COUNTY
MAIDSTONE
KENTISH RAG
GREEN-SAND FORMATION
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
Incertae sedis
*Protemys serrata
_____________________________________________________________________________________
ENGLAND
KENT COUNTY
MAIDSTONE
UPPER CHALK
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: LACERTILIA OWEN, 1842
GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961
MICRORDER: SCINCOMORPHA CAMP, 1923
ORDER: ANGUIMORPHA FURBRINGER, 1900
SUBORDER: DIPLOGLOSSA COPE, 1864
INFRAORDER: THECAGLOSSA COPE, 1890
PARVORDER: MONSTERSAURIA NORELL & GAO, 1977
Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)
Family: MOSASAURIDAE Gervais, 1853
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
ENGLAND
KENT COUNTY
NEAR MAIDSTONE
IGUANODON QUARRY
LOWER GREENSAND
APTIAN-LOWERMOST ALBIAN STAGE
MIDDLE GALLIC SUBEPOCH
UPPER EARLY CRETACEOUS EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: EURYAPSIDA COLBERT, 1945
SUPERORDER: SAUROPTERYGIA OWEN 1860
GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994
ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
INFRAORDER: PLESIOSAUROMORPHA NOVA
EUPLESIOSAURIA O’KEEFE, 2001
Family: PLESIOSAURIDAE Gray, 1825
Incertae sedis
non Plesiosaurus, P. latispinus
_____________________________________________________________________________________
ENGLAND
KENT COUNTY
NEAR MAIDSTONE
KENTISH RAG
LOWER GREENSAND FORMTION
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
SUBORDER: ANKYLOPOLLEXIA SERENO, 1986
PARVORDER: HADROSAURIFORMES SERENO, 1997
Superfamily: IGUANODONTIDEA Cope, 1869 (sensu Sereno, 1986)
Family: IGUANODONTIDAE Cope, 1869
Mantellodon carpenteri
= *Iguanodon mantelli
= Therosaurus mantelli
= Iguanodon mantelliIncertae sedis
Gen. sp indet.
_____________________________________________________________________________________
ENGLAND
KENT COUNTY
NEAR MAIDSTONE
CHALK
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
SUPERORDER: TAPINOPLATIA PETERS, 2000
GRANDORDER: CHARACIOPODA PETERS, 2000
MICRORDER: FENESTRASAURIA PETERS, 2000
ORDER: PTEROSAURIA KAUP, 1834
SUBORDER: EUPTERODACTYLOIDEA BENNETT, 1994
Incertae sedis
Gen sp indet.
_____________________________________________________________________________________
ENGLAND
KENT COUNTY
WOULDHAM
PETER’S PIT
GREY CHALK FORMATION
HOLASTER SUBGLOBOSUS ZONE
CENOMANIAN STAGE
UPPER MIDDLE GALLIC SUBEPOCH
LOWERMOST GULF EPOCH
MIDDLE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
SUPERORDER: TAPINOPLATIA PETERS, 2000
GRANDORDER: CHARACIOPODA PETERS, 2000
MICRORDER: FENESTRASAURIA PETERS, 2000
ORDER: PTEROSAURIA KAUP, 1834
SUBORDER: EUPTERODACTYLOIDEA BENNETT, 1994
Superfamily: ORNITHOCHEIROIDEA Seeley, 1870 (sensu Seeley, 1891)
Family: ORNITHOCHEIRIDAE Seeley, 1870
Incertae sedis
Ornithocheirus sp
_____________________________________________________________________________________
ENGLAND
KENT COUNTY
ISLE OF SHEPPEY
LONDON CLAY
EARLY EOCENE EPOCH
PALEOGENE PERIOD
LOWER TERTIARY SUBERA
LOWER CENOZOIC ERA
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
ORDER: EUSUCHIA emend HUXLEY, 1875
INFRAORDER: CROCODYLIA emend GMELIN, 1877
Family: GAVIALIDAE Cuvier, 1807
Subfamily: THORACOSAURINAE Cope, 1875 (sensu Nopcsa, 1928)
Eosuchus sp
PARVORDER: BREVIROSTRES
Superfamily: CROCODILOIDEA Cuvier, 1807 (sensu Fitzinger, 1826)
Family: CROCODYLIDAE Cuvier, 1807
Subfamily: TOMISTOMINAE Eastman, 1902 (sensu Kalin, 1955)
*Kentisuchus spenceri
= Crocodylus, *C. spenceri
= Kentisuchus toliapicus
= Crocodilus toliapicus
= *Crocodilus delucii
= *Crocodilus champsoides
= *Crocodilus arduini
_____________________________________________________________________________________
ENGLAND
KENT COUNTY
QUARRY AT SOUTHBOROUGH
HASTINGS SANDS
LOWER WEALDEN
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
Incertae sedis
*Thecospondylus horneri
_____________________________________________________________________________________