SOUTHERN UNITED KINGDOM

ENGLAND

SUSSEX COUNTY

FRESHFIELDS QUARRY

WEALDEN

BARREMIAN STAGE

LOWER GALLIC SUBEPOCH

MIDDLE EARLY CRETACOEUS EPOCH

EARLY CRETACEOUS

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: ORNITHISCHIFORMES COOPER, 1985

MAGNORDER: ORNITHISCHIA SEELEY, 1888

SUPERORDER: EURYOPDA SERENO, 1986

GRANDORDER: ARMATOSAURIA ZHAO, 1983 (emend)

ORDER: ANKYLOSAURIA OSBORN, 1923

SUBORDER: SCELIDOSAURIA COOPER, 1985

INFRAORDER: ANKYLOSAURA OSBORN, 1923

Family: POLACANTHIDAE Weiland, 1911

Polacanthus foxii

_____________________________________________________________________________________

ENGLAND

SUSSEX COUNTY

BURNWASH

WEALDEN

VALANGINIAN STAGE

MIDDLE NEOCOMIAN SUBEPOCH

EARLY CRETACEOUS EPOCH

EARLY CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: CHELONOMORPHA KUHN, 1960

SUPERCOHORT: TESTUDINES LINNAEUS, 1758

COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988

MAGANORDER: CRYPTODIRA GAFFNEY & MEYLAN, 1988

ORDER: CRYPTODIRA COPE, 1868

GRANDORDER: DAIOCRYPTODIRA GAFFNEY & MEYLAN, 1988

MICRORDER: EUCRYPTODIRA GAFFNEY, 1975

Family: PLESIOCHELYIDAE Rutimeyer, 1873

Hylaeochelys belli

_____________________________________________________________________________________

ENGLAND

SUSSEX COUNTY

CHUCKFIELD

WEALDEN

VALANGINIAN STAGE

MIDDLE NEOCOMIAN SUBEPOCH

EARLY CRETACEOUS EPOCH

EARLY CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: CHELONOMORPHA KUHN, 1960

SUPERCOHORT: TESTUDINES LINNAEUS, 1758

COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988

MAGANORDER: CRYPTODIRA GAFFNEY & MEYLAN, 1988

ORDER: CRYPTODIRA COPE, 1868

GRANDORDER: DAIOCRYPTODIRA GAFFNEY & MEYLAN, 1988

MICRORDER: EUCRYPTODIRA GAFFNEY, 1975

Family: PLESIOCHELYIDAE Rutimeyer, 1873

Hylaeochelys belli

_____________________________________________________________________________________

SOUTHERN UNITED KINGDOM

ENGLAND

SUSSEX COUNTY

BEXHILL

UPPER TURNBRIDGE WELLS  FORMATION

HAUTERVIAN STAGE

LOWER NEOCOMIAN SUBEPOCH

LOWER EARLY CRETACEOUS EPOCH

EARLY CRETACEOUS

_____________________________________________________________________________________

SOUTHERN UNITED KINGDOM

ENGLAND

SUSSEX COUNTY

BEXHILL

WADHURST CLAY FORMATION

HAUTERVIAN OR VALANGINIAN STAGE

LOWER-MIDDLE NEOCOMIAN SUBEPOCH

LOWER EARLY CRETACEOUS EPOCH

EARLY CRETACEOUS

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: ORNITHISCHIFORMES COOPER, 1985

MAGNORDER: ORNITHISCHIA SEELEY, 1888

SUPERORDER: EURYOPDA SERENO, 1986

GRANDORDER: ARMATOSAURIA ZHAO, 1983 (emend)

ORDER: ANKYLOSAURIA OSBORN, 1923

SUBORDER: SCELIDOSAURIA COOPER, 1985

INFRAORDER: ANKYLOSAURA OSBORN, 1923

Family: POLACANTHIDAE Weiland, 1911

Polacanthus foxii

_____________________________________________________________________________________

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: CHELONOMORPHA KUHN, 1960

SUPERCOHORT: TESTUDINES LINNAEUS, 1758

COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988

MAGANORDER: CRYPTODIRA GAFFNEY & MEYLAN, 1988

SUPERORDER: SELMACRYPTODIRA GAFFNEY, HUTCHISON, JENKINS & MEEKR, 1987

GRANDORDER: DAIOCRYPTODIRA GAFFNEY & MEYLAN, 1988

MICRORDER: CENTROCRYPTODIRA GAFFNEY & MEYLAN, 1988

ORDER: POLYCRPTODIRA GAFFNEY & MEYLAN, 1988

SUBORDER: PROCOELOCRYPTODIRA GAFFNEY & MEYLAN, 1988

INFRAORDER: CHELONOIDEA GAFFNEY & MEYLAN, 1988

Superfamily: EUCHELONIOIDEA Gaffney & Meylan, 1988

Family: CHELONIIDAE Oppel, 1811

Subfamily: CHELONINAE Oppel, 1811

Chelone sp

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: LACERTILIA OWEN, 1842

GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961

MICRORDER: SCINCOMORPHA CAMP, 1923

ORDER: ANGUIMORPHA FURBRINGER, 1900

SUBORDER: DIPLOGLOSSA COPE, 1864

INFRAORDER: THECAGLOSSA COPE, 1890

PARVORDER: MONSTERSAURIA NORELL & GAO, 1977

Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)

Family: MOSASAURIDAE Gervais, 1853

Incertae sedis

Gen. sp indet.

Infrafamily: NATANIA

Subfamily: RUSSELLOSAURINAE Bell, 1993

Tribe: PLIOPLATECARPINI Dollo, 1884 (sensu Russell, 1967)

Platecarpus sp

Gen. sp indet.

Subfamily: MOSASAURINAE Gervais, 1853 (sensu Williston, 1897)

Tribe: MOSASAURINI Gervais, 1853 (sensu Russell, 1967)

Clidastes sp

Tribe: LEIODONTINI Lingham-Soliar, 1993

Leiodon anceps

SUSSEX COUNTY

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

INFRACLASS: EURYAPSIDA COLBERT, 1945

SUPERORDER: SAUROPTERYGIA OWEN, 1860

GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994

ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835

SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835

INFRAORDER: PLIOSAUROMORPHA NOVA

Superfamily: PLIOSAUROIDEA Seeley, 1874

Family: PLIOSAURIDAE Seeley, 1874

Incertae sedis

Polyptychodon sp

_____________________________________________________________________________________

SUSSEX COUNTY

CHALK

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

INFRACLASS: EURYAPSIDA COLBERT, 1945

SUPERORDER: SAUROPTERYGIA OWEN, 1860

GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994

ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835

SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835

INFRAORDER: PLIOSAUROMORPHA NOVA

Superfamily: PLIOSAUROIDEA Seeley, 1874

Family: PLIOSAURIDAE Seeley, 1874

Polyptychodon interruptus

_____________________________________________________________________________________

SUSSEX COUNTY

LOWER CHALK MARL

CENOMANIAN STAGE

UPPER MIDDLE GALLIC SUBEPOCH

LOWERMOST GULF EPOCH

MIDDLE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

INFRACLASS: EURYAPSIDA COLBERT, 1945

SUPERORDER: SAUROPTERYGIA OWEN, 1860

GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994

ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835

SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835

INFRAORDER: PLIOSAUROMORPHA NOVA

Superfamily: PLIOSAUROIDEA Seeley, 1874

Family: PLIOSAURIDAE Seeley, 1874

Polyptychodon interruptus

Incertae sedis

non Polyptychodon, P. continus
= Polyptychodon mackesoni

_____________________________________________________________________________________

ENGLAND

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

INFRACLASS: EURYAPSIDA COLBERT, 1945

SUPERORDER: SAUROPTERYGIA OWEN, 1860

GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994

ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835

SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835

INFRAORDER: PLIOSAUROMORPHA NOVA

Superfamily: PLIOSAUROIDEA Seeley, 1874

Family: PLIOSAURIDAE Seeley, 1874

Polyptychodon interruptus

SUSSEX

WEALDEN FORMATION

EARLY CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990

COHORT: CROCODYLOTARSI BENTON & CLARK, 1988

MAGNORDER: SUCHIA KREBS, 1974

SUPERORDER: CROCODYLOMORPHA WALKER, 1968

GRANDORDER: CROCODILIFORMES HAY, 1930

MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983

ORDER: MESOSUCHIA EMEND HUXLEY, 1875

PARVORDER: METASUCHIA BENTON & CLARK, 1988

INFRAORDER: METAMESOSUCHIA HULKE, 1878

NEOSUCHIA CLARK, IN BENTON & CLARK, 1988

Family: GONIOPHOLIDIDAE Cope, 1875

Goniopholis crassidens

_____________________________________________________________________________________

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990

COHORT: CROCODYLOTARSI BENTON & CLARK, 1988

MAGNORDER: SUCHIA KREBS, 1974

SUPERORDER: CROCODYLOMORPHA WALKER, 1968

GRANDORDER: CROCODILIFORMES HAY, 1930

MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983

ORDER: MESOSUCHIA EMEND HUXLEY, 1875

PARVORDER: METASUCHIA BENTON & CLARK, 1988

INFRAORDER: METAMESOSUCHIA HULKE, 1878

NEOSUCHIA CLARK, IN BENTON & CLARK, 1988

Family: GONIOPHOLIDIDAE Cope, 1875

Goniopholis crassidens

SUSSEX COUNTY

UPPER CHALK PT AT HOUGHTON

CHALK

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

INFRACLASS: EURYAPSIDA COLBERT, 1945

SUPERORDER: SAUROPTERYGIA OWEN 1860

GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994

ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835

SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835

INFRAORDER: PLESIOSAUROMORPHA NOVA

EUPLESIOSAURIA O’KEEFE, 2001

Family: PLESIOSAURIDAE Gray, 1825

Incertae sedis

non Plesiosaurus, P. pachyomus

_____________________________________________________________________________________

SUSSEX COUNTY

UPPER CHALK MARL

TURONIAN STAGE

UPPERMOST GALLIC SUBEPOCH

LOWER GULF EPOCH-CAMPANIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

INFRACLASS: EURYAPSIDA COLBERT, 1945

SUPERORDER: SAUROPTERYGIA OWEN, 1860

GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994

ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835

SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835

INFRAORDER: PLIOSAUROMORPHA NOVA

Superfamily: PLIOSAUROIDEA Seeley, 1874

Family: PLIOSAURIDAE Seeley, 1874

Polyptychodon interruptus

Incertae sedis

Crymocetus bernardi
= Plesiosaurus bernardi
= Plesiosaurus bernardi
= Cimoliasaurus bernardi

_____________________________________________________________________________________

SUSSEX COUNTY

NEAR LEWES

BERWICK

BERWICK BRICK PIT (CUCKMERE BRICK COMPANY)

UPPER WEALD CLAY

WEALDEN GROUP

BARREMIAN STAGE

UPPER NEOCOMIAN SUBEPOCH

LOWER EARLY CRETACEOUS EPOCH

EARLY CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

INFRACLASS: EURYAPSIDA COLBERT, 1945

SUPERORDER: SAUROPTERYGIA OWEN 1860

GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994

ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835

SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835

INFRAORDER: PLIOSAUROMORPHA NOVA

Superfamily: PLIOSAUROIDEA Seeley, 1874

Family: LEPTOCLEIDIDAE White, 1940

*Leptocleidus superstes

Note: Lagoonal or close inshore deposits.

_____________________________________________________________________________________

SUSSEX COUNTY

NEAR ARUNDEL

HOUGHTON

UPPER CHALK

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

INFRACLASS: EURYAPSIDA COLBERT, 1945

SUPERORDER: SAUROPTERYGIA OWEN, 1860

GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994

ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835

Incertae sedis

Gen. sp indet.

SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835

INFRAORDER: PLIOSAUROMORPHA NOVA

Superfamily: PLIOSAUROIDEA Seeley, 1874

Family: PLIOSAURIDAE Seeley, 1874

Incertae sedis

non Plesiosaurus, P. bernardi

_____________________________________________________________________________________

SUSSEX COUNTY

FAIRLIGHT COVE

ASHDOWN BEDS

UPPER BERRIASIAN STAGE

LOWER NEOCOMIAN SUBEPOCH

LOWER EARLY CRETACEOUS EPOCH

EARLY CRETACEOUS

ICHNOSTEM: TETRAPODIPIDA NOVA

ICHNOSUPERCLASS: AMNIOTAIPIDII NOVA

ICHNOCLASS: REPTILIPEDIA VIALOV, 1966

ICHNOSUBCLASS: DIAPSIDIPIDII NOVA

ICHNOSTEM-GROUP: NEODIAPSIDIPEDII NOVA

ICHNOINFRACLASS: NEODIAPSIDIPEDII NOVA

ICHNOSUBDIVISION: ARCHOSAURIPEDIDII NOVA

ICHNOLEGION: DINOSAUROMORPHIPEDA NOVA

ICHNOSUBLEGION: DINOSAURIFORMEIPEDIDII NOVA

ICHNOSUPERCOHORT: DINOSAUROPEDIDA NOVA

ICHNOCOHORT: THEROPODIPEDIA NOVA

Incerate sedis

Gen. sp indet.

ICHNOSUPERORDER: NEOTHEROPODOPEDIDA NOVA

ICHNOORDER: NEOTETANURIPEDIDII NOVA

ICHNOSUBORDER: AVETHEROPODIPIDA NOVA

ICHNOMICRORDER: COELUROSAURIPIDIDA NOVA

Incertae sedis

Gen. sp indet.

ICHNOCOHORT: ORNITHISCHIFORMEIPIDA NOVA

ICHNOMAGNORDER: ORNITHISCHIPIDA NOVA

ICHNOSUPERORDER: NEORNITHISCHIPIDA NOVA

ICHNOGRANDORDER: ORNITHOPODIPDIA NOVA

ICHNOMICRORDER: EUORNITHOPODIPIDA NOVA

ICHNOORDER: IGUANODONTIPIDIA NOVA

ICHNOSUBORDER: ANKYLOPOLLEXIPEDIDA NOVA

ICHNOINFRAORDER: STYRACOSTERNIPIDA NOVA

ICHNOPARVORDER: HADROSAURIFORMIPEDIDA NOVA

Ichnosuperfamily: IGUANODONTIDEA Nova

Ichnofamily: IGUANODONTIPODIDAE emend Vylov, 1988

Gen. sp indet.

_____________________________________________________________________________________

SUSSEX COUNTY

BEXHILL

COODEN BEACH RAILWAY STATION

COODEN BEACH

INTERTIDAL LEVEL

TUNBRIDGE WELLS SAND

LATE VALANGINIAN STAGE

MIDDLE NEOCOMIAN SUBEPOCH

EARLY CRETACEOUS EPOCH

EARLY CRETACEOUS

ICHNOSTEM: TETRAPODIPIDA NOVA

ICHNOSUPERCLASS: AMNIOTAIPIDII NOVA

ICHNOCLASS: REPTILIPEDIA VIALOV, 1966

ICHNOSUBCLASS: DIAPSIDIPIDII NOVA

ICHNOSTEM-GROUP: NEODIAPSIDIPEDII NOVA

ICHNOINFRACLASS: NEODIAPSIDIPEDII NOVA

ICHNOSUBDIVISION: ARCHOSAURIPEDIDII NOVA

ICHNOLEGION: DINOSAUROMORPHIPEDA NOVA

ICHNOSUBLEGION: DINOSAURIFORMEIPEDIDII NOVA

ICHNOSUPERCOHORT: DINOSAUROPEDIDA NOVA

ICHNOCOHORT: ORNITHISCHIFORMEIPIDA NOVA

ICHNOMAGNORDER: ORNITHISCHIPIDA NOVA

ICHNOSUPERORDER: NEORNITHISCHIPIDA NOVA

ICHNOGRANDORDER: ORNITHOPODIPDIA NOVA

ICHNOMICRORDER: EUORNITHOPODIPIDA NOVA

ICHNOORDER: IGUANODONTIPIDIA NOVA

ICHNOSUBORDER: ANKYLOPOLLEXIPEDIDA NOVA

ICHNOINFRAORDER: STYRACOSTERNIPIDA NOVA

ICHNOPARVORDER: HADROSAURIFORMIPEDIDA NOVA

Ichnosuperfamily: IGUANODONTIDEA Nova

Ichnofamily: IGUANODONTIPODIDAE emend Vylov, 1988

Gen. sp indet.

_____________________________________________________________________________________

SUSSEX COUNTY COUNTY

UPPER CHALK

TURONIAN STAGE

UPPERMOST GALLIC SUBEPOCH

LOWER GULF EPOCH-CAMPANIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

INFRACLASS: EURYAPSIDA COLBERT, 1945

SUPERORDER: SAUROPTERYGIA OWEN, 1860

GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994

ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835

SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835

INFRAORDER: PLIOSAUROMORPHA NOVA

Superfamily: PLIOSAUROIDEA Seeley, 1874

Family: PLIOSAURIDAE Seeley, 1874

Incertae sedis

*Crymocetus bernardi
= Plesiosaurus bernardi
= Plesiosaurus bernardi
= Cimoliasaurus bernardi

_____________________________________________________________________________________

ENGLAND

SUSSEX COUNTY

NEAR LEWES

SCADDLESCOMBE CHALK-PIT

CHALK

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

INFRACLASS: EURYAPSIDA COLBERT, 1945

SUPERORDER: SAUROPTERYGIA OWEN, 1860

GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994

ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835

SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835

INFRAORDER: PLIOSAUROMORPHA NOVA

Superfamily: PLIOSAUROIDEA Seeley, 1874

Family: PLIOSAURIDAE Seeley, 1874

Incertae sedis

Crymocetus bernardi
= Plesiosaurus bernardi
= Plesiosaurus bernardi
= Cimoliasaurus bernardi

_____________________________________________________________________________________

ENGLAND

SUSSEX COUNTY

SOUTHERAM

CHALK

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

INFRACLASS: EURYAPSIDA COLBERT, 1945

SUPERORDER: SAUROPTERYGIA OWEN, 1860

GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994

ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835

SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835

INFRAORDER: PLIOSAUROMORPHA NOVA

Superfamily: PLIOSAUROIDEA Seeley, 1874

Family: PLIOSAURIDAE Seeley, 1874

Incertae sedis

Crymocetus bernardi
= Plesiosaurus bernardi
= Plesiosaurus bernardi
= Cimoliasaurus bernardi

_____________________________________________________________________________________

ENGLAND

SUSSEX COUNTY

STEYNING

CHALK

EARLY CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

INFRACLASS: EURYAPSIDA COLBERT, 1945

SUPERORDER: SAUROPTERYGIA OWEN 1860

GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994

ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835

SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835

INFRAORDER: PLESIOSAUROMORPHA NOVA

EUPLESIOSAURIA O’KEEFE, 2001

Superfamily: ELASMOSAUROIDEA Cope, 1869

Family: ELASMOSAURIDAE Cope, 1869

Mauisaurus gardneri
= Plesiosaurus constrictus

_____________________________________________________________________________________

ENGLAND

SUSSEX COUNTY

STEYNING

CHALK-PIT

MIDDLE CHALK

TURONIAN STAGE

UPPERMOST GALLIC SUBEPOCH

LOWER GULF EPOCH

MIDDLE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

INFRACLASS: EURYAPSIDA COLBERT, 1945

SUPERORDER: SAUROPTERYGIA OWEN 1860

GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994

ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835

SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835

INFRAORDER: PLESIOSAUROMORPHA NOVA

EUPLESIOSAURIA O’KEEFE, 2001

Family: PLESIOSAURIDAE Gray, 1825

Incertae sedis

non Plesiosaurus, *P. constrictus

_____________________________________________________________________________________

ENGLAND

SUSSEX COUNTY

SOUTHERAM

CHALK

TURONIAN STAGE

UPPERMOST GALLIC SUBEPOCH

LOWER GULF EPOCH

MIDDLE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

INFRACLASS: EURYAPSIDA COLBERT, 1945

SUPERORDER: SAUROPTERYGIA OWEN 1860

GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994

ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835

SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835

INFRAORDER: PLESIOSAUROMORPHA NOVA

EUPLESIOSAURIA O’KEEFE, 2001

Family: PLESIOSAURIDAE Gray, 1825

Incertae sedis

non Plesiosaurus, P. constrictus

_____________________________________________________________________________________

ENGLAND

PROBABLY SUSSEX COUNTY

UNKNOWN

WEALDEN FORMATION

EARLY CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: ORNITHISCHIFORMES COOPER, 1985

MAGNORDER: ORNITHISCHIA SEELEY, 1888

SUPERORDER: NEORNITHISCHIA COOPER, 1985

GRANDORDER: ORNITHOPODA MARSH, 1871

MICRORDER: EUORNITHOPODA SERENO, 1986

ORDER: IGUANODONTIA DOLLO, 1888

SUBORDER: ANKYLOPOLLEXIA SERENO, 1986

PARVORDER: HADROSAURIFORMES SERENO, 1997

Superfamily: IGUANODONTIDEA Cope, 1869 (sensu Sereno, 1986)

Family: IGUANODONTIDAE Cope, 1869

Iguanodon anglicus
= Iguanodon mantelli
= Therosaurus mantelli
= Iguanodon mantelli

_____________________________________________________________________________________

ENGLAND

SOUTH COAST OF SUSSEX COUNTY

WEALDEN FORMATION

EARLY CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: ORNITHISCHIFORMES COOPER, 1985

MAGNORDER: ORNITHISCHIA SEELEY, 1888

SUPERORDER: NEORNITHISCHIA COOPER, 1985

GRANDORDER: ORNITHOPODA MARSH, 1871

MICRORDER: EUORNITHOPODA SERENO, 1986

ORDER: IGUANODONTIA DOLLO, 1888

SUBORDER: ANKYLOPOLLEXIA SERENO, 1986

PARVORDER: HADROSAURIFORMES SERENO, 1997

Superfamily: IGUANODONTIDEA Cope, 1869 (sensu Sereno, 1986)

Family: IGUANODONTIDAE Cope, 1869

Iguanodon anglicus
= Iguanodon mantelli
= Therosaurus mantelli
= Iguanodon mantelli

_____________________________________________________________________________________

ENGLAND

SUSSEX COUNTY

WEALDEN FORMATION

EARLY CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

Incertae sedis

Gen. sp indet.

COHORT: THEROPODA EMEND MARSH, 1881

Incertae sedis

Gen. sp indet.

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: TETANURAE GAUTHIER, 1986

ORDER: SPINOSAURIA OLSHEVSKY, 1991

Family: BARYONYCHIDAE Charig & Milner, 1986

*Suchosaurus cultridens

ORDER: NEOTETANURAE SERENO, WILSON, LARSSON, DUTHEIL & SUES, 1994

SUBORDER: CARNOSAURIA HUENE, 1920

Superfamily: Incertae sedis

Family: MEGALOSAURIDAE Huxley, 1889

Megalosaurus bucklandii

COHORT: ORNITHISCHIFORMES COOPER, 1985

SUPERORDER: SAUROPODOMORPHA HUENE, 1932

GRANDORDER: BRONTOSAURIA OLSHEVSKY, 1991 (emend)

MICRORDER: SAUROPODA MARSH, 1878

Incertae sedis

Gen. sp indet.

ORDER: EUSAUROPODA

SUBORDER: NEOSAUROPODA BONAPARTE, 1986

INFRAORDER: MACRONARIA WILSON & SERENO, 1998

PARVORDER: TITANOSAURIFORMES SALGADO ET AL, 1997

Family: BRACHIOSAURIDAE Riggs, 1904

Subfamily: BRACHIOSAURINAE Riggs, 1904 (sensu Janensch, 1929)

Pelorosaurus conybearei
= Pleurocelus valdensis
= Astrodon valdensis
= Astrodon (Pleurocelus) valdensis

MAGNORDER: ORNITHISCHIA SEELEY, 1888

SUPERORDER: NEORNITHISCHIA COOPER, 1985

GRANDORDER: ORNITHOPODA MARSH, 1871

MICRORDER: EUORNITHOPODA SERENO, 1986

ORDER: IGUANODONTIA DOLLO, 1888

SUBORDER: ANKYLOPOLLEXIA SERENO, 1986

PARVORDER: HADROSAURIFORMES SERENO, 1997

Superfamily: IGUANODONTIDEA Cope, 1869 (sensu Sereno, 1986)

Family: IGUANODONTIDAE Cope, 1869

Iguanodon anglicus
= Iguanodon mantelli
= Therosaurus mantelli
= Iguanodon mantelli
Iguanodon sp

SUPERORDER: EURYOPDA SERENO, 1986

GRANDORDER: ARMATOSAURIA ZHAO, 1983 (emend)

ORDER: ANKYLOSAURIA OSBORN, 1923

SUBORDER: SCELIDOSAURIA COOPER, 1985

INFRAORDER: ANKYLOSAURA OSBORN, 1923

Family: POLACANTHIDAE Weiland, 1911

Hylaeosaurus armatus
= Hylaeosaurus oweni

GRANDORDER: STEGOSAUROMORPHA COOPER, 1985 (EMEND)

ORDER: STEGOSAURIA MARSH, 1877

Incertae sedis

Gen. sp indet.

____________________________________________________________________________________

ENGLAND

SUSSEX COUNTY

HEATHFIELD

UPPER WEALDEN FORMATION

EARLY CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: ORNITHISCHIFORMES COOPER, 1985

MAGNORDER: ORNITHISCHIA SEELEY, 1888

SUPERORDER: NEORNITHISCHIA COOPER, 1985

GRANDORDER: ORNITHOPODA MARSH, 1871

MICRORDER: EUORNITHOPODA SERENO, 1986

ORDER: IGUANODONTIA DOLLO, 1888

SUBORDER: DRYOMORPHA SERNEO, 1986

Family: DRYOSAURIDAE Milner & Norman, 1984

Valdosaurus canaliculatus
= Dryosaurus canaliculatus

_____________________________________________________________________________________

ENGLAND

SUSSEX COUNTY

RIVERMORE

KNELLSONTE

WEALDEN FORMATION

EARLY CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: ORNITHISCHIFORMES COOPER, 1985

MAGNORDER: ORNITHISCHIA SEELEY, 1888

SUPERORDER: NEORNITHISCHIA COOPER, 1985

GRANDORDER: ORNITHOPODA MARSH, 1871

MICRORDER: EUORNITHOPODA SERENO, 1986

ORDER: IGUANODONTIA DOLLO, 1888

SUBORDER: ANKYLOPOLLEXIA SERENO, 1986

PARVORDER: HADROSAURIFORMES SERENO, 1997

Superfamily: IGUANODONTIDEA Cope, 1869 (sensu Sereno, 1986)

Family: IGUANODONTIDAE Cope, 1869

Mantellisaurus atherfieldensis
= Iguanodon? atherfieldensis

_____________________________________________________________________________________

ENGLAND

SUSSEX COUNTY

TETHAM

WEALDEN FORMATION

EARLY CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: ORNITHISCHIFORMES COOPER, 1985

MAGNORDER: ORNITHISCHIA SEELEY, 1888

SUPERORDER: NEORNITHISCHIA COOPER, 1985

GRANDORDER: ORNITHOPODA MARSH, 1871

MICRORDER: EUORNITHOPODA SERENO, 1986

ORDER: IGUANODONTIA DOLLO, 1888

SUBORDER: ANKYLOPOLLEXIA SERENO, 1986

PARVORDER: HADROSAURIFORMES SERENO, 1997

Superfamily: IGUANODONTIDEA Cope, 1869 (sensu Sereno, 1986)

Family: IGUANODONTIDAE Cope, 1869

Mantellisaurus atherfieldensis
= Iguanodon? atherfieldensis
= *Cetiosaurus brachyurus

____________________________________________________________________________________

ENGLAND

SUSSEX COUNTY

TUNBRIDGE WELLS

UPPER WEALDEN FORMATION

EARLY CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: TETANURAE GAUTHIER, 1986

ORDER: NEOTETANURAE SERENO, WILSON, LARSSON, DUTHEIL & SUES, 1994

SUBORDER: AVETHEROPODA PAUL, 1988

Superfamily: ALLOSAUROIDEA Marsh, 1878 (sensu Currie & Zhao, 1994)

Family: ALLOSAURIDAE Marsh, 1878

Incertae sedis

Becklespinax altispinax
= Acrocanthosaurus altispinax

_____________________________________________________________________________________