CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
3 KM WEST OF THE ROYAL TYRRELL MUSEUM OF PALEONTOLOGY
HORSETHIEF MEMBER
HORSESHOE CAYON FORMATION
EDMONTONAIN AGE
LATE CAMPANIAN-EARLY MAASTRICHTIAN STAGE
UPPERMOST SENONIAN SUBEPOCH
UPPER UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA PAUL, 2001
AVEPECTORA PAUL, 2001
SUBORDER: MANIRAPTORA GAUTHIER, 1986
INFRAORDER: OVIRAPTOROSAURIA BARSBOLD, 1976
Family: CAENAGNATHIDAE Sternberg, 1940
*Apatoraptor pennatus
_____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
PREHISTORIC PARK BONEBED
HORSESHOE CANYON FORMATION
EDMONTON GROUP
EDMONTONIAN AGE
UPPERMOST CAMPANIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
SUBORDER: ANKYLOPOLLEXIA SERENO, 1986
INFRAORDER: STYRACOSTERNA SERENO, 1986
SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)
Family: HADROSAURIDAE COPE, 1869
Subfamily: HADROSAURINAE Cope, 1869 (sensu Lambe, 1918)
Tribe: EDMONTOSAURINI Brett-Surman vide Chapman & Brett-Surman, 1990
Edmontosaurus regalis
_____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
FOX COULEE BONEBED
HORSESHOE CANYON FORMATION
EDMONTON GROUP
EDMONTONIAN AGE
UPPERMOST CAMPANIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
SUBORDER: ANKYLOPOLLEXIA SERENO, 1986
INFRAORDER: STYRACOSTERNA SERENO, 1986
SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)
Family: HADROSAURIDAE COPE, 1869
Subfamily: HADROSAURINAE Cope, 1869 (sensu Lambe, 1918)
Tribe: EDMONTOSAURINI Brett-Surman vide Chapman & Brett-Surman, 1990
Edmontosaurus regalis
_____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
EAST BANK OF THE RED DEER RIVER
BETWEEN MORRIN AND THE TOLMAN BRIDGE
HORSESHOE CANYON FORMATION
EDMONTON GROUP
EDMONTONIAN AGE
LOWER MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: MARGINOCEPHALIA SERENO, 1986
ORDER: CERATOPIA MARSH, 1890
SUBORDER: NEOCERATOPIA SERENO, 1986
INFRAORDER: CORONOSAURIA SERENO, 1986
PARVORDER: CERATOPOMORPHA Wolfe & Kirkland, 1998 (emend)
Superfamily: CERATOPOIDEA Marsh, 1888 (sensu Hay, 1902)
Family: CERATOPIDAE Marsh, 1888
Subfamily: CERATOPIINAE Marsh, 1888 (sensu Abel, 1919)
Tribe: CHASMOSAURINI Lambe, 1915
Anchiceratops ornatus
_____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
RED DEER RIVER
BUFFALO JUMP PROVINCIAL PARK
DRY ISLAND
HORSESHOE CANYON FORMAITON
EDMONTON GROUP
EDMONTONIAN AGE
LOWER MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
SUBORDER: ORNITHOMIMOSAURIA BARSBOLD, 1976
Family: ORNITHOMIMIDAE Marsh, 1890
Gen. sp indet.
____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
RED DEER RIVER
ABOUT 1 M NW OF THE MOUTH OF BIG VALLEY CREEK
BUFFALO JUMP
DRY ISLAND
HORSESHOE CANYON FORMAITON
EDMONTON GROUP
EDMONTONIAN AGE
LOWER MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA PAUL, 2002
SUBORDER: COELURIA MARSH, 1881
Superfamily: TYRANNOSAUROIDEA Osborn, 1906 (sensu Bonaparte et. al., 1990)
Family: TYRANNOSAURIDAE Osborn, 1906
Subfamily: ALBERTOSAURINAE Currie, Hurum & Sabath, 2003
Albertosaurus sarcophagus
AVEPECTORA PAUL, 2002
SUBORDER: METORNITHES PERLE, NORELL, CHIAPPE & CLARK, 1993
INFRAORDER: ALVAREZSAURIA BONAPARTE, 1991
Family: ALVAREZSAURIDAE Bonaparte, 1991
*Albertonykus borealis
____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
NEAR DRUMHELLER
TOLMAN BRIDGE
WESTERN (RIGHT) BANK OF RED DEER RIVER
HORSESHOE CANYON FORMATION
EDMONTON GROUP
EDMONTONIAN AGE
LOWER MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: EURYAPSIDA COLBERT, 1945
SUPERORDER: SAUROPTERYGIA OWEN 1860
GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994
ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
INFRAORDER: PLESIOSAUROMORPHA NOVA
EUPLESIOSAURIA O’KEEFE, 2001
Superfamily: ELASMOSAUROIDEA nova
Family: ELASMOSAURIDAE Cope, 1869
Incertae sedis
Gen. sp indet.
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
SUBORDER: ANKYLOPOLLEXIA SERENO, 1986
INFRAORDER: STYRACOSTERNA SERENO, 1986
SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)
Family: HADROSAURIDAE COPE, 1869
Subfamily: HADROSAURINAE Cope, 1869 (sensu Lambe, 1918)
Tribe: EDMONTOSAURINI Brett-Surman vide Chapman & Brett-Surman, 1990
Edmontosaurus sp?
_____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
NEAR DRUMHELLER
EASTERN BANK OF RED DEER RIVER AT ABOUT THE MIDPOINT BETWEEN THE TOLMAN AND MORRING BRIDGE
HORSESHOE CANYON FORMATION
EDMONTON GROUP
EDMONTONIAN AGE
LOWER MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: EURYAPSIDA COLBERT, 1945
SUPERORDER: SAUROPTERYGIA OWEN 1860
GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994
ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
INFRAORDER: PLESIOSAUROMORPHA NOVA
EUPLESIOSAURIA O’KEEFE, 2001
Superfamily: ELASMOSAUROIDEA nova
Family: ELASMOSAURIDAE Cope, 1869
Incertae sedis
*Leurospondylus ultimus
_____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
ABOUT 5 M S OF TOLMAN BRIDGE
HORSESHOE CANYON FORMATION
EDMONTON GROUP
EDMONTONIAN AGE
LOWER MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: EURYOPDA SERENO, 1986
GRANDORDER: ARMATOSAURIA ZHAO, 1983 (emend)
ORDER: ANKYLOSAURIA OSBORN, 1923
INFRAORDER: ANKYLOSAURA OSBORN, 1923
Family: ANKYLOSAURIDAE Brown, 1908
Subfamily: ANKYLOSAURINAE Brown, 1908
Euoplocephalus tutus
= Stereocephalus tutus
= Palaeoscincus tutus
_____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
3.5 MILES BELOW TOLMAN FERRY
EAST BANK OF THE RED DEER RIVER
HORSESHOE CANYON FORMATION (LOWER EDMONTON FORMATION)
EDMONTON GROUP
EDMONTONIAN AGE
LOWER MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: MARGINOCEPHALIA SERENO, 1986
ORDER: CERATOPIA MARSH, 1890
SUBORDER: NEOCERATOPIA SERENO, 1986
INFRAORDER: CORONOSAURIA SERENO, 1986
Family: PROTOCERATOPIDAE Granger & Gregory, 1923
Subfamily: LEPTOCERATOPINAE Nopcsa, 1923
Montanoceratops cerorhynchus
_____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
RED DEER RIVER
NEAR THE TOLMAN FERRY
HORSESHOE CANYON FORMATION
EDMONTON GROUP
EDMONTONIAN AGE
LOWER MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
SUBORDER: ANKYLOPOLLEXIA SERENO, 1986
INFRAORDER: STYRACOSTERNA SERENO, 1986
SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)
Family: HADROSAURIDAE COPE, 1869
Subfamily: HADROSAURINAE Cope, 1869 (sensu Lambe, 1918)
Tribe: SAUROLOPHINI Nopcsa, 1917 (sensu Brett-Surman vide Chapman & Brett-Surman, 1990)
*Saurolophus osborni
Saurolophus sp
____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
TOLMAN FERRY
"EDMONTON FORMATION"
EDMONTON GROUP
EDMONTONIAN AGE
LOWER MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
SUBORDER: ANKYLOPOLLEXIA SERENO, 1986
INFRAORDER: STYRACOSTERNA SERENO, 1986
SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)
Family: HADROSAURIDAE COPE, 1869
Incertae sedis
Gen. sp indet.
____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
RED DEER RIVER
30 FT ABOVE RIVER AT TOLMAN FERRY
HORSESHOE CANYON FORMATION
EDMONTON GROUP
EDMONTONIAN AGE
LOWER MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
SUBORDER: ANKYLOPOLLEXIA SERENO, 1986
INFRAORDER: STYRACOSTERNA SERENO, 1986
SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)
Family: HADROSAURIDAE COPE, 1869
Subfamily: LAMBEOSAURINAE Parks, 1923
Tribe: CORYTHOSAURINI Brett-Surman vide Chapman & Brett-Surman, 1990
Hypacrosaurus altispinus
____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
RED DEER RIVER
NEAR TOLMAN FERRY
HORSESHOE CANYON (EDMONTON?) FORMATION
EDMONTON GROUP
EDMONTONIAN AGE
LOWER MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
SUBORDER: ANKYLOPOLLEXIA SERENO, 1986
INFRAORDER: STYRACOSTERNA SERENO, 1986
SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)
Family: HADROSAURIDAE COPE, 1869
Incertae sedis
Gen. sp indet.
Subfamily: LAMBEOSAURINAE Parks, 1923
Tribe: CORYTHOSAURINI Brett-Surman vide Chapman & Brett-Surman, 1990
Hypacrosaurus altispinus
____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
LEFT BANK OF RED DEER RIVER
1 1/2 M ABOVE TOLMAN
A SANDSTONE NEAR THE TOP OF THE BEDS
SCOLLARD FORMATION
UPPERMOST EDMONTON GROUP
LANCIAN AGE
UPPERMOST MAASTRICHTIAN STAGE
UPPERMOST SENONIAN SUBEPOCH
UPPER UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
SUBORDER: ANKYLOPOLLEXIA SERENO, 1986
INFRAORDER: STYRACOSTERNA SERENO, 1986
SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)
Family: HADROSAURIDAE COPE, 1869
Incertae sedis
Gen. sp indet.
SUPERORDER: EURYOPDA SERENO, 1986
GRANDORDER: ARMATOSAURIA ZHAO, 1983 (emend)
ORDER: ANKYLOSAURIA OSBORN, 1923
INFRAORDER: ANKYLOSAURA OSBORN, 1923
Family: ANKYLOSAURIDAE Brown, 1908
Subfamily: ANKYLOSAURINAE Brown, 1908
Ankylosaurus magniventris
____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
RED DEER RIVER
2 M ABOVE TOLMAN FERRY
HORSESHOE CANYON FORMATION
EDMONTON GROUP
EDMONTONIAN AGE
LOWER MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
SUBORDER: ANKYLOPOLLEXIA SERENO, 1986
INFRAORDER: STYRACOSTERNA SERENO, 1986
SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)
Family: HADROSAURIDAE COPE, 1869
Subfamily: LAMBEOSAURINAE Parks, 1923
Tribe: CORYTHOSAURINI Brett-Surman vide Chapman & Brett-Surman, 1990
Hypacrosaurus altispinus
____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
190 FT ABOVE THE RED DEER RIVER
3 M ABOVE TOLMAN
SCOLLARD FORMATION
UPPERMOST EDMONTON GROUP
LANCIAN AGE
UPPERMOST MAASTRICHTIAN STAGE
UPPERMOST SENONIAN SUBEPOCH
UPPER UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: MARGINOCEPHALIA SERENO, 1986
ORDER: CERATOPIA MARSH, 1890
SUBORDER: NEOCERATOPIA SERENO, 1986
INFRAORDER: CORONOSAURIA SERENO, 1986
Family: PROTOCERATOPIDAE Granger & Gregory, 1923
Subfamily: LEPTOCERATOPINAE Nopcsa, 1923
*Leptoceratops gracilis
_____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
RED DEER RIVER
3 M ABOVE TOLMAN FERRY
HORSESHOE CANYON (EDMONTON?) FORMATION
EDMONTON GROUP
EDMONTONIAN AGE
LOWER MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
SUBORDER: ANKYLOPOLLEXIA SERENO, 1986
INFRAORDER: STYRACOSTERNA SERENO, 1986
SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)
Family: HADROSAURIDAE COPE, 1869
Incertae sedis
Gen. sp indet.
____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
RED DEER RIVER
4 M ABOVE TOLMAN FERRY
HORSESHOE CANYON FORMATION
EDMONTON GROUP
EDMONTONIAN AGE
LOWER MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
SUBORDER: ANKYLOPOLLEXIA SERENO, 1986
INFRAORDER: STYRACOSTERNA SERENO, 1986
SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)
Family: HADROSAURIDAE COPE, 1869
Subfamily: LAMBEOSAURINAE Parks, 1923
Tribe: CORYTHOSAURINI Brett-Surman vide Chapman & Brett-Surman, 1990
*Hypacrosaurus altispinus
____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
EAST SIDE OF RED DEER RIVER
5 M ABOVE TOLMAN FERRY
HORSESHOE CANYON FORMATION
EDMONTON GROUP
EDMONTONIAN AGE
LOWER MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
SUBORDER: ANKYLOPOLLEXIA SERENO, 1986
INFRAORDER: STYRACOSTERNA SERENO, 1986
SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)
Family: HADROSAURIDAE COPE, 1869
Subfamily: LAMBEOSAURINAE Parks, 1923
Tribe: CORYTHOSAURINI Brett-Surman vide Chapman & Brett-Surman, 1990
Hypacrosaurus altispinus
= *Cheneosaurus tolmanensis (Juveinle)
____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
7 M BELOW TOLMAN FERRY
HORSESHOE CANYON FORMATION
EDMONTON GROUP
EDMONTONIAN AGE
LOWER MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: MARGINOCEPHALIA SERENO, 1986
ORDER: CERATOPIA MARSH, 1890
SUBORDER: NEOCERATOPIA SERENO, 1986
INFRAORDER: CORONOSAURIA SERENO, 1986
PARVORDER: CERATOPOMORPHA Wolfe & Kirkland, 1998 (emend)
Superfamily: CERATOPOIDEA Marsh, 1888 (sensu Hay, 1902)
Family: CERATOPIDAE Marsh, 1888
Subfamily: CERATOPIINAE Marsh, 1888 (sensu Abel, 1919)
Tribe: CHASMOSAURINI Lambe, 1915
*Anchiceratops ornatus
_____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
ON BOTH BANKS OF THE RED DEER RIVER
3-10 MILES BELOW TOLMAN
HORSESHOE CANYON (EDMONTON?) FORMATION
EDMONTON GROUP
EDMONTONIAN AGE
LOWER MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: MARGINOCEPHALIA SERENO, 1986
ORDER: CERATOPIA MARSH, 1890
SUBORDER: NEOCERATOPIA SERENO, 1986
INFRAORDER: CORONOSAURIA SERENO, 1986
PARVORDER: CERATOPOMORPHA Wolfe & Kirkland, 1998 (emend)
Superfamily: CERATOPOIDEA Marsh, 1888 (sensu Hay, 1902)
Family: CERATOPIDAE Marsh, 1888
Incertae sedis
Gen. sp indet.
Subfamily: CENTROSAURINAE Lambe, 1915
Centrosaurus sp
_____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
RED DEER RIVER
16 MILES BELOW TOLMAN FERRY
HORSESHOE CANYON FORMATION
EDMONTON GROUP
EDMONTONIAN AGE
LOWER MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
SUBORDER: ANKYLOPOLLEXIA SERENO, 1986
INFRAORDER: STYRACOSTERNA SERENO, 1986
SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)
Family: HADROSAURIDAE COPE, 1869
Subfamily: LAMBEOSAURINAE Parks, 1923
Tribe: CORYTHOSAURINI Brett-Surman vide Chapman & Brett-Surman, 1990
Hypacrosaurus altispinus
____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
DRUMHELLER
DRUMHELLER MARINE TONGUE LOCALITY
HORSESHOE CANYON FORMATION
EDMONTON GROUP
EDMONTONIAN AGE
LOWER MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?
SUPERORDER: BATRACHIA BRONGNIART, 1800
ORDER: CAUDATA SCOPOLI, 1777
Incertae sedis
Gen. sp indet.
SUPERORDER: SALIENTIA LAURENTI, 1768
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
DRUMHELLER
STRANGER’S RANCH
HORSESHOE CANYON FORMATION
EDMONTON GROUP
LANCIAN AGE
LOWER MAASTRICHTIAN STAGE
UPPERMOST SENONIAN SUBEPOCH
UPPER UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
ORDER: EUSUCHIA emend HUXLEY, 1875
INFRAORDER: CROCODYLIA emend GMELIN, 1877
PARVORDER: BREVIROSTRES
Superfamily: ALLIGATORIDEA Buffetaut, 1985
GLOBIDONTA BROUCHU, 1999
Family: ALLIGATORIDAE Gray, 1844
*Stangerochampsa mccabei
_____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
NEAR DRUMHELLER
HORSESHOE CANYON FORMATION
EDMONTON GROUP
EDMONTONIAN AGE
LOWER MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: EURYAPSIDA COLBERT, 1945
SUPERORDER: SAUROPTERYGIA OWEN 1860
GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994
ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
NEAR DRUMHELLER
RED DEER RIVER
SCOLLARD FORMATION
UPPERMOST EDMONTON GROUP
LANCIAN AGE
UPPERMOST MAASTRICHTIAN STAGE
UPPERMOST SENONIAN SUBEPOCH
UPPER UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
SUBORDER: ANKYLOPOLLEXIA SERENO, 1986
INFRAORDER: STYRACOSTERNA SERENO, 1986
SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)
Family: HADROSAURIDAE COPE, 1869
Subfamily: HADROSAURINAE Cope, 1869 (sensu Lambe, 1918)
Tribe: EDMONTOSAURINI Brett-Surman vide Chapman & Brett-Surman, 1990
Edmontosaurus regalis
____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
DRUMHELLER
SW CORNER OF LEGAL SUBDIVISIION 6
HORSESHOE CANYON FORMATION
EDMONTON GROUP
EDMONTONIAN AGE
LOWER MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?
SUPERORDER: BATRACHIA BRONGNIART, 1800
ORDER: CAUDATA SCOPOLI, 1777
SUBORDER: PORTEOIDEA (NOBEL 1931) EMEND EDWARDS, 1976
Family: BATRACHOSAUROIDIDAE Auffenberg, 1958
Opisthotriton sp
SUBORDER: AMBYSTOMATOIDEA NOBLE, 1931
Family: SCAPHERPETONTIDAE Auffenberg & Goin, 1959
Scapherpeton sp
____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
DRUMHELLER
RED DEER RIVER VALLEY
EDMONTON GROUP
LANCIAN AGE
LOWER MAASTRICHTIAN STAGE
UPPERMOST SENONIAN SUBEPOCH
UPPER UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT CHORISTODERA NOVA
ORDER: CHORISTODERA COPE, 1876
Family: CHAMPSOSAURIDAE Cope, 1884
Champsosaurus albertensis?
____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
DRUMHELLER
HORSESHOE CANYON FORMATION
EDMONTON GROUP
LANCIAN AGE
LOWER MAASTRICHTIAN STAGE
UPPERMOST SENONIAN SUBEPOCH
UPPER UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGANORDER: CRYPTODIRA GAFFNEY & MEYLAN, 1988
SUPERORDER: SELMACRYPTODIRA GAFFNEY, HUTCHISON, JENKINS & MEEKR, 1987
GRANDORDER: DAIOCRYPTODIRA GAFFNEY & MEYLAN, 1988
MICRORDER: CENTROCRYPTODIRA GAFFNEY & MEYLAN, 1988
ORDER: POLYCRPTODIRA GAFFNEY & MEYLAN, 1988
SUBORDER: PROCOELOCRYPTODIRA GAFFNEY & MEYLAN, 1988
INFRAORDER: CHELOMACRYPTODIRA GAFFNEY & MEYLAN, 1988
PARVORDER: TRIONYCHIDA GRAY, 1870
Superfamily: TRIONYCHIDEA Bell, 1828 (sensu Gray, 1870)
Family: NANHSIUNGCHELYIDAE Yeh, 1966
Basilemys sp
Family: TRIONYCHIDAE Bell, 1828
Aspideretes sp
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
ORDER: EUSUCHIA emend HUXLEY, 1875
Family: BOREALOSUCHIDAE emend BROCHU, 1999
Borealosuchus sp
= Leidyosuchus spINFRAORDER: CROCODYLIA emend GMELIN, 1877
PARVORDER: BREVIROSTRES
Superfamily: ALLIGATORIDEA Buffetaut, 1985
GLOBIDONTA BROUCHU, 1999
Family: ALLIGATORIDAE Gray, 1844
Albertochampsa sp
____________________________________________________________________________________
CANADA
SOUTHERN ALBERTA PROVINCE
WHEATLAND COUNTY
DRUMHELLER
KNEEHILLS CREEK
HORSESHOE CANYON FORMAITON
EDMONTON GROUP
EDMONTONIAN AGE
LOWER MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA PAUL, 2002
SUBORDER: COELURIA MARSH, 1881
Superfamily: TYRANNOSAUROIDEA Osborn, 1906 (sensu Bonaparte et. al., 1990)
Family: TYRANNOSAURIDAE Osborn, 1906
Subfamily: ALBERTOSAURINAE Currie, Hurum & Sabath, 2003
*Albertosaurus sarcophagus
_____________________________________________________________________________________
CANADA
SOUTHERN ALBERTA PROVINCE
WHEATLAND COUNTY
DRUMHELLER
EAST BANK OF THE RED DEER RIVER
HORSESHOE CANYON FORMAITON
EDMONTON GROUP
EDMONTONIAN AGE
LOWER MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA PAUL, 2002
SUBORDER: COELURIA MARSH, 1881
Superfamily: TYRANNOSAUROIDEA Osborn, 1906 (sensu Bonaparte et. al., 1990)
Family: TYRANNOSAURIDAE Osborn, 1906
Subfamily: ALBERTOSAURINAE Currie, Hurum & Sabath, 2003
Albertosaurus sarcophagus
_____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
NEAR DRUMHELLER
OPPOSITE THE MOUTH OF THREE HILLS CREEK
SCOLLARD FORMATION
UPPERMOST EDMONTON GROUP
LANCIAN AGE
UPPERMOST MAASTRICHTIAN STAGE
UPPERMOST SENONIAN SUBEPOCH
UPPER UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
SUBORDER: ANKYLOPOLLEXIA SERENO, 1986
INFRAORDER: STYRACOSTERNA SERENO, 1986
SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)
Family: HADROSAURIDAE COPE, 1869
Subfamily: HADROSAURINAE Cope, 1869 (sensu Lambe, 1918)
Tribe: EDMONTOSAURINI Brett-Surman vide Chapman & Brett-Surman, 1990
*Edmontosaurus regalis
____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
DRUMHELLER
LESS THAN 1.6 KM (I M) FROM THE TYRRELL MUSEUM
HORSESHOE CANYON FORMATION
EDMONTON GROUP
EDMONTONIAN AGE
LOWER MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA PAUL, 2002
SUBORDER: COELURIA MARSH, 1881
Superfamily: TYRANNOSAUROIDEA Osborn, 1906 (sensu Bonaparte et. al., 1990)
Family: TYRANNOSAURIDAE Osborn, 1906
Subfamily: ALBERTOSAURINAE Currie, Hurum & Sabath, 2003
Albertosaurus sarcophagus
____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
ABOUT 30 KM N OF DRUMHELLER
HALFWAY UP A STEEP CLIFF
HORSESHOE CANYON FORMATION
EDMONTON GROUP
EDMONTONIAN AGE
LOWER MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA PAUL, 2002
SUBORDER: COELURIA MARSH, 1881
Superfamily: TYRANNOSAUROIDEA Osborn, 1906 (sensu Bonaparte et. al., 1990)
Family: TYRANNOSAURIDAE Osborn, 1906
Subfamily: ALBERTOSAURINAE Currie, Hurum & Sabath, 2003
Albertosaurus sarcophagus
= Albertosaurus arctunguis
= Deinodon arctunguis
____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
DRUMHELLER
1 M FROM THE TYRRELL MUSEUM
HORSESHOE CANYON FORMATION
EDMONTON GROUP
EDMONTONIAN AGE
LOWER MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA PAUL, 2002
SUBORDER: COELURIA MARSH, 1881
Superfamily: TYRANNOSAUROIDEA Osborn, 1906 (sensu Bonaparte et. al., 1990)
Family: TYRANNOSAURIDAE Osborn, 1906
Subfamily: TYRANNOSAURINAE Osborn, 1906
Genus nova
____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
DRUMHELLER
SW CORNER OF LEGAL SUBDIVISION 6
HORSESHOE CANYON FORMATION
EDMONTON GROUP
EDMONTONIAN AGE
LOWER MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA PAUL, 2002
SUBORDER: COELURIA MARSH, 1881
Superfamily: TYRANNOSAUROIDEA Osborn, 1906 (sensu Bonaparte et. al., 1990)
Family: TYRANNOSAURIDAE Osborn, 1906
Subfamily: ALBERTOSAURINAE Currie, Hurum & Sabath, 2003
Albertosaurus sp
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: MARGINOCEPHALIA SERENO, 1986
ORDER: CERATOPIA MARSH, 1890
SUBORDER: NEOCERATOPIA SERENO, 1986
INFRAORDER: CORONOSAURIA SERENO, 1986
PARVORDER: CERATOPOMORPHA Wolfe & Kirkland, 1998 (emend)
Superfamily: CERATOPOIDEA Marsh, 1888 (sensu Hay, 1902)
Family: CERATOPIDAE Marsh, 1888
Subfamily: PACHYRHINOSAURINAE [nomen novum ex PACHYRHINOSAURIDAE C. M. Sternberg, 1950]
Pachyrhinosaurus canadensis
_____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
5 M FROM DRUMHELLER
MICHICHI CREEK
SCOLLARD FORMATION
UPPERMOST EDMONTON GROUP
LANCIAN AGE
UPPERMOST MAASTRICHTIAN STAGE
UPPERMOST SENONIAN SUBEPOCH
UPPER UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
SUBORDER: ANKYLOPOLLEXIA SERENO, 1986
INFRAORDER: STYRACOSTERNA SERENO, 1986
SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)
Family: HADROSAURIDAE COPE, 1869
Subfamily: HADROSAURINAE Cope, 1869 (sensu Lambe, 1918)
Tribe: EDMONTOSAURINI Brett-Surman vide Chapman & Brett-Surman, 1990
Anatosaurus annectens
= *Thespesius edmontonensis
= Thespesius edmontoni
= Anatosaurus edmontonensis
= Anatosaurus edmontoni
= Edmontosaurus edmontonensis
= Edmontosaurus edmontoni
= Trachodon edmontonensis
= Trachodon edmontoni
____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
MICHICHI CREEK
HORSESHOE CANYON FORMATION
EDMONTON GROUP
EDMONTONIAN AGE
LOWER MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: MAMMALIA LINNAEUS, 1758
SUBCLASS: PROTOTHERIA GILL, 1872
INFRACLASS: TRECHNOTHERIA McKENNA, 1975
LEGION: CLADOTHERIA McKENNA, 1975
SUBLEGION: ZATHERIA MCKENNA, 1975
INFRALEGION: TRIBOSPHENIDA (McKENNA, 1975) McKENNA & BELL, 1997
SUPERCOHORT: THERIA (PARKER & HASWELL, 1987) McKENNA & BELL, 1997
COHORT: MARSUPIALIA (ILLIGER, 1811) McKENNA & BELL, 1997
ORDER: STAGODONTIA ARCHER, 1984
Family: STAGODONTIDAE Marsh, 1889
*Didelphodon coyi
____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
RTMP LOCALITY HC-4
HORSESHOE CANYON FORMATION
EDMONTON GROUP
EDMONTONIAN AGE
LOWER MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: MAMMALIA LINNAEUS, 1758
SUBCLASS: PROTOTHERIA GILL, 1872
INFRACLASS: TRECHNOTHERIA McKENNA, 1975
LEGION: CLADOTHERIA McKENNA, 1975
SUBLEGION: ZATHERIA MCKENNA, 1975
INFRALEGION: TRIBOSPHENIDA (McKENNA, 1975) McKENNA & BELL, 1997
SUPERCOHORT: THERIA (PARKER & HASWELL, 1987) McKENNA & BELL, 1997
COHORT: MARSUPIALIA (ILLIGER, 1811) McKENNA & BELL, 1997
ORDER: STAGODONTIA ARCHER, 1984
Family: STAGODONTIDAE Marsh, 1889
Didelphodon coyi
____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
NEAR DRUMHELLER
3 M ABOVE MUNSON FERRY
SCOLLARD FORMATION
UPPERMOST EDMONTON GROUP
LANCIAN AGE
UPPERMOST MAASTRICHTIAN STAGE
UPPERMOST SENONIAN SUBEPOCH
UPPER UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
SUBORDER: ANKYLOPOLLEXIA SERENO, 1986
INFRAORDER: STYRACOSTERNA SERENO, 1986
SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)
Family: HADROSAURIDAE COPE, 1869
Subfamily: HADROSAURINAE Cope, 1869 (sensu Lambe, 1918)
Tribe: EDMONTOSAURINI Brett-Surman vide Chapman & Brett-Surman, 1990
Anatosaurus annectens
= Thespesius edmontonensis
= Thespesius edmontoni
= Anatosaurus edmontonensis
= Anatosaurus edmontoni
= Edmontosaurus edmontonensis
= Edmontosaurus edmontoni
= Trachodon edmontonensis
= Trachodon edmontoni
____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
DRUMHELLER
1.5 M S OF MUNSON FERRY
SW CORNER OF LEGAL SUBDIVISION 6
HORSESHOE CANYON FORMATION
EDMONTON GROUP
EDMONTONIAN AGE
LOWER MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
SUBORDER: ANKYLOPOLLEXIA SERENO, 1986
INFRAORDER: STYRACOSTERNA SERENO, 1986
SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)
Family: HADROSAURIDAE COPE, 1869
Subfamily: HADROSAURINAE Cope, 1869 (sensu Lambe, 1918)
Tribe: EDMONTOSAURINI Brett-Surman vide Chapman & Brett-Surman, 1990
Edmontosaurus regalis
____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
DRUMHELLER VALLEY BADLANDS
RTMP LOCALITY 2000 (L2000)
HORSESHOE CANYON FORMATION
EDMONTON GROUP
LANCIAN AGE
LOWER MAASTRICHTIAN STAGE
UPPERMOST SENONIAN SUBEPOCH
UPPER UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
Incertae sedis
Gen. sp indet.
MAGANORDER: CRYPTODIRA GAFFNEY & MEYLAN, 1988
SUPERORDER: SELMACRYPTODIRA GAFFNEY, HUTCHISON, JENKINS & MEEKR, 1987
GRANDORDER: DAIOCRYPTODIRA GAFFNEY & MEYLAN, 1988
MICRORDER: BAENOIDEA WILLIAMS, 1960
Family: BAENIDAE Cope, 1882
Subfamily: BAENINAE Cope, 1882 (sensu Williams, 1950)
Gen. sp indet.
MICRORDER: CENTROCRYPTODIRA GAFFNEY & MEYLAN, 1988
ORDER: POLYCRPTODIRA GAFFNEY & MEYLAN, 1988
SUBORDER: PROCOELOCRYPTODIRA GAFFNEY & MEYLAN, 1988
INFRAORDER: CHELOMACRYPTODIRA GAFFNEY & MEYLAN, 1988
PARVORDER: TRIONYCHIDA GRAY, 1870
Superfamily: TRIONYCHIDEA Bell, 1828 (sensu Gray, 1870)
Family: TRIONYCHIDAE Bell, 1828
Gen. sp indet.
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT CHORISTODERA NOVA
ORDER: CHORISTODERA COPE, 1876
Family: CHAMPSOSAURIDAE Cope, 1884
Gen sp indet.
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
SUPERORDER: TAPINOPLATIA PETERS, 2000
GRANDORDER: CHARACIOPODA PETERS, 2000
MICRORDER: FENESTRASAURIA PETERS, 2000
ORDER: PTEROSAURIA KAUP, 1834
Incertae sedis
Gen. sp indet.
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
ORDER: Incertae sedis
Ricardoestesia sp
Gen. sp indet.
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA PAUL, 2002
SUBORDER: COELURIA MARSH, 1881
Superfamily: TYRANNOSAUROIDEA Osborn, 1906 (sensu Bonaparte et. al., 1990)
Family: TYRANNOSAURIDAE Osborn, 1906
Subfamily: ALBERTOSAURINAE Currie, Hurum & Sabath, 2003
Albertosaurus sp
AVEPECTORA PAUL, 2002
SUBORDER: MANIRAPTORA GAUTHIER, 1986
INFRAORDER: PARVES SERENO, 1997
PARVORDER: DEINONYCHOSAURIA COLBERT & D. A. RUSSELL, 1969
Incerate sedis
Paronychodon sp
Family: DROMAEOSAURIDAE Russell, 1969
Incertae sedis
Gen. sp indet.
Subfamily: VELOCIRAPTORINAE Barsbold, 1978
cf. Saurornitholestes
Family: TROODONTIDAE Gilmore, 1924
Troodon sp
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
SUBORDER: ANKYLOPOLLEXIA SERENO, 1986
INFRAORDER: STYRACOSTERNA SERENO, 1986
SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)
Family: HADROSAURIDAE COPE, 1869
Incertae sedis
Gen. sp indet.
MICRORDER: MARGINOCEPHALIA SERENO, 1986
ORDER: CERATOPIA MARSH, 1890
SUBORDER: NEOCERATOPIA SERENO, 1986
INFRAORDER: CORONOSAURIA SERENO, 1986
PARVORDER: CERATOPOMORPHA Wolfe & Kirkland, 1998 (emend)
Superfamily: CERATOPOIDEA Marsh, 1888 (sensu Hay, 1902)
Family: CERATOPIDAE Marsh, 1888
Incertae sedis
Gen. sp indet.
SUPERORDER: EURYOPDA SERENO, 1986
GRANDORDER: ARMATOSAURIA ZHAO, 1983 (emend)
ORDER: ANKYLOSAURIA OSBORN, 1923
INFRAORDER: ANKYLOSAURA OSBORN, 1923
Family: ANKYLOSAURIDAE Brown, 1908
Incertae sedis
Gen. sp indet.
CLASS: AVES LINNAEUS, 1758
Incertae sedis
Gen. sp indet.
CLASS: MAMMALIA LINNAEUS, 1758
SUBCLASS: PROTOTHERIA GILL, 1872
INFRACLASS: TRECHNOTHERIA McKENNA, 1975
LEGION: CLADOTHERIA McKENNA, 1975
SUBLEGION: ZATHERIA MCKENNA, 1975
INFRALEGION: TRIBOSPHENIDA (McKENNA, 1975) McKENNA & BELL, 1997
SUPERCOHORT: THERIA (PARKER & HASWELL, 1987) McKENNA & BELL, 1997
COHORT: MARSUPIALIA (ILLIGER, 1811) McKENNA & BELL, 1997
ORDER: STAGODONTIA ARCHER, 1984
Family: PEDIOMYIDAE Simpson, 1927
Pediomys sp
____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
DRUMHELLER
SCOLLARD FORMATION
UPPERMOST EDMONTON GROUP
LANCIAN AGE
UPPERMOST MAASTRICHTIAN STAGE
UPPERMOST SENONIAN SUBEPOCH
UPPER UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
ORDER: EUSUCHIA emend HUXLEY, 1875
INFRAORDER: CROCODYLIA emend GMELIN, 1877
Incerate sedis
Gen. sp. indet.
____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
APPROXIMATLEY .6 M NE OF “BIGGEST LITTLE CHURCH”
DRUMHELLER
NORTH SIDE OF RED DEER RIVER
EDMONTON FORMATION?
EDMONTONIAN AGE
MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
6 M ABOVE EDMONTON
NORTH SASKATCHEWAN RIVER
HORSESHOE CANYON FORMATION
EDMONTON GROUP
EDMONTONIAN AGE
LOWER MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: EURYAPSIDA COLBERT, 1945
SUPERORDER: SAUROPTERYGIA OWEN 1860
GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994
ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
INFRAORDER: PLESIOSAUROMORPHA NOVA
EUPLESIOSAURIA O’KEEFE, 2001
Superfamily: ELASMOSAUROIDEA nova
Family: ELASMOSAURIDAE Cope, 1869
Incertae sedis
Leurospondylus ultimus
_____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
NEAR TOLMAN CROSSING
HORSESHOE CANYON FORMATION
LOWER EDMONTON GROUP
LANCIAN AGE
LOWER MAASTRICHTIAN STAGE
UPPERMOST SENONIAN SUBEPOCH
UPPER UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT CHORISTODERA NOVA
ORDER: CHORISTODERA COPE, 1876
Family: CHAMPSOSAURIDAE Cope, 1884
Incertae sedis
Champsosaurus sp
_____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
WEST SIDE OF THE RED DEER RIVER NEAR MORRIN BRIDGE
EDMONTON FORMATION
EDMONTONIAN AGE
UPPER CAMPANIAN STAGE- LOWER MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
ORDER: EUSUCHIA emend HUXLEY, 1875
Family: BOREALOSUCHIDAE emend BROCHU, 1999
Borealosuchus sternbergii
= Leidyosuchus sternbergii
_____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
NEAR DRUMHELLER
7 M NW OF MORRIN
SCOLLARD FORMATION
UPPERMOST EDMONTON GROUP
LANCIAN AGE
UPPERMOST MAASTRICHTIAN STAGE
UPPERMOST SENONIAN SUBEPOCH
UPPER UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
SUBORDER: ANKYLOPOLLEXIA SERENO, 1986
INFRAORDER: STYRACOSTERNA SERENO, 1986
SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)
Family: HADROSAURIDAE COPE, 1869
Subfamily: HADROSAURINAE Cope, 1869 (sensu Lambe, 1918)
Tribe: EDMONTOSAURINI Brett-Surman vide Chapman & Brett-Surman, 1990
Edmontosaurus regalis
____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
NW OF MORRIN
20 FT ABOVE THE RED DEER RIVER
HORSESHOE CANYON FORMATION
EDMONTON GROUP
EDMONTONIAN AGE
LOWER MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: EURYOPDA SERENO, 1986
GRANDORDER: ARMATOSAURIA ZHAO, 1983 (emend)
ORDER: ANKYLOSAURIA OSBORN, 1923
INFRAORDER: ANKYLOSAURA OSBORN, 1923
Family: NODOSAURIDAE Marsh, 1890
Subfamily: EDMONTONIINAE Russell, 1940
*Edmontonia longiceps
= Panoplosaurus longiceps
_____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
RED DEER RIVER
NEAR DRUMHELLER
SW MORRIN
HORSESHOE CANYON FORMATION
EDMONTON GROUP
EDMONTONIAN AGE
LOWER MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: MARGINOCEPHALIA SERENO, 1986
ORDER: CERATOPIA MARSH, 1890
SUBORDER: NEOCERATOPIA SERENO, 1986
INFRAORDER: CORONOSAURIA SERENO, 1986
PARVORDER: CERATOPOMORPHA Wolfe & Kirkland, 1998 (emend)
Superfamily: CERATOPOIDEA Marsh, 1888 (sensu Hay, 1902)
Family: CERATOPIDAE Marsh, 1888
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
RED DEER RIVER
NEAR DRUMHELLER
ON A LEDGE 1/4 M W OF MORRIN BRIDGE
HORSESHOE CANYON FORMATION
EDMONTON GROUP
EDMONTONIAN AGE
LOWER MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
Incertae sedis
Gen. sp indet.
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
SUBORDER: ANKYLOPOLLEXIA SERENO, 1986
INFRAORDER: STYRACOSTERNA SERENO, 1986
SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)
Family: HADROSAURIDAE COPE, 1869
Incertae sedis
Gen. sp indet.
____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
NEAR DRUMHELLER
APPROXIMATLEY 1 M NORTH OF MORRIN BRIDGE
EAST SIDE OF RED DEER RIVER
HORSESHOE CANYON FORMATION
EDMONTON GROUP
EDMONTONIAN AGE
LOWER MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
NEAR DRUMHELLER
NORTH WALL OF SECOND COULEE NORTH OF MORRIN BRIDGE
HORSESHOE CANYON FORMATION
EDMONTON GROUP
EDMONTONIAN AGE
LOWER MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
NEAR DRUMHELLER
APPROXIMATELY 2 M NE OF MORRIN BRIDGE
EAST SIDE OF RED DEER RIVER
HORSESHOE CANYON FORMATION
EDMONTON GROUP
EDMONTONIAN AGE
LOWER MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
NEAR DRUMHELLER
IN A COULEE APPROXIMATELY 1.75 M NNW OF MORRIN BRIDGE
HORSESHOE CANYON FORMATION
EDMONTON GROUP
EDMONTONIAN AGE
LOWER MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
NEAR DRUMHELLER
7 M W AND 2 MN OF MORRIN
HORSESHOE CANYON FORMATION
EDMONTON GROUP
EDMONTONIAN AGE
LOWER MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: EURYOPDA SERENO, 1986
GRANDORDER: ARMATOSAURIA ZHAO, 1983 (emend)
ORDER: ANKYLOSAURIA OSBORN, 1923
INFRAORDER: ANKYLOSAURA OSBORN, 1923
Family: ANKYLOSAURIDAE Brown, 1908
Subfamily: ANKYLOSAURINAE Brown, 1908
Euoplocephalus tutus
= Stereocephalus tutus
= Palaeoscincus tutus
_____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
NEAR DRUMHELLER
1.3 M N OF MORRIN BRIDGE
HORSESHOE CANYON FORMATION
EDMONTON GROUP
EDMONTONIAN AGE
LOWER MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
NEAR DRUMHELLER
1.3 M N OF MORRIN BRIDGE
HORSESHOE CANYON FORMATION
EDMONTON GROUP
EDMONTONIAN AGE
LOWER MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
RED DEER RIVER
NEAR DRUMHELLER
6 M W OF MORRIN
HORSESHOE CANYON FORMATION
EDMONTON GROUP
EDMONTONIAN AGE
LOWER MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: MARGINOCEPHALIA SERENO, 1986
ORDER: CERATOPIA MARSH, 1890
SUBORDER: NEOCERATOPIA SERENO, 1986
INFRAORDER: CORONOSAURIA SERENO, 1986
PARVORDER: CERATOPOMORPHA Wolfe & Kirkland, 1998 (emend)
Superfamily: CERATOPOIDEA Marsh, 1888 (sensu Hay, 1902)
Family: CERATOPIDAE Marsh, 1888
Subfamily: CERATOPIINAE Marsh, 1888 (sensu Abel, 1919)
Tribe: CHASMOSAURINI Lambe, 1915
Anchiceratops ornatus
_____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
RED DEER RIVER
NEAR DRUMHELLER
7 1/2 M SW OF MORRIN
HORSESHOE CANYON FORMATION
EDMONTON GROUP
EDMONTONIAN AGE
LOWER MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: MARGINOCEPHALIA SERENO, 1986
ORDER: CERATOPIA MARSH, 1890
SUBORDER: NEOCERATOPIA SERENO, 1986
INFRAORDER: CORONOSAURIA SERENO, 1986
PARVORDER: CERATOPOMORPHA Wolfe & Kirkland, 1998 (emend)
Superfamily: CERATOPOIDEA Marsh, 1888 (sensu Hay, 1902)
Family: CERATOPIDAE Marsh, 1888
Subfamily: CERATOPIINAE Marsh, 1888 (sensu Abel, 1919)
Tribe: CHASMOSAURINI Lambe, 1915
Anchiceratops ornatus
_____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
RED DEER RIVER
NEAR DRUMHELLER
12 NW OF MORRIN
HORSESHOE CANYON FORMATION
EDMONTON GROUP
EDMONTONIAN AGE
LOWER MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: MARGINOCEPHALIA SERENO, 1986
ORDER: CERATOPIA MARSH, 1890
SUBORDER: NEOCERATOPIA SERENO, 1986
INFRAORDER: CORONOSAURIA SERENO, 1986
PARVORDER: CERATOPOMORPHA Wolfe & Kirkland, 1998 (emend)
Superfamily: CERATOPOIDEA Marsh, 1888 (sensu Hay, 1902)
Family: CERATOPIDAE Marsh, 1888
Subfamily: CERATOPIINAE Marsh, 1888 (sensu Abel, 1919)
Tribe: CHASMOSAURINI Lambe, 1915
*Anchiceratops longirostris
_____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
WEST SIDE OF RED DEER RIVER
NEAR DRUMHELLER
6 M W OF MUNSON
SCOLLARD FORMATION
UPPERMOST EDMONTON GROUP
LANCIAN AGE
UPPERMOST MAASTRICHTIAN STAGE
UPPERMOST SENONIAN SUBEPOCH
UPPER UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: MARGINOCEPHALIA SERENO, 1986
ORDER: CERATOPIA MARSH, 1890
SUBORDER: NEOCERATOPIA SERENO, 1986
INFRAORDER: CORONOSAURIA SERENO, 1986
PARVORDER: CERATOPOMORPHA Wolfe & Kirkland, 1998 (emend)
Superfamily: CERATOPOIDEA Marsh, 1888 (sensu Hay, 1902)
Family: CERATOPIDAE Marsh, 1888
Subfamily: CERATOPIINAE Marsh, 1888 (sensu Abel, 1919)
Tribe: CHASMOSAURINI Lambe, 1915
Arrhinoceratops brachyops
_____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
NEAR DRUMHELLER
2 M ABOVE MUNSON FERRY
SCOLLARD FORMATION
UPPERMOST EDMONTON GROUP
LANCIAN AGE
UPPERMOST MAASTRICHTIAN STAGE
UPPERMOST SENONIAN SUBEPOCH
UPPER UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
SUBORDER: ANKYLOPOLLEXIA SERENO, 1986
INFRAORDER: STYRACOSTERNA SERENO, 1986
SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)
Family: HADROSAURIDAE COPE, 1869
Subfamily: HADROSAURINAE Cope, 1869 (sensu Lambe, 1918)
Tribe: EDMONTOSAURINI Brett-Surman vide Chapman & Brett-Surman, 1990
Edmontosaurus regalis
_____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
WEST SIDE OF THE RED DEER RIVER
NEAR DRUMHELLER
6 M W AND 3 M N OF MUNSON
SCOLLARD FORMATION
UPPERMOST EDMONTON GROUP
LANCIAN AGE
UPPERMOST MAASTRICHTIAN STAGE
UPPERMOST SENONIAN SUBEPOCH
UPPER UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: MARGINOCEPHALIA SERENO, 1986
ORDER: CERATOPIA MARSH, 1890
SUBORDER: NEOCERATOPIA SERENO, 1986
INFRAORDER: CORONOSAURIA SERENO, 1986
PARVORDER: CERATOPOMORPHA Wolfe & Kirkland, 1998 (emend)
Superfamily: CERATOPOIDEA Marsh, 1888 (sensu Hay, 1902)
Family: CERATOPIDAE Marsh, 1888
Subfamily: CERATOPIINAE Marsh, 1888 (sensu Abel, 1919)
Tribe: CHASMOSAURINI Lambe, 1915
Anchiceratops ornatus
Arrhinoceratops brachyops
_____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
STEPHEN’S SITE
HORSESHOE CANYON FORMATION
EDMONTON GROUP
EDMONTONIAN AGE
LOWER MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
ORDER: Incertae sedis
Ricardoestesia gilmorei
Ricardoestesia spAVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA PAUL, 2002
SUBORDER: COELURIA MARSH, 1881
Superfamily: TYRANNOSAUROIDEA Osborn, 1906 (sensu Bonaparte et. al., 1990)
Family: TYRANNOSAURIDAE Osborn, 1906
Gen. sp indet.
AVEPECTORA PAUL, 2002
SUBORDER: MANIRAPTORA GAUTHIER, 1986
INFRAORDER: PARVES SERENO, 1997
PARVORDER: DEINONYCHOSAURIA COLBERT & D. A. RUSSELL, 1969
Family: DROMAEOSAURIDAE Russell, 1969
Subfamily: DROMAEOSAURINAE Matthew & Brown, 1922
Dromaeosaurus albertensis
Subfamily: VELOCIRAPTORINAE Barsbold, 1978
cf. Saurornitholestes longstoni
Family: TROODONTIDAE Gilmore, 1924
cf. Troodon formosus
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
SUBORDER: ANKYLOPOLLEXIA SERENO, 1986
INFRAORDER: STYRACOSTERNA SERENO, 1986
SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)
Family: HADROSAURIDAE COPE, 1869
Subfamily: HADROSAURINAE Cope, 1869 (sensu Lambe, 1918)
Tribe: EDMONTOSAURINI Brett-Surman vide Chapman & Brett-Surman, 1990
cf. Edmontosaurus
MICRORDER: MARGINOCEPHALIA SERENO, 1986
ORDER: PACHYCEPHALOSAURIA MARYANSKA & OSMOLSKA, 1974
SUBORDER: DOMOCEPHALINIA SERENO, 1986
INFRAORDER: PACHYCEPHALOSAURIA (MARYANSKA & OSMOLSKA, 1974) COOPER, 1985
Family: PACHYCEPHALOSAURIDAE Sternberg, 1945
Gen. sp indet.
ORDER: CERATOPIA MARSH, 1890
SUBORDER: NEOCERATOPIA SERENO, 1986
INFRAORDER: CORONOSAURIA SERENO, 1986
PARVORDER: CERATOPOMORPHA Wolfe & Kirkland, 1998 (emend)
Superfamily: CERATOPOIDEA Marsh, 1888 (sensu Hay, 1902)
Family: CERATOPIDAE Marsh, 1888
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
JANET’S SITE
HORSESHOE CANYON FORMATION
EDMONTON GROUP
EDMONTONIAN AGE
LOWER MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
ORDER: Incertae sedis
Ricardoestesia gilmorei
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA PAUL, 2002
SUBORDER: COELURIA MARSH, 1881
Superfamily: TYRANNOSAUROIDEA Osborn, 1906 (sensu Bonaparte et. al., 1990)
Family: TYRANNOSAURIDAE Osborn, 1906
Gen. sp indet.
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
SUBORDER: ANKYLOPOLLEXIA SERENO, 1986
INFRAORDER: STYRACOSTERNA SERENO, 1986
SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)
Family: HADROSAURIDAE COPE, 1869
Gen. sp indet.
_____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
PREHISTORIC PARK QUARRY
HORSESHOE CANYON FORMATION
EDMONTON GROUP
EDMONTONIAN AGE
LOWER MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGANORDER: CRYPTODIRA GAFFNEY & MEYLAN, 1988
SUPERORDER: SELMACRYPTODIRA GAFFNEY, HUTCHISON, JENKINS & MEEKR, 1987
GRANDORDER: DAIOCRYPTODIRA GAFFNEY & MEYLAN, 1988
MICRORDER: CENTROCRYPTODIRA GAFFNEY & MEYLAN, 1988
ORDER: POLYCRPTODIRA GAFFNEY & MEYLAN, 1988
SUBORDER: PROCOELOCRYPTODIRA GAFFNEY & MEYLAN, 1988
INFRAORDER: CHELOMACRYPTODIRA GAFFNEY & MEYLAN, 1988
PARVORDER: TRIONYCHIDA GRAY, 1870
Superfamily: TRIONYCHIDEA Bell, 1828 (sensu Gray, 1870)
Family: TRIONYCHIDAE Bell, 1828
Aspideretoides sp
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT CHORISTODERA NOVA
ORDER: CHORISTODERA COPE, 1876
Family: CHAMPSOSAURIDAE Cope, 1884
Incertae sedis
Cf. Champsosaurus sp
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
Incertae sedis
Gen. sp indet.
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA PAUL, 2002
SUBORDER: COELURIA MARSH, 1881
Superfamily: TYRANNOSAUROIDEA Osborn, 1906 (sensu Bonaparte et. al., 1990)
Family: TYRANNOSAURIDAE Osborn, 1906
Subfamily: ALBERTOSAURINAE Currie, Hurum & Sabath, 2003
Cf. Albertosaurus sp
AVEPECTORA PAUL, 2002
SUBORDER: MANIRAPTORA GAUTHIER, 1986
INFRAORDER: PARVES SERENO, 1997
PARVORDER: DEINONYCHOSAURIA COLBERT & D. A. RUSSELL, 1969
Family: DROMAEOSAURIDAE Russell, 1969
Subfamily: VELOCIRAPTORINAE Barsbold, 1978
cf. Saurornitholestes
Family: TROODONTIDAE Gilmore, 1924
Troodon sp
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
SUBORDER: ANKYLOPOLLEXIA SERENO, 1986
INFRAORDER: STYRACOSTERNA SERENO, 1986
SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)
Family: HADROSAURIDAE COPE, 1869
Subfamily: HADROSAURINAE Cope, 1869 (sensu Lambe, 1918)
Tribe: EDMONTOSAURINI Brett-Surman vide Chapman & Brett-Surman, 1990
Cf. Edmontosaurus
MICRORDER: MARGINOCEPHALIA SERENO, 1986
ORDER: CERATOPIA MARSH, 1890
SUBORDER: NEOCERATOPIA SERENO, 1986
INFRAORDER: CORONOSAURIA SERENO, 1986
PARVORDER: CERATOPOMORPHA Wolfe & Kirkland, 1998 (emend)
Superfamily: CERATOPOIDEA Marsh, 1888 (sensu Hay, 1902)
Family: CERATOPIDAE Marsh, 1888
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
IN A COULEE TRIBUTARY IN THE RED DEER RIVER VALLEY
SCOLLARD FORMATION
UPPERMOST EDMONTON GROUP
LANCIAN AGE
UPPERMOST MAASTRICHTIAN STAGE
UPPERMOST SENONIAN SUBEPOCH
UPPER UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: MARGINOCEPHALIA SERENO, 1986
ORDER: CERATOPIA MARSH, 1890
SUBORDER: NEOCERATOPIA SERENO, 1986
INFRAORDER: CORONOSAURIA SERENO, 1986
PARVORDER: CERATOPOMORPHA Wolfe & Kirkland, 1998 (emend)
Superfamily: CERATOPOIDEA Marsh, 1888 (sensu Hay, 1902)
Family: CERATOPIDAE Marsh, 1888
Subfamily: CERATOPIINAE Marsh, 1888 (sensu Abel, 1919)
Tribe: TOROSAURINI Nopcsa, 1915
Torosaurus sp
_____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
RED DEER RIVER
ABOUT 1 M NW OF THE MOUTH OF BIG VALLEY CREEK
HORSESHOE CANYON FORMAITON
EDMONTON GROUP
EDMONTONIAN AGE
LOWER MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
SUBORDER: ANKYLOPOLLEXIA SERENO, 1986
INFRAORDER: STYRACOSTERNA SERENO, 1986
SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)
Family: HADROSAURIDAE COPE, 1869
Subfamily: LAMBEOSAURINAE Parks, 1923
Tribe: CORYTHOSAURINI Brett-Surman vide Chapman & Brett-Surman, 1990
Hypacrosaurus altispinus
= Cheneosaurus tolmanensis (Juvenile)
____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
RED DEER RIVER
AT A POINT ON THE EAST SIDE
HORSESHOE CANYON FORMAITON
EDMONTON GROUP
EDMONTONIAN AGE
LOWER MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
SUBORDER: ORNITHOMIMOSAURIA BARSBOLD, 1976
Family: ORNITHOMIMIDAE Marsh, 1890
Ornithomimus antiquus
= Dromiceiomimus brevitertius
= *Struthiomimus ingens
= Dromiceiomimus ingens
= Ornithomimus ingens
_____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
75 FT ABOVE W BANK OF RED DEER RIVER
HORSESHOE CANYON FORMATION
EDMONTON GROUP
EDMONTONIAN AGE
LOWER MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: EURYOPDA SERENO, 1986
GRANDORDER: ARMATOSAURIA ZHAO, 1983 (emend)
ORDER: ANKYLOSAURIA OSBORN, 1923
INFRAORDER: ANKYLOSAURA OSBORN, 1923
Family: ANKYLOSAURIDAE Brown, 1908
Subfamily: ANKYLOSAURINAE Brown, 1908
Euoplocephalus tutus
= Stereocephalus tutus
= Palaeoscincus tutus
_____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND PROVINCE
APPROXIMATELY 9 M ABOVE WEST BANK OF THE RED DEER RIVER
?SECTION 19
HORSESHOE CANYON FORMATION
EDMONTON GROUP
EDMONTONIAN AGE
LOWER MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
SUBORDER: ORNITHOMIMOSAURIA BARSBOLD, 1976
Family: ORNITHOMIMIDAE Marsh, 1890
Ornithomimus antiquus
= Dromiceiomimus brevitertius
= Struthiomimus brevitertius
= Ornithomimus brevitertius
_____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
55 M ABOVE EAST BANK OF RED DEER RIVER
NORTHEAST QUARTER OF SEC. 28, TP 34, R21, WEST OF THE 4TH M
HORSESHOE CANYON FORMATION
EDMONTON GROUP
EDMONTONIAN AGE
LOWER MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
SUBORDER: ORNITHOMIMOSAURIA BARSBOLD, 1976
Family: ORNITHOMIMIDAE Marsh, 1890
Ornithomimus antiquus
= Dromiceiomimus brevitertius
= Struthiomimus brevitertius
= Ornithomimus brevitertius
_____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
APPROXIMATELY 44 M ABOVE WEST BANK OF THE RED DEER RIVER
SEC. 28, TP32, R21, WEST OF 4TH M
HORSESHOE CANYON FORMATION
EDMONTON GROUP
EDMONTONIAN AGE
LOWER MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
SUBORDER: ORNITHOMIMOSAURIA BARSBOLD, 1976
Family: ORNITHOMIMIDAE Marsh, 1890
Ornithomimus antiquus
= Dromiceiomimus brevitertius
= Struthiomimus brevitertius
= Ornithomimus brevitertius
_____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
150 FT ABOVE THE W BANK OF RED DEER RIVER
SECTION 32 (?)
HORSESHOE CANYON FORMATION
EDMONTON GROUP
EDMONTONIAN AGE
LOWER MAASTRICHTIAN STAGE
UPPER SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: EURYOPDA SERENO, 1986
GRANDORDER: ARMATOSAURIA ZHAO, 1983 (emend)
ORDER: ANKYLOSAURIA OSBORN, 1923
INFRAORDER: ANKYLOSAURA OSBORN, 1923
Family: ANKYLOSAURIDAE Brown, 1908
Subfamily: ANKYLOSAURINAE Brown, 1908
Euoplocephalus sp
_____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
RED DEER RIVER VALLEY
IN A COULEE TRIBUTARY
SCOLLARD FORMATION
UPPERMOST EDMONTON GROUP
LANCIAN AGE
UPPERMOST MAASTRICHTIAN STAGE
UPPERMOST SENONIAN SUBEPOCH
UPPER UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA PAUL, 2002
SUBORDER: COELURIA MARSH, 1881
Superfamily: TYRANNOSAUROIDEA Osborn, 1906 (sensu Bonaparte et. al., 1990)
Family: TYRANNOSAURIDAE Osborn, 1906
Subfamily: TYRANNOSAURINAE Osborn, 1906
Tribe: TYRANNOSAURINI Osborn, 1906 (sensu Olshevsky, Ford & Yamamoto, 1995)
Tyrannosaurus rex
_____________________________________________________________________________________
CANADA
ALBERTA PROVINCE
WHEATLAND COUNTY
RED DEER RIVER
NW 1/4, SEC 2, TP34, R22, W 4TH MER
SCOLLARD FORMATION
UPPERMOST EDMONTON GROUP
LANCIAN AGE
UPPERMOST MAASTRICHTIAN STAGE
UPPERMOST SENONIAN SUBEPOCH
UPPER UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: MARGINOCEPHALIA SERENO, 1986
ORDER: CERATOPIA MARSH, 1890
SUBORDER: NEOCERATOPIA SERENO, 1986
INFRAORDER: CORONOSAURIA SERENO, 1986
PARVORDER: CERATOPOMORPHA Wolfe & Kirkland, 1998 (emend)
Superfamily: CERATOPOIDEA Marsh, 1888 (sensu Hay, 1902)
Family: CERATOPIDAE Marsh, 1888
Subfamily: CERATOPIINAE Marsh, 1888 (sensu Abel, 1919)
Tribe: TRICERATOPINI nova
*Nedoceratops albertensis
= Triceratops albertensis