KANSAS
GRAHAM COUNTY
3 M S OF THE SOLOMON
BENTON FORMATION
TURONIAN STAGE
UPPERMOST GALLIC SUBEPOCH
LOWER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: EURYAPSIDA COLBERT, 1945
SUPERORDER: SAUROPTERYGIA OWEN 1860
GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994
ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
INFRAORDER: PLESIOSAUROMORPHA NOVA
EUPLESIOSAURIA O’KEEFE, 2001
Superfamily: CRYPTOCLEIDOIDEA Williston, 1925
TRICLEDIA O’KEEFE , 2001
Family: POLYCOTYLIDAE Cope, 1869
Trinacromerium bentonianum
= Trinacromerium anonymum
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KANSASGRAHAM COUNTY
LOW CHALK
NIOBRARA FORMATION
UPPER CONIACIAN STAGE
LOWERMOST SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: LACERTILIA OWEN, 1842
GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961
MICRORDER: SCINCOMORPHA CAMP, 1923
ORDER: ANGUIMORPHA FURBRINGER, 1900
SUBORDER: DIPLOGLOSSA COPE, 1864
INFRAORDER: THECAGLOSSA COPE, 1890
PARVORDER: MONSTERSAURIA NORELL & GAO, 1977
Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)
Family: MOSASAURIDAE Gervais, 1853
Infrafamily: NATANIA
Subfamily: TYLOSAURINAE Williston, 1897
Tylosaurus kansasensis
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KANSAS
GRAHAM COUNTY
NORTH BANK OF THE SMOKY HILL RIVER
SMOKY HILL MEMBER
NIOBRARA FORMATION
SANTONIAN OR CAMPANIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: LACERTILIA OWEN, 1842
GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961
MICRORDER: SCINCOMORPHA CAMP, 1923
ORDER: ANGUIMORPHA FURBRINGER, 1900
SUBORDER: DIPLOGLOSSA COPE, 1864
INFRAORDER: THECAGLOSSA COPE, 1890
PARVORDER: MONSTERSAURIA NORELL & GAO, 1977
Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)
Family: MOSASAURIDAE Gervais, 1853
Infrafamily: NATANIA
Subfamily: RUSSELLOSAURINAE
Tribe: PLIOPLATECARPINI Dollo, 1884 (sensu Russell, 1967)
*Platecarpus ictericus
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KANSAS
GRAHAM COUNTY
UPPER SMOKY HILL MEMBER
NIOBRARA FORMATION
SANTONIAN OR CAMPANIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
SUPERORDER: TAPINOPLATIA PETERS, 2000
GRANDORDER: CHARACIOPODA PETERS, 2000
MICRORDER: FENESTRASAURIA PETERS, 2000
ORDER: PTEROSAURIA KAUP, 1834
SUBORDER: EUPTERODACTYLOIDEA BENNETT, 1994
Superfamily: PTERANODONTIDEA Marsh, 1876
Family: PTERANODONTIDAE Marsh, 1876
Incertae sedis
non Pteranodon, P. velox
= Pterodactylus velox
= *Pteranodon comptus
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KANSAS
GRAHAM COUNTY
2 M NE PENOKEE
UPPER SMOKY HILL MEMBER
NIOBRARA FORMATION
SANTONIAN OR CAMPANIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
SUPERORDER: TAPINOPLATIA PETERS, 2000
GRANDORDER: CHARACIOPODA PETERS, 2000
MICRORDER: FENESTRASAURIA PETERS, 2000
ORDER: PTEROSAURIA KAUP, 1834
SUBORDER: EUPTERODACTYLOIDEA BENNETT, 1994
Superfamily: PTERANODONTIDEA Marsh, 1876
Family: PTERANODONTIDAE Marsh, 1876
Incertae sedis
Pteranodon sp
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KANSAS
GRAHAM COUNTY
BETWEEN BOGUE AND HILL CITY
UPPER SMOKY HILL MEMBER
NIOBRARA FORMATION
SANTONIAN OR CAMPANIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
SUPERORDER: TAPINOPLATIA PETERS, 2000
GRANDORDER: CHARACIOPODA PETERS, 2000
MICRORDER: FENESTRASAURIA PETERS, 2000
ORDER: PTEROSAURIA KAUP, 1834
SUBORDER: EUPTERODACTYLOIDEA BENNETT, 1994
Superfamily: PTERANODONTIDEA Marsh, 1876
Family: PTERANODONTIDAE Marsh, 1876
Incertae sedis
Pteranodon sp
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KANSAS
GRAHAM COUNTY
1 MILE WEST OF BOGUE
LOWER SMOKY HILL MEMBER
NIOBRARA FORMATION
SANTONIAN OR CAMPANIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
SUPERORDER: TAPINOPLATIA PETERS, 2000
GRANDORDER: CHARACIOPODA PETERS, 2000
MICRORDER: FENESTRASAURIA PETERS, 2000
ORDER: PTEROSAURIA KAUP, 1834
SUBORDER: EUPTERODACTYLOIDEA BENNETT, 1994
Superfamily: PTERANODONTIDEA Marsh, 1876
Family: PTERANODONTIDAE Marsh, 1876
*Geosternbergia sternbergi
= Pteranodon sternbergi
= Pteranodon (Sternbergia) sternbergi
= Pteranodon (Geosternbergia) sternbergi
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KANSAS
GRAHAM COUNTY
2 M NE OF PENOKEE
LOWER SMOKY HILL MEMBER
NIOBRARA FORMATION
SANTONIAN OR CAMPANIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
SUPERORDER: TAPINOPLATIA PETERS, 2000
GRANDORDER: CHARACIOPODA PETERS, 2000
MICRORDER: FENESTRASAURIA PETERS, 2000
ORDER: PTEROSAURIA KAUP, 1834
SUBORDER: EUPTERODACTYLOIDEA BENNETT, 1994
Superfamily: PTERANODONTIDEA Marsh, 1876
Family: PTERANODONTIDAE Marsh, 1876
*Geosternbergia walkeri
= Pteranodon (Sternbergia) walkeri
= Pteranodon walkeri
= Pteranodon (Geosternbergia) walkeri
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KANSAS
GRAHAM COUNTY
KUVP GRA-2
SMOKY HILL CHALK MEMBER
NIOBRARA FORMATION
CONIACIAN STAGE
LOWER SENONIAN SUBEPOCH
LOWER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AVES LINNAEUS, 1758
AVEBREVICAUDA PAUL, 2001
SUBCLASS: ODONTORNITHES MARSH, 1873
INFRACLASS: ODONTOHOLCAE NOVA
MAGNORDER: HESPERORNITHES
ORDER: HESPERORNITHIFORMES FURBRINGER, 1888
Family: HESPERORNITHIDAE Marsh, 1872
Subfamily: HESPERORNITHINAE Marsh, 1872
*Parahesperornis alexi
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KANSAS
GRAHAM COUNTY
NEAR SOLOMON RIVER
SMOKY HILL MEMBER
NIOBRARA FORMATION
CONIACIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AVES LINNAEUS, 1758
AVEBREVICAUDA PAUL, 2001
SUBCLASS: ODONTORNITHES MARSH, 1873
INFRACLASS: ODONTOTORMAE MARSH, 1880
ORDER: ICHTHYORNTHIFORMES MARSH, 1873
Family: ICHTHYORNITHIDAE Marsh, 1873
Ichthyornis dispar
= *Ichthyornis validus
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KANSAS
GRAHAM COUNTY
SMOKY HILL MEMBER
NIOBRARA FORMATION
CONIACIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AVES LINNAEUS, 1758
AVEBREVICAUDA PAUL, 2001
SUBCLASS: ODONTORNITHES MARSH, 1873
INFRACLASS: ODONTOTORMAE MARSH, 1880
ORDER: ICHTHYORNTHIFORMES MARSH, 1873
Family: ICHTHYORNITHIDAE Marsh, 1873
Ichthyornis dispar
= Ichthyornis victor
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