CHOUTEAU COUNTY

EGG CULTCH SITE (MOR LOCALTY TM. 7M-008

TWO MEDICINE FORMATION

JUDITHIAN AGE

UPPER CAMPANIAN STAGE

SENONIAN SUBEPOCH

UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: TETANURAE GAUTHIER, 1986

MICRORDER: COELUROSAURIA HUENE, 1914

ORDER: MANIRAPTORFORMES HOLTZ, 1996

AVETHEROPODA PAUL, 2001

SUBORDER: COELURIA MARSH, 1881

Superfamily: TYRANNOSAUROIDEA Osborn, 1906 (sensu Bonaparte et. al., 1990)

Family: TYRANNOSAURIDAE Osborn, 1906

Subfamily: TYRANNOSAURINAE Osborn, 1906

Tribe: TYRANNOSAURINI Osborn, 1906 (sensu Olshevsky, Ford & Yamamoto, 1995)

Daspletosaurus horneri

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SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: AMPHIBIA LINNAEUS, 1758

SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?

ORDER: ALLOCAUDATA FOX & NAYLOR, 1982

Family: ALBANERPETONTIDAE Fox & Naylor, 1982

Albanerpeton nexuosus

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: TETANURAE GAUTHIER, 1986

MICRORDER: COELUROSAURIA HUENE, 1914

ORDER: MANIRAPTORFORMES HOLTZ, 1996

AVETHEROPODA PAUL, 2001

SUBORDER: COELURIA MARSH, 1881

Superfamily: TYRANNOSAUROIDEA Osborn, 1906 (sensu Bonaparte et. al., 1990)

Family: TYRANNOSAURIDAE Osborn, 1906

Aublysodon mirandus
= Ornithomimus mirandus

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: AMPHIBIA LINNAEUS, 1758

SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?

ORDER: ALLOCAUDATA FOX & NAYLOR, 1982

Family: ALBANERPETONTIDAE Fox & Naylor, 1982

Gen. sp indet.

SUPERORDER: BATRACHIA BRONGNIART, 1800

ORDER: CAUDATA SCOPOLI, 1777

SUBORDER: PORTEOIDEA (NOBEL 1931) EMEND EDWARDS, 1976

Family: BATRACHOSAUROIDIDAE Auffenberg, 1958

Prodesmodon copei

SUBORDER: SALAMANDROIDEA NOBLE, 1931

Family: SIRENIDAE Gray, 1825

Habrosaurus dilatus
= Adelphesiren olivae

SUBCLASS: DIAPSIDA OSBORN, 1904

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA DUMERIL & BIBRON, 1839

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA OPPEL, 1811

SUPERORDER: LACERTILIA OWEN, 1842

GRANDORDER: ASCALABOTA GILMORE, 1928

ORDER: IGUANIA LATREILLE, 1825

SUBORDER: DRACOMORPHA NORTHCUTT, 1978

INFRAORDER: TEIUMORPHA ALIFANOV, 2000

Family: TEIIDAE Gray, 1827

Subfamily: CHAMOPSINAE Denton & O’Neill 1995

Leptochamops denticulatus

GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961

MICRORDER: SCINCOMORPHA CAMP, 1923

ORDER: ANGUIMORPHA FURBRINGER, 1900

SUBORDER: PLATYNOTA GILMORE, 1928

INFRAORDER: HELODERMOIDA COPE, 1900

Family: HELODERMATIDAE Gray, 1837

Paraderma bogerti

SUBORDER: DIPLOGLOSSA COPE, 1864

INFRAORDER: XENOSAUIRA FURBRINGER, 1900

Family: XENOSAURIDAE Cope, 1886

Exostinus lancensis

INFRAORDER: OPHISAUROMORPHA ALIFANOV, 2000

Family: ANGUIDAE Gray, 1825

Subfamily: GLYPTOSAURINAE Marsh, 1872

Tribe: MELANOSAURINI Sullivan, 1979

Odaxosaurus piger

INFRAORDER: THECAGLOSSA COPE, 1890

PARVORDER: MONSTERSAURIA NORELL & GAO, 1977

Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)

Family: PARASANIWIDAE Estes, 1964

Parasaniwa wyomingensis

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: TETANURAE GAUTHIER, 1986

MICRORDER: COELUROSAURIA HUENE, 1914

ORDER: MANIRAPTORFORMES HOLTZ, 1996

AVETHEROPODA PAUL, 2001

AVEPECTORA PAUL, 2001

SUBORDER: MANIRAPTORA GAUTHIER, 1986

INFRAORDER: PARVES SERENO, 1997

PARVORDER: DEINONYCHOSAURIA COLBERT & D. A. RUSSELL, 1969

Incertae sedis

Paronychodon lacustris

Family: DROMAEOSAURIDAE Russell, 1969

Subfamily: DROMAEOSAURINAE Matthew & Brown, 1922

Dromaeosaurus sp

Family: TROODONTIDAE Gilmore, 1924

Troodon formosus

COHORT: ORNITHISCHIFORMES COOPER, 1985

MAGNORDER: ORNITHISCHIA SEELEY, 1888

SUPERORDER: NEORNITHISCHIA COOPER, 1985

GRANDORDER: ORNITHOPODA MARSH, 1871

INCERTAE SEDIS

ORDER: PACHYCEPHALOSAURIA MARYANSKA & OSMOLSKA, 1974

SUBORDER: DOMOCEPHALINIA SERENO, 1986

INFRAORDER: PACHYCEPHALOSAURIA (MARYANSKA & OSMOLSKA, 1974) COOPER, 1985

Family: PACHYCEPHALOSAURIDAE Sternberg, 1945

Subfamily: STEGOCERINAE Sereno, 1998 (emend)

Stegoceras validum
= Stegoceras validus
= Troodon validus

MICRORDER: EUORNITHOPODA SERENO, 1986

ORDER: HYPSILOPHODONTIA COOPER, 1983

Family: HYPSILOPHODONTIDAE Dollo, 1882

Incertae sedis? Or new families? (pending further research)

Orodromeus makelai
= Thescelosaurus cf. neglectus

ORDER: IGUANODONTIA DOLLO, 1888

SUBORDER: ANKYLOPOLLEXIA SERENO, 1986

INFRAORDER: STYRACOSTERNA SERENO, 1986

PARVORDER: HADROSAURIFORMES SERENO, 1997

SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)

Family: HADROSAURIDAE COPE, 1869

Subfamily: HADROSAURINAE Cope, 1869 (sensu Lambe, 1918)

Tribe: HADROSAURINI Cope, 1869 (sensu Brett-Surman vide Chapman & Brett-Surman, 1990)

Hadrosaurus? breviceps

Subfamily: LAMBEOSAURINAE Parks, 1923

Tribe: CORYTHOSAURINI Brett-Surman vide Chapman & Brett-Surman, 1990

Lambeosaurus lambei
= Trachodon altidens
= Didanodon altidens
= Procheneosaurus altidens
= Pteropelyx altidens
= Thespesius altidens

MICRORDER: MARGINOCEPHALIA SERENO, 1986

ORDER: CERATOPIA MARSH, 1890

SUBORDER: NEOCERATOPIA SERENO, 1986

INFRAORDER: CORONOSAURIA SERENO, 1986

PARVORDER: CERATOPOMORPHA Wolfe & Kirkland, 1998 (emend)

Superfamily: CERATOPOIDEA Marsh, 1888 (sensu Hay, 1902)

Incertae sedis

Gen. sp indet.

SUPERORDER: EURYOPDA SERENO, 1986

GRANDORDER: ARMATOSAURIA ZHAO, 1983 (emend)

ORDER: ANKYLOSAURIA OSBORN, 1923

SUBORDER: SCELIDOSAURIA COOPER, 1985

INFRAORDER: ANKYLOSAURA OSBORN, 1923

Family: NODOSAURIDAE Marsh, 1890

Palaeoscincus costatus

CLASS: MAMMALIA LINNAEUS, 1758

SUBCLASS: THERIIFORMES (ROWE, 1988) McKENNA & BELL, 1997

INFRACLASS: ALLOTHERIA (MARSH, 1880) McKENNA & BELL, 1997

ORDER: MULTITUBERCULATA COPE, 1884

Incertae sedis

Gen. sp indet.

SUBORDER: CIMOLODONTA McKENNA, 1975

Incertae sedis

*Cimexomys judithae
*Cimexomys magnus

Paracimexomys priscus

Superfamily: PTILODONTOIDEA Cope, 1887 (sensu McKenna & Bell, 1997)

Family: CIMOLODONTIDAE Marsh, 1889

Cimolodon nitidus
Cimolodon sp

Family: PTILODONTIDAE Cope, 1887 (sensu McKenna & Bell, 1997)

Subfamily: NEOPLAGIAULACINAE Ameghino, 1890 (sensu McKenna & Bell, 1997)

Mesodma formosa
= Halodon formosus
Mesodma premaeva
= Ptilodus primaeuvs
Mesodma sp

Superfamily: Incertae sedis

Family: CIMOLOMYIDAE Marsh, 1889

Cimolomys clarki

Family: TAENIOLABIDIDAE Granger & Simpson, 1929

Meniscoessus major
= Cimolomys major

SUBCLASS: PROTOTHERIA GILL, 1872

INFRACLASS: TRECHNOTHERIA McKENNA, 1975

LEGION: CLADOTHERIA McKENNA, 1975

SUBLEGION: ZATHERIA MCKENNA, 1975

INFRALEGION: TRIBOSPHENIDA (McKENNA, 1975) McKENNA & BELL, 1997

SUPERCOHORT: THERIA (PARKER & HASWELL, 1987) McKENNA & BELL, 1997

COHORT: MARSUPIALIA (ILLIGER, 1811) McKENNA & BELL, 1997

ORDER: STAGODONTIA ARCHER, 1984

Family: STAGODONTIDAE Marsh, 1889

?Boreodon cutleri
= Eodelphis cutleri

Family: PEDIOMYIDAE Simpson, 1927

Pediomys sp

Protolambda clemensi
= *Pediomys clemensi

MAGNORDER: AMERIDELPHIA (SZALAY, in ARCHER, ED, 1982) McKENNA & BELL, 1997

ORDER: DIDELPHIMORPHIA GILL, 1872

Family: DIDELPHIDAE Gray 1821

Turgidodon praesagus
Turgidodon rhaister
= Alaphadon rhaister

Subfamily: ALPHADONTIDAE Marshall, Case & Woodburne, 1990

*Alphadon halleyi

Subfamily: ECTOCENTROCRISTINAE Rigby & Wolberg, 1987

*Ectocentrocristus foxi

COHORT: PLACENTALIA (OWEN, 1837) McKENNA & BELL, 1997

MAGNORDER: EPITHERIA (McKENNA, 1975) McKENNA & BELL, 1997

SUPERORDER: PREPTOTHERIA McKENNA, 1975

GRANDORDER: UNGULATOMORPHA ARCHIBALD, 1996

Family: GYPSONICTIPIDAE Van Valen, 1967

*Gypsonictops lewisi

OOSUPERCOHORT: DINOSAURIOVUM VIALOV, 1975

Incertae sedis

Gen. sp indet.

CHOUTEAU COUNTY

CLAYBALL HILL

JUDITH RIVER FORMATION

JUDITHIAN AGE

MIDDLE CAMPANIAN STAGE

MIDDLE SENONIAN SUBEPOCH

UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: MAMMALIA LINNAEUS, 1758

SUBCLASS: THERIIFORMES (ROWE, 1988) McKENNA & BELL, 1997

INFRACLASS: ALLOTHERIA (MARSH, 1880) McKENNA & BELL, 1997

ORDER: MULTITUBERCULATA COPE, 1884

SUBORDER: CIMOLODONTA McKENNA, 1975

Superfamily: Incertae sedis

Family: CIMOLOMYIDAE Marsh, 1889

*Cimolomys clarki

CHOUTEAU COUNTY

FORT BENTON

FROM THE VICINITY OF THE JUDITH RIVER

JUDITH RIVER FORMATION

JUDITHAN AGE

MIDDLE CAMPANIAN STAGE

MIDDLE SENONIAN SUBEPOCH

UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: TETANURAE GAUTHIER, 1986

MICRORDER: COELUROSAURIA HUENE, 1914

ORDER: MANIRAPTORFORMES HOLTZ, 1996

AVETHEROPODA PAUL, 2001

SUBORDER: COELURIA MARSH, 1881

Superfamily: TYRANNOSAUROIDEA Osborn, 1906 (sensu Bonaparte et. al., 1990)

Family: TYRANNOSAURIDAE Osborn, 1906

*Aublysodon mirandus
= Ornithomimus mirandus

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CHOUTEAU COUNTY

FORT BENTON

MISSOURI RIVER

JUDITH RIVER FORMATION

JUDITHAN AGE

MIDDLE CAMPANIAN STAGE

MIDDLE SENONIAN SUBEPOCH

UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: TETANURAE GAUTHIER, 1986

MICRORDER: COELUROSAURIA HUENE, 1914

ORDER: MANIRAPTORFORMES HOLTZ, 1996

AVETHEROPODA PAUL, 2001

SUBORDER: COELURIA MARSH, 1881

Superfamily: TYRANNOSAUROIDEA Osborn, 1906 (sensu Bonaparte et. al., 1990)

Family: TYRANNOSAURIDAE Osborn, 1906

*Deinodon horridus
= Gorgosaurus horridus
= Megalosaurus horridus

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MONTANA

CHOTEAU COUNTY

FORT BENTON

JUDITH RIVER FORMATION

JUDITHAN AGE

MIDDLE CAMPANIAN STAGE

MIDDLE SENONIAN SUBEPOCH

UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: ORNITHISCHIFORMES COOPER, 1985

MAGNORDER: ORNITHISCHIA SEELEY, 1888

SUPERORDER: EURYOPDA SERENO, 1986

GRANDORDER: ARMATOSAURIA ZHAO, 1983 (emend)

ORDER: ANKYLOSAURIA OSBORN, 1923

SUBORDER: SCELIDOSAURIA COOPER, 1985

INFRAORDER: ANKYLOSAURA OSBORN, 1923

Family: NODOSAURIDAE Marsh, 1890

*Palaeoscincus costatus

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CHOETAU COUNTY

CHOETAU

PEEBLES RANCH

JUDITH RIVER FORMATION

JUDITHAN AGE

MIDDLE CAMPANIAN STAGE

MIDDLE SENONIAN SUBEPOCH

UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: TETANURAE GAUTHIER, 1986

MICRORDER: COELUROSAURIA HUENE, 1914

ORDER: MANIRAPTORFORMES HOLTZ, 1996

AVETHEROPODA PAUL, 2001

SUBORDER: COELURIA MARSH, 1881

Superfamily: TYRANNOSAUROIDEA Osborn, 1906 (sensu Bonaparte et. al., 1990)

Family: TYRANNOSAURIDAE Osborn, 1906

Subfamily: ALBERTOSAURINAE Currie, Hurum & Sabath, 2003

Albertosaurus sp

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