PENNSYLVANIA
YORK COUNTY
NEAR YOCUMTOWN
LATE TRIASSIC
ICHNOSTEM: TETRAPODIPIDA NOVA
ICHNOSUPERCLASS: AMNIOTAIPIDII NOVA
ICHNOCLASS: REPTILIPEDIA VIALOV, 1966
ICHNOSUBCLASS: DIAPSIDIPIDII NOVA
ICHNOSTEM-GROUP: NEODIAPSIDIPEDII NOVA
ICHNOINFRACLASS: NEODIAPSIDIPEDII NOVA
ICHNOSUBDIVISION: ARCHOSAURIPEDIDII NOVA
ICHNOLEGION: DINOSAUROMORPHIPEDA NOVA
ICHNOSUBLEGION: DINOSAURIFORMEIPEDIDII NOVA
ICHNOSUPERCOHORT: DINOSAUROPEDIDA NOVA
Incerate sedis
Gen. sp indet.
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PENNSYLVANIAYORK COUNTY
LITTLE CONEWAGO CREEK AT ZIONS VIEW
NEW OXFORD FORMATION
NEWARK SUPERGROUP
LATE CARNIAN STAGE
LATE TRIASSIC EPOCH
LATE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
____________________________________________________________________________________SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LABYRINTHODONTIA OWEN, 1860
SUPERORDER: TEMNOSPONDYLIA ZITTEL, 1887
MICROORDER: LIMNARCHIA YATES & WARREN, 2000
ORDER: TREMATOSAURIA PIVETEAU, 1955
Superfamily: METOPOSAUROIDEA Watson, 1919
Family: METOPOSAURIDAE Watson, 1919
Subfamily: METOPOSAURINAE Watson, 1919
Koskinonodon perfectus
= Buettneria perfecta
PENNSYLANIA
YORK COUNTY
PHOENIXVILLE TUNNEL
LATE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
SUPERORDER: TAPINOPLATIA PETERS, 2000
GRANDORDER: CHARACIOPODA PETERS, 2000
MICRORDER: FENESTRASAURIA PETERS, 2000
ORDER: PTEROSAURIA KAUP, 1834
Incertae sedis
Gen. sp indet.
Note: may belong to a prolacertain
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PENNSYLVANIA
YORK COUNTY
NEWARK GROUP
LATE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
SUPERORDER: PARASUCHIMORPHA NOVA
ORDER: PARASUCHIA HUXLEY, 1875
Family: ANGISTORHINIDAE Huene, 1936
Subfamily: RUTIODONTINAE Long & Murry, 1995
Rutiodon carolinense
= Rutiodon manhattanensis
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PENNSYLVANIA
YORK COUNTY
BLACK LIMESTONE AND MARL IN THE PHOENIXVILLE TUNNEL
CARNIAN STAGE
LATE TRIASSIC EPOCH
LATE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
____________________________________________________________________________________SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
SUPERORDER: PARASUCHIMORPHA NOVA
ORDER: PARASUCHIA HUXLEY, 1875
Incertae sedis
Clepsysaurus pennsylvanicus
= Rutiodon pennsylvanicus
*Clepsysaurus veatleianus
= *Belodon priscus*Palaeoctonus appalachianus
*Palaeoctonus aulacodus
= Suchoprion aulacodus
*Suchoprion cyphodon
= Palaeoctonus cyphodon
*Suchoprion sulcidensFamily: ANGISTORHINIDAE Huene, 1936
Subfamily: RUTIODONTINAE Long & Murry, 1995
Rutiodon carolinense
= Belodon caroliensis
= Mystriosuchus caroliensis
= Palaeosaurus caroliensis
= Phytosaurus caroliensis
= Palaeosaurus carolinensis
PENNSYLVANIA
YORK COUNTY
ALONG THE LITTLE CONEWAGO CREEK
LATE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
SUPERORDER: PARASUCHIMORPHA NOVA
ORDER: PARASUCHIA HUXLEY, 1875
Incertae sedis
Genus et sp indet.
Family: ANGISTORHINIDAE Huene, 1936
Subfamily: RUTIODONTINAE Long & Murry, 1995
Rutiodon carolinense
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PENNSYLVANIA
YORK COUNTY
LITTLE CONEWAGA CREEK
LOC. 5
LATE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
SUPERORDER: PARASUCHIMORPHA NOVA
ORDER: PARASUCHIA HUXLEY, 1875
Family: ANGISTORHINIDAE Huene, 1936
Subfamily: RUTIODONTINAE Long & Murry, 1995
Rutiodon carolinense
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PENNSYLVANIA
YORK COUNTY
LITTLE CONEWAGA CREEK
LOC. 6
LATE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
SUPERORDER: PARASUCHIMORPHA NOVA
ORDER: PARASUCHIA HUXLEY, 1875
Family: ANGISTORHINIDAE Huene, 1936
Subfamily: RUTIODONTINAE Long & Murry, 1995
Rutiodon carolinense
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PENNSYLVANIA
YORK COUNTY
LITTLE CONEWAGA CREEK
LOC. 8
LATE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
SUPERORDER: PARASUCHIMORPHA NOVA
ORDER: PARASUCHIA HUXLEY, 1875
Family: ANGISTORHINIDAE Huene, 1936
Subfamily: RUTIODONTINAE Long & Murry, 1995
Rutiodon carolinense
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PENNSYLVANIA
YORK COUNTY
LITTLE CONEWAGA CREEK
LOC. 10
LATE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
SUPERORDER: PARASUCHIMORPHA NOVA
ORDER: PARASUCHIA HUXLEY, 1875
Family: ANGISTORHINIDAE Huene, 1936
Subfamily: RUTIODONTINAE Long & Murry, 1995
Rutiodon carolinense
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PENNSYLVANIA
YORK COUNTY
LITTLE CONEWAGA CREEK
LOC.11
LATE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
SUPERORDER: PARASUCHIMORPHA NOVA
ORDER: PARASUCHIA HUXLEY, 1875
Family: ANGISTORHINIDAE Huene, 1936
Subfamily: RUTIODONTINAE Long & Murry, 1995
Rutiodon carolinense
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PENNSYLVANIA
YORK COUNTY
PHOENIXVILLE TUNNEL
NEW OXFORD FORMATION
NEWARK SUPERGROUP
LATE CARNIAN STAGE
LATE TRIASSIC EPOCH
LATE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
SUPERORDER: SAUROPODOMORPHA HUENE, 1932
GRANDORDER: BRONTOSAURIA OLSHEVSKY, 1991 (emend)
MICRORDER: SAUROPODA MARSH, 1878
Family: ANCHISAURIDAE Marsh, 1885
Anchisaurus polyzelus
= *Palaeosaurus fraserianus
= Clepsysaurus fraserianus
= Palaeosauriscus fraserianus
= Thecodontosaurus fraserianus
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PENNSYLVANIA
YORK COUNTY
PHOENIXVILLE-TUNNEL
NEAR EMIGGSVILLE
NEW OXFORD FORMATION
NEWARK SUPERGROUP
LATE CARNIAN STAGE
LOWER LATE TRIASSIC EPOCH
LATE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: PSEUDOSUCHIA ZITTEL 1887-1890
GRANDORDER: RAUISUCHIFORMES PARRISH, 1993
*Galtonia gibbidens
= Thecodontosaurus gibbidens
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PENNSYLVANIA
YORK COUNTY
GETTYSBURG SHALE
LATE TRIASSIC
ICHNOSTEM: TETRAPODIPIDA NOVA
ICHNOSUPERCLASS: AMNIOTAIPIDII NOVA
ICHNOCLASS: REPTILIPEDIA VIALOV, 1966
ICHNOSUBCLASS: DIAPSIDIPIDII NOVA
ICHNOSTEM-GROUP: NEODIAPSIDIPEDII NOVA
ICHNOINFRACLASS: NEODIAPSIDIPEDII NOVA
ICHNOSUBDIVISION: ARCHOSAURIPEDIDII NOVA
ICHNOLEGION: DINOSAUROMORPHIPEDA NOVA
ICHNOSUBLEGION: DINOSAURIFORMEIPEDIDII NOVA
ICHNOSUPERCOHORT: DINOSAUROPEDIDA NOVA
Incerate sedis
Gen. sp indet.
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PENNSYLVANIA
YORK COUNTY
NEW CUMBERLAND QUADRANGLE
HESS’ OR WENTZ’S MILL ON BIG SPRING RUN ABOUT 1 MILE ENE OF YOCUMTOWN,
UPPER PART OF THE CONEWAGO FORMATION
NEW OXFORD OF STONES
LATE TRIASSIC
ICHNOSTEM: TETRAPODIPIDA NOVA
ICHNOSUPERCLASS: AMNIOTAIPIDII NOVA
ICHNOCLASS: REPTILIPEDIA VIALOV, 1966
ICHNOSUBCLASS: DIAPSIDIPIDII NOVA
ICHNOSTEM-GROUP: NEODIAPSIDIPEDII NOVA
ICHNOINFRACLASS: NEODIAPSIDIPEDII NOVA
ICHNOSUBDIVISION: ARCHOSAURIPEDIDII NOVA
ICHNOLEGION: DINOSAUROMORPHIPEDA NOVA
ICHNOSUBLEGION: DINOSAURIFORMEIPEDIDII NOVA
ICHNOSUPERCOHORT: DINOSAUROPEDIDA NOVA
ICHNOCOHORT: THEROPODIPEDIA NOVA
ICHNOSUPERORDER: NEOTHEROPODOPEDIDA NOVA
ICHNOORDER: NEOTETANURIPEDIDII NOVA
ICHNOSUBORDER: AVETHEROPODIPIDA NOVA
ICHNOMICRORDER: COELUROSAURIPIDIDA NOVA
Ichnofamily: GRALLATORIDAE emend Lull, 1904
Gen. sp indet.
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PENNSYLVANIA
YORK COUNTY
NEW CUMBERLAND QUADRANGLE
NEAR YOUCUMTOWN
UNSPECIFIED LOCALITY
LATE TRIASSIC
ICHNOSTEM: TETRAPODIPIDA NOVA
ICHNOSUPERCLASS: AMNIOTAIPIDII NOVA
ICHNOCLASS: REPTILIPEDIA VIALOV, 1966
ICHNOSUBCLASS: DIAPSIDIPIDII NOVA
ICHNOSTEM-GROUP: NEODIAPSIDIPEDII NOVA
ICHNOINFRACLASS: NEODIAPSIDIPEDII NOVA
ICHNOSUBDIVISION: ARCHOSAURIPEDIDII NOVA
ICHNOLEGION: DINOSAUROMORPHIPEDA NOVA
ICHNOSUBLEGION: DINOSAURIFORMEIPEDIDII NOVA
ICHNOSUPERCOHORT: DINOSAUROPEDIDA NOVA
ICHNOCOHORT: THEROPODIPEDIA NOVA
ICHNOSUPERORDER: NEOTHEROPODOPEDIDA NOVA
ICHNOORDER: NEOTETANURIPEDIDII NOVA
ICHNOSUBORDER: AVETHEROPODIPIDA NOVA
ICHNOMICRORDER: COELUROSAURIPIDIDA NOVA
Ichnofamily: GRALLATORIDAE emend Lull, 1904
Gen. sp indet.
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PENNSYLVANIA
YORK COUNTY
GETTYSBURG BASIN FROM NEAR GOLDSBORO
GETTYSBURG SHALE
LATE TRIASSIC
ICHNOSTEM: TETRAPODIPIDA NOVA
ICHNOSUPERCLASS: AMNIOTAIPIDII NOVA
ICHNOCLASS: REPTILIPEDIA VIALOV, 1966
ICHNOSUBCLASS: DIAPSIDIPIDII NOVA
ICHNOSTEM-GROUP: NEODIAPSIDIPEDII NOVA
ICHNOINFRACLASS: NEODIAPSIDIPEDII NOVA
ICHNOSUBDIVISION: ARCHOSAURIPEDIDII NOVA
ICHNOLEGION: DINOSAUROMORPHIPEDA NOVA
ICHNOSUBLEGION: DINOSAURIFORMEIPEDIDII NOVA
Incertae sedis
Ichnofamily: ATREIPODIDAE emend Olsen & Baird, 1986
Atreipus milfordensis = Grigandipus ? (Anchisauripus) milfordensis
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