SOUTH DAKOTA

BUTTE COUNTY

TOOTH DRAW DEPOSIT

HELL CREEK FORMATION

LANCIAN AGE

UPPER MAASTRICHTIAN STAGE

UPPERMOST SENONIAN SUBEPOCH

UPPER UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: AMPHIBIA LINNAEUS, 1758

SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?

SUPERORDER: BATRACHIA BRONGNIART, 1800

ORDER: CAUDATA SCOPOLI, 1777

SUBORDER: PORTEOIDEA (NOBEL 1931) EMEND EDWARDS, 1976

Family: BATRACHOSAUROIDIDAE Auffenberg, 1958

Opisthotriton sp

SUBORDER: AMBYSTOMATOIDEA NOBLE, 1931

Family: SCAPHERPETONTIDAE Auffenberg & Goin, 1959

Scapherpeton sp

SUBORDER: SALAMANDROIDEA NOBLE, 1931

Family: SIRENIDAE Gray, 1825

Habrosaurus sp

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: CHELONOMORPHA KUHN, 1960

SUPERCOHORT: TESTUDINES LINNAEUS, 1758

COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988

MAGANORDER: CRYPTODIRA GAFFNEY & MEYLAN, 1988

SUPERORDER: SELMACRYPTODIRA GAFFNEY, HUTCHISON, JENKINS & MEEKR, 1987

GRANDORDER: DAIOCRYPTODIRA GAFFNEY & MEYLAN, 1988

MICRORDER: BAENOIDEA WILLIAMS, 1960

Incertae sedis

Saxochelys sp

Family: BAENIDAE Cope, 1882

Peckemys sp

Subfamily: EUBAENINAE Gaffney, 1972

Plesiobaena sp

Family: NEURANKYLIDAE Lambe, 1902

Neurankylus sp

MICRORDER: CENTROCRYPTODIRA GAFFNEY & MEYLAN, 1988

ORDER: POLYCRPTODIRA GAFFNEY & MEYLAN, 1988

SUBORDER: CHELYDROIDEA AGASSIZ, 1857

Family: CHELYDRIDAE Agassiz, 1857

Tullochelys sp

SUBORDER: PROCOELOCRYPTODIRA GAFFNEY & MEYLAN, 1988

INFRAORDER: CHELOMACRYPTODIRA GAFFNEY & MEYLAN, 1988

PARVORDER: TRIONYCHIDA GRAY, 1870

Superfamily: TRIONYCHIDEA Bell, 1828 (sensu Gray, 1870)

Family: NANHSIUNGCHELYIDAE Yeh, 1966

Basilemys sp

Family: TRIONYCHIDAE Bell, 1828

Apalone sp

Aspideretoides sp

Axestemys sp

Gilmoremys sp

Family: ADOCIDAE Cope, 1870

Adocus sp

SUBCLASS: DIAPSIDA OSBORN, 1904

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA DUMERIL & BIBRON, 1839

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA OPPEL, 1811

SUPERORDER: LACERTILIA OWEN, 1842

GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961

MICRORDER: SCINCOMORPHA CAMP, 1923

ORDER: ANGUIMORPHA FURBRINGER, 1900

SUBORDER: DIPLOGLOSSA COPE, 1864

INFRAORDER: THECAGLOSSA COPE, 1890

PARVORDER: MONSTERSAURIA NORELL & GAO, 1977

Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)

Family: VARANIDAE Gray, 1827

Cemetarius sp

Palaeosaniwa sp

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUPERCOHORT CHORISTODERA NOVA

ORDER: CHORISTODERA COPE, 1876

Family: CHAMPSOSAURIDAE Cope, 1884

Champsosaurus cf C. laramiensis

SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990

COHORT: CROCODYLOTARSI BENTON & CLARK, 1988

MAGNORDER: SUCHIA KREBS, 1974

SUPERORDER: CROCODYLOMORPHA WALKER, 1968

GRANDORDER: CROCODILIFORMES HAY, 1930

MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983

ORDER: EUSUCHIA emend HUXLEY, 1875

Family: BOREALOSUCHIDAE emend BROCHU, 1999

Borealosuchus sp

INFRAORDER: CROCODYLIA emend GMELIN, 1877

Family: GAVIALIDAE Cuvier, 1807

Subfamily: THORACOSAURINAE Cope, 1875 (sensu Nopcsa, 1928)

Thoracosaurus sp

INFRAORDER: CROCODYLIA emend GMELIN, 1877

PARVORDER: BREVIROSTRES

Superfamily: ALLIGATORIDEA Buffetaut, 1985

GLOBIDONTA BROUCHU, 1999

Family: ALLIGATORIDAE Gray, 1844

Brachychampsa sp

SUPERCOHORT: AVEMETATARSALIA BENTON, 1999

COHORT: PROLACERTIFORMES CAMP, 1945 NOVA

SUPERORDER: TAPINOPLATIA PETERS, 2000

GRANDORDER: CHARACIOPODA PETERS, 2000

MICRORDER: FENESTRASAURIA PETERS, 2000

ORDER: PTEROSAURIA KAUP, 1834

SUBORDER: EUPTERODACTYLOIDEA BENNETT, 1994

Superfamily: ORNITHOCHEIROIDEA Seeley, 1870 (sensu Seeley, 1891)

Family: ORNITHOCHEIRIDAE Seeley, 1870

Incertae sedis

Gen. sp indet.

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

Incertae sedis

Ricardoestesia sp

Gen. sp indet.

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: TETANURAE GAUTHIER, 1986

MICRORDER: COELUROSAURIA HUENE, 1914

ORDER: MANIRAPTORFORMES HOLTZ, 1996

AVETHEROPODA PAUL, 2001

SUBORDER: COELURIA MARSH, 1881

Superfamily: TYRANNOSAUROIDEA Osborn, 1906 (sensu Bonaparte et. al., 1990)

Family: TYRANNOSAURIDAE Osborn, 1906

Subfamily: TYRANNOSAURINAE Osborn, 1906

Tribe: TYRANNOSAURINI Osborn, 1906 (sensu Olshevsky, Ford & Yamamoto, 1995)

Nanotyrannus sp

Tyrannosaurus rex

SUBORDER: ORNITHOMIMOSAURIA BARSBOLD, 1976

Family: ORNITHOMIMIDAE Marsh, 1890

Ornithomimus sp

AVEPECTORA PAUL, 2001

SUBORDER: MANIRAPTORA GAUTHIER, 1986

INFRAORDER: OVIRAPTOROSAURIA BARSBOLD, 1976

Family: CAENAGNATHIDAE Sternberg, 1940

Anzu sp

INFRAORDER: PARVES SERENO, 1997

PARVORDER: DEINONYCHOSAURIA COLBERT & D. A. RUSSELL, 1969

Incertae sedis

Paronychodon sp

Family: DROMAEOSAURIDAE Russell, 1969

Dakotaraptor steini.

Subfamily: VELOCIRAPTORINAE Barsbold, 1978

Acheroraptor sp

Saurornitholestes sp

Family: TROODONTIDAE Gilmore, 1924

Pectinodon sp

Troodon sp

SUBORDER: METORNITHES PERLE, NORELL, CHIAPPE & CLARK, 1993

INFRAORDER: ALVAREZSAURIA BONAPARTE, 1991

Family: ALVAREZSAURIDAE Bonaparte, 1991

Albertonykus sp

COHORT: ORNITHISCHIFORMES COOPER, 1985

MAGNORDER: ORNITHISCHIA SEELEY, 1888

Incertae sedis

Gen. sp indet.

SUPERORDER: NEORNITHISCHIA COOPER, 1985

GRANDORDER: ORNITHOPODA MARSH, 1871

MICRORDER: EUORNITHOPODA SERENO, 1986

ORDER: IGUANODONTIA DOLLO, 1888

Family: THESCELOSAURIDAE Sternberg, 1937

Thescelosaurus sp

SUBORDER: ANKYLOPOLLEXIA SERENO, 1986

INFRAORDER: STYRACOSTERNA SERENO, 1986

PARVORDER: HADROSAURIFORMES SERENO, 1997

SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)

Family: HADROSAURIDAE COPE, 1869

Subfamily: HADROSAURINAE Cope, 1869 (sensu Lambe, 1918)

Tribe: EDMONTOSAURINI Brett-Surman vide Chapman & Brett-Surman, 1990

Anatosaurus annectens

MICRORDER: MARGINOCEPHALIA SERENO, 1986

ORDER: PACHYCEPHALOSAURIA MARYANSKA & OSMOLSKA, 1974

SUBORDER: DOMOCEPHALINIA SERENO, 1986

INFRAORDER: PACHYCEPHALOSAURIA (MARYANSKA & OSMOLSKA, 1974) COOPER, 1985

Family: PACHYCEPHALOSAURIDAE Sternberg, 1945

Subfamily: PACHYCEPHALOSAURINAE Sternberg, 1945 (sensu Sereno, 1998)

Dracorex hogwartsia

Gen. sp indet.

ORDER: CERATOPIA MARSH, 1890

SUBORDER: NEOCERATOPIA SERENO, 1986

INFRAORDER: CORONOSAURIA SERENO, 1986

Family: PROTOCERATOPIDAE Granger & Gregory, 1923

Subfamily: LEPTOCERATOPINAE Nopcsa, 1923

Leptoceratops sp

PARVORDER: CERATOPOMORPHA Wolfe & Kirkland, 1998 (emend)

Superfamily: CERATOPOIDEA Marsh, 1888 (sensu Hay, 1902)

Family: CERATOPIDAE Marsh, 1888

Subfamily: CERATOPIINAE Marsh, 1888 (sensu Abel, 1919)

Tribe: TOROSAURINI Nopcsa, 1915

Torosaurus sp

Tribe: TRICERATOPINI nova

Triceratops sp

SUPERORDER: EURYOPDA SERENO, 1986

GRANDORDER: ARMATOSAURIA ZHAO, 1983 (emend)

ORDER: ANKYLOSAURIA OSBORN, 1923

INFRAORDER: ANKYLOSAURA OSBORN, 1923

Family: NODOSAURIDAE Marsh, 1890

Subfamily: EDMONTONIINAE Russell, 1940

Denverosauurs sp

Family: ANKYLOSAURIDAE Brown, 1908

Subfamily: ANKYLOSAURINAE Brown, 1908

Ankylosaurus magniventris

CLASS: AVES LINNAEUS, 1758

Incertae sedis

Gen sp indet.

CLASS: MAMMALIA LINNAEUS, 1758

SUBCLASS: THERIIFORMES (ROWE, 1988) McKENNA & BELL, 1997

INFRACLASS: ALLOTHERIA (MARSH, 1880) McKENNA & BELL, 1997

ORDER: MULTITUBERCULATA COPE, 1884

SUBORDER: CIMOLODONTA McKENNA, 1975

Superfamily: PTILODONTOIDEA Cope, 1887 (sensu McKenna & Bell, 1997)

Family: PTILODONTIDAE Cope, 1887 (sensu McKenna & Bell, 1997)

Subfamily: NEOPLAGIAULACINAE Ameghino, 1890 (sensu McKenna & Bell, 1997)

Mesodma sp

SUBCLASS: PROTOTHERIA GILL, 1872

INFRACLASS: TRECHNOTHERIA McKENNA, 1975

LEGION: CLADOTHERIA McKENNA, 1975

SUBLEGION: ZATHERIA MCKENNA, 1975

INFRALEGION: TRIBOSPHENIDA (McKENNA, 1975) McKENNA & BELL, 1997

SUPERCOHORT: THERIA (PARKER & HASWELL, 1987) McKENNA & BELL, 1997

COHORT: MARSUPIALIA (ILLIGER, 1811) McKENNA & BELL, 1997

ORDER: STAGODONTIA ARCHER, 1984

Family: STAGODONTIDAE Marsh, 1889

Didelphodon sp

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SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: CHELONOMORPHA KUHN, 1960

SUPERCOHORT: TESTUDINES LINNAEUS, 1758

COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988

MAGANORDER: CRYPTODIRA GAFFNEY & MEYLAN, 1988

SUPERORDER: SELMACRYPTODIRA GAFFNEY, HUTCHISON, JENKINS & MEEKR, 1987

GRANDORDER: DAIOCRYPTODIRA GAFFNEY & MEYLAN, 1988

MICRORDER: CENTROCRYPTODIRA GAFFNEY & MEYLAN, 1988

ORDER: POLYCRPTODIRA GAFFNEY & MEYLAN, 1988

SUBORDER: PROCOELOCRYPTODIRA GAFFNEY & MEYLAN, 1988

INFRAORDER: CHELONOIDEA GAFFNEY & MEYLAN, 1988

Superfamily: EUCHELONIOIDEA Gaffney & Meylan, 1988

Family: CHELONIIDAE Oppel, 1811

Gen. sp indet.

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SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: LACERTILIA OWEN, 1842

GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961

MICRORDER: SCINCOMORPHA CAMP, 1923

ORDER: ANGUIMORPHA FURBRINGER, 1900

SUBORDER: DIPLOGLOSSA COPE, 1864

INFRAORDER: THECAGLOSSA COPE, 1890

PARVORDER: MONSTERSAURIA NORELL & GAO, 1977

Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)

Family: MOSASAURIDAE Gervais, 1853

Incertae sedis

Gen. sp indet.

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SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: LACERTILIA OWEN, 1842

GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961

MICRORDER: SCINCOMORPHA CAMP, 1923

ORDER: ANGUIMORPHA FURBRINGER, 1900

SUBORDER: DIPLOGLOSSA COPE, 1864

INFRAORDER: THECAGLOSSA COPE, 1890

PARVORDER: MONSTERSAURIA NORELL & GAO, 1977

Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)

Family: MOSASAURIDAE Gervais, 1853

Incertae sedis

Gen. sp indet.

INFRACLASS: EURYAPSIDA COLBERT, 1945

SUPERORDER: SAUROPTERYGIA OWEN 1860

GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994

ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835

SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835

INFRAORDER: PLESIOSAUROMORPHA NOVA

EUPLESIOSAURIA O’KEEFE, 2001

Superfamily: CRYPTOCLEIDOIDEA Williston, 1925

TRICLEDIA O’KEEFE , 2001

Family: POLYCOTYLIDAE Cope, 1869

Gen. sp indet.

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SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: LACERTILIA OWEN, 1842

GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961

MICRORDER: SCINCOMORPHA CAMP, 1923

ORDER: ANGUIMORPHA FURBRINGER, 1900

SUBORDER: DIPLOGLOSSA COPE, 1864

INFRAORDER: THECAGLOSSA COPE, 1890

PARVORDER: MONSTERSAURIA NORELL & GAO, 1977

Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)

Family: MOSASAURIDAE Gervais, 1853

Subfamily: DOLICHOSAURIDAE Gervais, 1852

Tribe: DOLICHOSAURINI Gervais, 1852

Coniasaurus crassidens

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: LACERTILIA OWEN, 1842

GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961

MICRORDER: SCINCOMORPHA CAMP, 1923

ORDER: ANGUIMORPHA FURBRINGER, 1900

SUBORDER: DIPLOGLOSSA COPE, 1864

INFRAORDER: THECAGLOSSA COPE, 1890

PARVORDER: MONSTERSAURIA NORELL & GAO, 1977

Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)

Family: MOSASAURIDAE Gervais, 1853

Subfamily: DOLICHOSAURIDAE Gervais, 1852

Tribe: DOLICHOSAURINI Gervais, 1852

Coniasaurus crassidens

INFRACLASS: EURYAPSIDA COLBERT, 1945

SUPERORDER: SAUROPTERYGIA OWEN, 1860

GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994

ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835

SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835

INFRAORDER: PLIOSAUROMORPHA NOVA

Superfamily: CRYPTOCLEIDOIDEA Williston, 1925

TRICLEDIA O’KEEFE , 2001

Family: POLYCOTYLIDAE Cope, 1869

Subfamily: PALMULA Albright, Gillette & Titus, 2007a

Genus: Nova

Superfamily: PLIOSAUROIDEA Seeley, 1874

Family: PLIOSAURIDAE Seeley, 1874

Polyptychodon interruptus

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SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: LACERTILIA OWEN, 1842

GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961

MICRORDER: SCINCOMORPHA CAMP, 1923

ORDER: ANGUIMORPHA FURBRINGER, 1900

SUBORDER: DIPLOGLOSSA COPE, 1864

INFRAORDER: THECAGLOSSA COPE, 1890

PARVORDER: MONSTERSAURIA NORELL & GAO, 1977

Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)

Family: MOSASAURIDAE Gervais, 1853

Subfamily: DOLICHOSAURIDAE Gervais, 1852

Tribe: DOLICHOSAURINI Gervais, 1852

Coniasaurus crassidens

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

INFRACLASS: EURYAPSIDA COLBERT, 1945

SUPERORDER: SAUROPTERYGIA OWEN, 1860

GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994

ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835

SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835

Family: Nova?

Gen sp nova?

(cross between pliosaurs and polycotylids)

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: ORNITHISCHIFORMES COOPER, 1985

MAGNORDER: ORNITHISCHIA SEELEY, 1888

SUPERORDER: NEORNITHISCHIA COOPER, 1985

GRANDORDER: ORNITHOPODA MARSH, 1871

MICRORDER: EUORNITHOPODA SERENO, 1986

ORDER: IGUANODONTIA DOLLO, 1888

Family: THESCELOSAURIDAE Sternberg, 1937

Thescelosaurus neglectus

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SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: TETANURAE GAUTHIER, 1986

MICRORDER: COELUROSAURIA HUENE, 1914

ORDER: MANIRAPTORFORMES HOLTZ, 1996

AVETHEROPODA PAUL, 2001

SUBORDER: COELURIA MARSH, 1881

Superfamily: TYRANNOSAUROIDEA Osborn, 1906 (sensu Bonaparte et. al., 1990)

Family: TYRANNOSAURIDAE Osborn, 1906

Subfamily: TYRANNOSAURINAE Osborn, 1906

Tribe: TYRANNOSAURINI Osborn, 1906 (sensu Olshevsky, Ford & Yamamoto, 1995)

Tyrannosaurus rex

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NE BUTTE COUNTY

RUBY SITE

HELL CREEK FORMATION

LANCIAN AGE

UPPER MAASTRICHTIAN STAGE

UPPERMOST SENONIAN SUBEPOCH

UPPER UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: TETANURAE GAUTHIER, 1986

MICRORDER: COELUROSAURIA HUENE, 1914

ORDER: MANIRAPTORFORMES HOLTZ, 1996

AVETHEROPODA PAUL, 2001

SUBORDER: COELURIA MARSH, 1881

Superfamily: TYRANNOSAUROIDEA Osborn, 1906 (sensu Bonaparte et. al., 1990)

Family: TYRANNOSAURIDAE Osborn, 1906

Subfamily: ALBERTOSAURINAE Currie, Hurum & Sabath, 2003

Gorgosaurus sp

Subfamily: TYRANNOSAURINAE Osborn, 1906

Tribe: TYRANNOSAURINI Osborn, 1906 (sensu Olshevsky, Ford & Yamamoto, 1995)

Tyrannosaurus rex
Tyrannosaurus sp

COHORT: ORNITHISCHIFORMES COOPER, 1985

MAGNORDER: ORNITHISCHIA SEELEY, 1888

SUPERORDER: NEORNITHISCHIA COOPER, 1985

GRANDORDER: ORNITHOPODA MARSH, 1871

MICRORDER: EUORNITHOPODA SERENO, 1986

ORDER: IGUANODONTIA DOLLO, 1888

Family: THESCELOSAURIDAE Sternberg, 1937

Thescelosaurus sp

SUBORDER: ANKYLOPOLLEXIA SERENO, 1986

INFRAORDER: STYRACOSTERNA SERENO, 1986

PARVORDER: HADROSAURIFORMES SERENO, 1997

SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)

Family: HADROSAURIDAE COPE, 1869

Incertae sedis

Gen. sp indet.

MICRORDER: MARGINOCEPHALIA SERENO, 1986

ORDER: PACHYCEPHALOSAURIA MARYANSKA & OSMOLSKA, 1974

SUBORDER: DOMOCEPHALINIA SERENO, 1986

INFRAORDER: PACHYCEPHALOSAURIA (MARYANSKA & OSMOLSKA, 1974) COOPER, 1985

Family: PACHYCEPHALOSAURIDAE Sternberg, 1945

Gen. sp indet.

ORDER: CERATOPIA MARSH, 1890

SUBORDER: NEOCERATOPIA SERENO, 1986

INFRAORDER: CORONOSAURIA SERENO, 1986

PARVORDER: CERATOPOMORPHA Wolfe & Kirkland, 1998 (emend)

Superfamily: CERATOPOIDEA Marsh, 1888 (sensu Hay, 1902)

Family: CERATOPIDAE Marsh, 1888

Subfamily: CERATOPIINAE Marsh, 1888 (sensu Abel, 1919)

Tribe: TRICERATOPINI nova

Triceratops horridus
Triceratops sp

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