SOUTH DAKOTA

CUSTER COUNTY

SHARON SPRINGS MEMBER

LOWER PIERRE SHALE FORMATION

LOWER MIDDLE CAMPANIAN STAGE

SENONIAN SUBEPOCH

UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: LACERTILIA OWEN, 1842

GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961

MICRORDER: SCINCOMORPHA CAMP, 1923

ORDER: ANGUIMORPHA FURBRINGER, 1900

SUBORDER: DIPLOGLOSSA COPE, 1864

INFRAORDER: THECAGLOSSA COPE, 1890

PARVORDER: MONSTERSAURIA NORELL & GAO, 1977

Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)

Family: MOSASAURIDAE Gervais, 1853

Infrafamily: NATANIA

Subfamily: RUSSELLOSAURINAE Bell, 1993

Tribe: PLIOPLATECARPINI Dollo, 1884 (sensu Russell, 1967)

Latoplatecarpus nichollsae
= Plioplatecarpus nichollsae

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SOUTH DAKOTA

CUSTER COUNTY

HESPERORNIS LOCALITY

SDSM LOCALITY V8121

SHARON SPRINGS FORMATION

LOWER PIERRE SHALE GROUP

LOWER CAMPANIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUPERCOHORT: AVEMETATARSALIA BENTON, 1999

COHORT: PROLACERTIFORMES CAMP, 1945 NOVA

SUPERORDER: TAPINOPLATIA PETERS, 2000

GRANDORDER: CHARACIOPODA PETERS, 2000

MICRORDER: FENESTRASAURIA PETERS, 2000

ORDER: PTEROSAURIA KAUP, 1834

SUBORDER: EUPTERODACTYLOIDEA BENNETT, 1994

Superfamily: PTERANODONTIDEA Marsh, 1876

Family: PTERANODONTIDAE Marsh, 1876

Pteranodon longiceps

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SOUTH DAKOTA

CUSTER COUNTY

SOUTH FORK OF CHEYENNE RIVER

PIERRE FORMATION

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: CHELONOMORPHA KUHN, 1960

SUPERCOHORT: TESTUDINES LINNAEUS, 1758

COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988

MAGANORDER: CRYPTODIRA GAFFNEY & MEYLAN, 1988

SUPERORDER: SELMACRYPTODIRA GAFFNEY, HUTCHISON, JENKINS & MEEKR, 1987

GRANDORDER: DAIOCRYPTODIRA GAFFNEY & MEYLAN, 1988

MICRORDER: CENTROCRYPTODIRA GAFFNEY & MEYLAN, 1988

ORDER: POLYCRPTODIRA GAFFNEY & MEYLAN, 1988

SUBORDER: PROCOELOCRYPTODIRA GAFFNEY & MEYLAN, 1988

INFRAORDER: CHELONOIDEA GAFFNEY & MEYLAN, 1988

Superfamily: DERMOCHELYOIDEA Willams, 1950

Family: PROTOSTEGIDAE Cope, 1889

Subfamily: PROTOSTEGINAE Cope, 1889

Tribe: PROTOSTEGINI Cope, 1889 (sensu Hooks, 1998)

Subtribe: PROTOSTEGINA Cope, 1889 (sensu Hooks, 1998)

*Archelon ischyros
= Protostega ischyros

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SOUTH DAKOTA

CUSTER COUNTY

W SIDE NW 1/4 SW 1/4

SHARON SPRINGS

LOWER PIERRE

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: LACERTILIA OWEN, 1842

GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961

MICRORDER: SCINCOMORPHA CAMP, 1923

ORDER: ANGUIMORPHA FURBRINGER, 1900

SUBORDER: DIPLOGLOSSA COPE, 1864

INFRAORDER: THECAGLOSSA COPE, 1890

PARVORDER: MONSTERSAURIA NORELL & GAO, 1977

Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)

Family: MOSASAURIDAE Gervais, 1853

Infrafamily: NATANIA

Subfamily: RUSSELLOSAURINAE Bell, 1993

Tribe: PLIOPLATECARPINI Dollo, 1884 (sensu Russell, 1967)

Platecarpus somenensis

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SOUTH DAKOTA

CUSTER COUNTY

CHEYENNE RIVER

PIERRE FORMATION

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: LACERTILIA OWEN, 1842

GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961

MICRORDER: SCINCOMORPHA CAMP, 1923

ORDER: ANGUIMORPHA FURBRINGER, 1900

SUBORDER: DIPLOGLOSSA COPE, 1864

INFRAORDER: THECAGLOSSA COPE, 1890

PARVORDER: MONSTERSAURIA NORELL & GAO, 1977

Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)

Family: MOSASAURIDAE Gervais, 1853

Infrafamily: NATANIA

Subfamily: RUSSELLOSAURINAE Bell, 1993

Tribe: PLIOPLATECARPINI Dollo, 1884 (sensu Russell, 1967)

Prognathodon overtoni

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SOUTH DAKOTA

CUSTER COUNTY

CHEYENNE RIVER

BEARPAW FORMATION

MIDDLE CAMPANIAN STAGE

UPPER SENONIAN SUBEPOCH

UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: LACERTILIA OWEN, 1842

GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961

MICRORDER: SCINCOMORPHA CAMP, 1923

ORDER: ANGUIMORPHA FURBRINGER, 1900

SUBORDER: DIPLOGLOSSA COPE, 1864

INFRAORDER: THECAGLOSSA COPE, 1890

PARVORDER: MONSTERSAURIA NORELL & GAO, 1977

Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)

Family: MOSASAURIDAE Gervais, 1853

Infrafamily: NATANIA

Subfamily: MOSASAURINAE Gervais, 1853 (sensu Williston, 1897)

Tribe: MOSASAURINI Gervais, 1853 (sensu Russell, 1967)

Mosasaurus missouriensis
= *Mosasaurus horridus

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SOUTH DAKOTA

CUSTER COUNTY

7 1/2 M S OF FAIRBURN

UPPER SHARON SPRINGS

PIEERE SHALE

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: LACERTILIA OWEN, 1842

GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961

MICRORDER: SCINCOMORPHA CAMP, 1923

ORDER: ANGUIMORPHA FURBRINGER, 1900

SUBORDER: DIPLOGLOSSA COPE, 1864

INFRAORDER: THECAGLOSSA COPE, 1890

PARVORDER: MONSTERSAURIA NORELL & GAO, 1977

Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)

Family: MOSASAURIDAE Gervais, 1853

Infrafamily: NATANIA

Subfamily: MOSASAURINAE Gervais, 1853 (sensu Williston, 1897)

Tribe: GLOBIDENTINI Dollo, 1924 (sensu Russell, 1967)

*Globidens dakotensis

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SOUTH DAKOTA

CUSTER COUNTY

8 M S OF FAIRBURN

LOWER PIERRE FORMATION

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: LACERTILIA OWEN, 1842

GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961

MICRORDER: SCINCOMORPHA CAMP, 1923

ORDER: ANGUIMORPHA FURBRINGER, 1900

SUBORDER: DIPLOGLOSSA COPE, 1864

INFRAORDER: THECAGLOSSA COPE, 1890

PARVORDER: MONSTERSAURIA NORELL & GAO, 1977

Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)

Family: MOSASAURIDAE Gervais, 1853

Infrafamily: NATANIA

Subfamily: RUSSELLOSAURINAE Bell, 1993

Tribe: PLIOPLATECARPINI Dollo, 1884 (sensu Russell, 1967)

Platecarpus sp

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SOUTH DAKOTA

CUSTER COUNTY

FAIRBURN

SECOND CALCARENITE FROM BASE

ORMAN LAKE MEMBER

GREENHORN FORMATION

DUNVEGANOCERAS PONDI ZONE

EARLY UPPER CENOMANIAN STAGE

UPPER MIDDLE GALLIC SUBEPOCH

LOWERMOST EARLY CRETACEOUS EPOCH

MIDDLE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

INFRACLASS: EURYAPSIDA COLBERT, 1945

SUPERORDER: SAUROPTERYGIA OWEN, 1860

GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994

ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835

SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835

INFRAORDER: PLIOSAUROMORPHA NOVA

Superfamily: PLIOSAUROIDEA Seeley, 1874

Family: PLIOSAURIDAE Seeley, 1874

Polyptychodon interruptus

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SOUTH DAKOTA

CUSTER COUNTY

BUFFALO GAP

CALICO CANYON

LAKOTA FORMATION

BARREMIAN STAGE

LOWER GALLIC SUBEPOCH

GULF EPOCH

EARLY CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: ORNITHISCHIFORMES COOPER, 1985

MAGNORDER: ORNITHISCHIA SEELEY, 1888

SUPERORDER: NEORNITHISCHIA COOPER, 1985

GRANDORDER: ORNITHOPODA MARSH, 1871

MICRORDER: EUORNITHOPODA SERENO, 1986

ORDER: IGUANODONTIA DOLLO, 1888

SUBORDER: ANKYLOPOLLEXIA SERENO, 1986

INFRAORDER: STYRACOSTERNA SERENO, 1986

Incertae sedis

Osmakasaurus depresus
= *Camptosaurus? depressus

SUPERORDER: EURYOPDA SERENO, 1986

GRANDORDER: ARMATOSAURIA ZHAO, 1983 (emend)

ORDER: ANKYLOSAURIA OSBORN, 1923

SUBORDER: SCELIDOSAURIA COOPER, 1985

INFRAORDER: ANKYLOSAURA OSBORN, 1923

Family: POLACANTHIDAE Weiland, 1911

*Hoplitosaurus marshi
= Stegosaurus marshi
= Polacanthus marshi


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SOUTH DAKOTA

CUSTER COUNTY

NEARLY OPPOSITE CORRAL DRAW

W SIDE OF CHEYENNE RIVER

PIERRE FORMATION

CAMPANIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: ORNITHISCHIFORMES COOPER, 1985

MAGNORDER: ORNITHISCHIA SEELEY, 1888

SUPERORDER: NEORNITHISCHIA COOPER, 1985

GRANDORDER: ORNITHOPODA MARSH, 1871

MICRORDER: EUORNITHOPODA SERENO, 1986

ORDER: IGUANODONTIA DOLLO, 1888

SUBORDER: ANKYLOPOLLEXIA SERENO, 1986

INFRAORDER: STYRACOSTERNA SERENO, 1986

PARVORDER: HADROSAURIFORMES SERENO, 1997

SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)

Family: HADROSAURIDAE COPE, 1869

Subfamily: HADROSAURINAE Cope, 1869 (sensu Lambe, 1918)

Tribe: EDMONTOSAURINI Brett-Surman vide Chapman & Brett-Surman, 1990

Anatosaurus annectens
= *Claosaurus affinis
= Trachodon affinis


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SOUTHWESTERN SOUTH DAKOTA

CUSTER COUNTY

BUFFALO GAP

SHARON SPRINGS (LOWER) MEMBER)

PIERRE SHALE FORMATION

CAMPANIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: AVES LINNAEUS, 1758

AVEBREVICAUDA PAUL, 2001

SUBCLASS: ODONTORNITHES MARSH, 1873

INFRACLASS: ODONTOHOLCAE NOVA

MAGNORDER: HESPERORNITHES

ORDER: HESPERORNITHIFORMES FURBRINGER, 1888

Family: HESPERORNITHIDAE Marsh, 1872

Subfamily: HESPERORNITHINAE Marsh, 1872

*Hesperornis macdonaldi

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SOUTH DAKOTA

CUSTER COUNTY

5 MILES BY ROAD AND 2 3/4 MILES IN A STRAIGHT LINE S. 60?W OF HERMOSA

GRACE COOLIDGE CREEK SITE (FORMERLY SQUAW CREEK)

LAKOTA SANDSTONE

BARREMIAN-LOWER APTIAN STAGE

LOWER GALLIC SUBEPOCH

MIDDLE EARLY CRETACEOUS EPOCH

LATE CRETACEOUS

ICHNOSTEM: TETRAPODIPIDA NOVA

ICHNOSUPERCLASS: AMNIOTAIPIDII NOVA

ICHNOCLASS: REPTILIPEDIA VIALOV, 1966

ICHNOSUBCLASS: DIAPSIDIPIDII NOVA

ICHNOSTEM-GROUP: NEODIAPSIDIPEDII NOVA

ICHNOINFRACLASS: NEODIAPSIDIPEDII NOVA

ICHNOSUBDIVISION: ARCHOSAURIPEDIDII NOVA

ICHNOSUPERCOHORT: AVEMETATARSALIPEDII NOVA

ICHNOLEGION: DINOSAUROMORPHIPEDA NOVA

ICHNOSUBLEGION: DINOSAURIFORMEIPEDIDII NOVA

Incertae sedis

Ichnofamily: ATREIPODIDAE emend Olsen & Baird, 1986

Atreipus sp

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