HOWARD COUNTY
DOCKUM FORMATION
CARNIAN STAGE
LATE TRIASSIC EPOCH
LATE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LABYRINTHODONTIA OWEN, 1860
SUPERORDER: TEMNOSPONDYLIA ZITTEL, 1887
Incertae sedis
Dictyocephalus elegans
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
SUPERORDER: PARASUCHIMORPHA NOVA
ORDER: PARASUCHIA HUXLEY, 1875
Family: MYSTRIOSUCHIDAE Huene, 1915
Subfamily: MESORHININAE Huene, 1936
Parasuchus sp
= Palaeorhinus sp
_____________________________________________________________________________________
TEXAS
HOWARD COUNTY
BETWEEN DEVILS CREEK AND HYMAN SCOTT OIL FIELD
DOCKUM FORMATION
CARNIAN-?RHAETIAN STAGE
LATE TRIASSIC EPOCH
LATE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
_____________________________________________________________________________________SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: PSEUDOSUCHIA ZITTEL 1887-1890
GRANDORDER: RAUISUCHIFORMES PARRISH, 1993
ORDER: AETOSAURIA NICHOLSON & LYDEKKER, 1889
Family: STAGONOLEPIDIDAE Lydekker, 1887
Subfamily: DESMATOSUCHINAE Huene, 1942
Desmatosuchus sp
TEXAS
HOWARD COUNTY
3 M N OF OTIS CHALK
DOCKUM FORMATION
CARNIAN-?RHAETIAN STAGE
LATE TRIASSIC EPOCH
LATE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
_____________________________________________________________________________________SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: PSEUDOSUCHIA ZITTEL 1887-1890
GRANDORDER: RAUISUCHIFORMES PARRISH, 1993
ORDER: AETOSAURIA NICHOLSON & LYDEKKER, 1889
Family: STAGONOLEPIDIDAE Lydekker, 1887
Subfamily: DESMATOSUCHINAE Huene, 1942
Longosuchus meadei
= Typothorax meadei
TEXAS
HOWARD COUNTY
OTIS CHALK
DOCKUM FORMATION
CARNIAN STAGE
LATE TRIASSIC EPOCH
LATE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LABYRINTHODONTIA OWEN, 1860
SUPERORDER: TEMNOSPONDYLIA ZITTEL, 1887
MICROORDER: LIMNARCHIA YATES & WARREN, 2000
ORDER: TREMATOSAURIA PIVETEAU, 1955
Superfamily: METOPOSAUROIDEA Watson, 1919
Family: METOPOSAURIDAE Watson, 1919
Subfamily: APACHESAURINAE nova
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
MAGNORDER: PROTEROSUCHIFORMES nova
ORDER: PROTEROSUCHIA BROOM, 1906
Family: PROTEROSUCHIDAE Huene, 1914
Gen. sp indet.
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
SUPERORDER: PARASUCHIMORPHA NOVA
ORDER: PARASUCHIA HUXLEY, 1875
Family: MYSTRIOSUCHIDAE Huene, 1915
Subfamily: MESORHININAE Huene, 1936
Parasuchus sp
= Palaeorhinus spFamily: ANGISTORHINIDAE Huene, 1936
Subfamily: ANGISTORHININAE Huene, 1936 (sensu Camp, 1930)
Angistorhinus grandis
= Angistorhinus alticephalusMAGNORDER: SUCHIA KREBS, 1974
Family: SHUVOSAURIDAE Chatterjee, 1993
Shuvosaurus inexpectatus
= Chatterjeea elegansSUPERORDER: PSEUDOSUCHIA ZITTEL 1887-1890
GRANDORDER: RAUISUCHIFORMES PARRISH, 1993
ORDER: RAUISUCHIA Bonaparte, 1984
SUBORDER: RAUISUCHIA BONAPARTE, 1982
Family: POPOSAURIDAE Nopcsa, 1928
Poposaurus sp
_____________________________________________________________________________________
TEXAS
HOWARD COUNTY
NEAR OTIS CHALK
LOCALITY 9
COLORADO CITY FORMATION
CHINLE GROUP
OTISCHALKIAN
LATE CARNIAN STAGE
LOWER LATE TRIASSIC EPOCH
LATE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
SUPERORDER: PARASUCHIMORPHA NOVA
ORDER: PARASUCHIA HUXLEY, 1875
Family: ANGISTORHINIDAE Huene, 1936
Subfamily: ANGISTORHININAE Huene, 1936 (sensu Camp, 1930)
Angistorhinus grandis
_____________________________________________________________________________________
TEXAS
HOWARD COUNTY
OTIS CHALK
QUARRY 1
DOCKUM FORMATION
CARNIAN-?RHAETIAN STAGE
LATE TRIASSIC EPOCH
LATE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
SUPERORDER: PARASUCHIMORPHA NOVA
ORDER: PARASUCHIA HUXLEY, 1875
Family: MYSTRIOSUCHIDAE Huene, 1915
Subfamily: MESORHININAE Huene, 1936
Parasuchus bransoni
= Palaeorhinus bransoni
= Parasuchus (Paleorhinus) bransoniMAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: PSEUDOSUCHIA ZITTEL 1887-1890
GRANDORDER: RAUISUCHIFORMES PARRISH, 1993
ORDER: AETOSAURIA NICHOLSON & LYDEKKER, 1889
Family: STAGONOLEPIDIDAE Lydekker, 1887
Subfamily: DESMATOSUCHINAE Huene, 1942
Longosuchus meadei
= Typothorax meadei
_____________________________________________________________________________________
TEXAS
HOWARD COUNTY
OTIS CHALK
LOCALITY 2
DOCKUM FORMATION
CARNIAN-?RHAETIAN STAGE
LATE TRIASSIC EPOCH
LATE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
SUPERORDER: PARASUCHIMORPHA NOVA
ORDER: PARASUCHIA HUXLEY, 1875
Family: ANGISTORHINIDAE Huene, 1936
Subfamily: ANGISTORHININAE Huene, 1936 (sensu Camp, 1930)
Angistorhinus grandis
= Angistorhinus alticephalus
_____________________________________________________________________________________
TEXAS
HOWARD COUNTY
OTIS CHALK
LOCALITY 9
DOCKUM FORMATION
CARNIAN-?RHAETIAN STAGE
LATE TRIASSIC EPOCH
LATE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
SUPERORDER: PARASUCHIMORPHA NOVA
ORDER: PARASUCHIA HUXLEY, 1875
Family: ANGISTORHINIDAE Huene, 1936
Subfamily: ANGISTORHININAE Huene, 1936 (sensu Camp, 1930)
Angistorhinus grandis
= Angistorhinus alticephalus
_____________________________________________________________________________________
TEXAS
HOWARD COUNTY
ABOUT 20 M SE OF BIG SPRINGS
NEAR OTIS CHALK
DOCKUM FORMATION
CARNIAN-?RHAETIAN STAGE
LATE TRIASSIC EPOCH
LATE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
SUPERORDER: PARASUCHIMORPHA NOVA
ORDER: PARASUCHIA HUXLEY, 1875
Family: ANGISTORHINIDAE Huene, 1936
Subfamily: ANGISTORHININAE Huene, 1936 (sensu Camp, 1930)
*Brachysuchus megalodon
= Palaeorhinus megalodon
= Phytosaurus megalodon
_____________________________________________________________________________________
TEXAS
HOWARD COUNTY
N OF OTIS CHALK
NMMNHLOCALITY 3357
COLORADO CITY MEMBER
CHINLE GROUP
LATE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: PSEUDOSUCHIA ZITTEL 1887-1890
GRANDORDER: RAUISUCHIFORMES PARRISH, 1993
ORDER: AETOSAURIA NICHOLSON & LYDEKKER, 1889
Family: STAGONOLEPIDIDAE Lydekker, 1887
Subfamily: STAGONOLEPINAE Lydekker, 1887 (sensu Heckert & Lucas, 2000)
*Coahomasuchus kahleorum
_____________________________________________________________________________________
TEXAS
HOWARD COUNTY
256 MILES (40 KM) SOUTHEAST OF BIG SPRINGS
ROY LAMB RANCH
“PRE-TECOVAS HORIZON”
DOCKUM GROUP
MIDDLE CARNIAN STAGE
LOWER LATE TRIASSIC EPOCH
LATE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
SUPERORDER: PARASUCHIMORPHA NOVA
ORDER: PARASUCHIA HUXLEY, 1875
Family: ANGISTORHINIDAE Huene, 1936
Subfamily: ANGISTORHININAE Huene, 1936 (sensu Camp, 1930)
Angistorhinus grandis
= *Angistorhinus alticephalus
_____________________________________________________________________________________
TEXAS
HOWARD COUNTY
NMMNH LOCALITY 860 (TRILOPHOSAURUS QUARRIES)
COLORADO CITY MEMBER
DOCKUM FORMATION
CHINLE GROUP
UPPER OTISCHALKIAN
LATE CARNIAN STAGE
LOWER LATE TRIASSIC EPOCH
LATE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
Gen. sp indet.
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LABYRINTHODONTIA OWEN, 1860
SUPERORDER: TEMNOSPONDYLIA ZITTEL, 1887
MICROORDER: LIMNARCHIA YATES & WARREN, 2000
ORDER: TREMATOSAURIA PIVETEAU, 1955
Superfamily: METOPOSAUROIDEA Watson, 1919
Family: METOPOSAURIDAE Watson, 1919
Gen. sp indet.
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
Gen. sp indet.
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
ARCHOSAURIFORMES indet.
Protecovasaurus lucasi
_____________________________________________________________________________________
TEXAS
HOWARD COUNTY
TRILOPHOSAURUS QUARRY 1
OTIS CHALK LOCALITY
DOCKUM FORMATION
CARNIAN STAGE
LATE TRIASSIC EPOCH
LATE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LABYRINTHODONTIA OWEN, 1860
SUPERORDER: TEMNOSPONDYLIA ZITTEL, 1887
MICROORDER: LIMNARCHIA YATES & WARREN, 2000
ORDER: TREMATOSAURIA PIVETEAU, 1955
Superfamily: TREMATOSAUROIDEA Watson, 1919 (sensu Save-Soderbergh, 1935)
Family: ALMASAURIDAE Dutuit, 1972
Latiscopus sp
Superfamily: METOPOSAUROIDEA Watson, 1919
Incertae sedis
Metoposaurus sp
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: PARAREPTILIA OLSON 1947
INFRACLASS: SAUROPSIDA HUXLEY, 1864
ORDER: PROCOLOPHONOMORPHA ROMER, 1964
SUBORDER: PROCOLPHOMORPHA SEELEY, 1888
Family: PROCOLOPHONIDAE, Seeley, 1888
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
TEXAS
HOWARD COUNTY
NMMNH LOCALITY 3101
COLORADO CITY FORMATION
CHINLE GROUP
OTISCHALKIAN
MIDDLE CARNIAN STAGE
LOWER LATE TRIASSIC EPOCH
LATE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
CLADE NAME: EUCROCOPODA EZCURRA, 2016
ORDER: PROTEROCHAMPSIA EMEND BENTON & NORMAN, in BENTON, 1985
Family: DOSWELLIDAE Weems, 1980
*Ankylosuchus chinlegroupensis
_____________________________________________________________________________________
TEXAS
HOWARD COUNTY
TMM LOCALITY 31025
TMM SITE 1
TRILOPHOSAURUS SITE
“PRE-TECOVAS HORIZON”
DOCKUM GROUP
MIDDLE CARNIAN STAGE
LOWER LATE TRIASSIC EPOCH
LATE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LABYRINTHODONTIA OWEN, 1860
SUPERORDER: TEMNOSPONDYLIA ZITTEL, 1887
MICROORDER: LIMNARCHIA YATES & WARREN, 2000
ORDER: TREMATOSAURIA PIVETEAU, 1955
Superfamily: METOPOSAUROIDEA Watson, 1919
Family: METOPOSAURIDAE Watson, 1919
Subfamily: METOPOSAURINAE Watson, 1919
Koskinonodon perfectus
= Buettneria perfectaCLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
ORDER: TRILOPHOSAURIA ROMER, 1956
Family: TRILOPHOSAURIDAE Gregory, 1945
Trilophosaurus buettneri
SUPERCOHORT: RHYNOCEPHALIA nova
ORDER: RHYNOCEPHALIA BUTHER, 1867
SUBORDER: RHYNCHOSAUROIDEA GERVAIS, 1889
Family: RHYNCHOSAURIDAE Huxley, 1887
*Otischalkia elderae
= Dockumensia beckorumSUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
SUPERORDER: PARASUCHIMORPHA NOVA
ORDER: PARASUCHIA HUXLEY, 1875
Family: MYSTRIOSUCHIDAE Huene, 1915
Subfamily: MESORHININAE Huene, 1936
Parasuchus bransoni
= Palaeorhinus bransoni
= Parasuchus (Paleorhinus) bransoniMAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: PSEUDOSUCHIA ZITTEL 1887-1890
GRANDORDER: RAUISUCHIFORMES PARRISH, 1993
ORDER: RAUISUCHIA Bonaparte, 1984
SUBORDER: RAUISUCHIA BONAPARTE, 1982
Family: DOSWELLIDAE Weems, 1980
Doswellia kaltenbachi
Family: POPOSAURIDAE Nopcsa, 1928
Poposaurus langstoni
= *Lythrosuchus langstoni
_____________________________________________________________________________________
TEXAS
HOWARD COUNTY
TMM LOCALITY 31098
TMM SITE 2
“PRE’TECOVAS HORIZON”
DOCKUM GROUP
MIDDLE CARNIAN STAGE
LOWER LATE TRIASSIC EPOCH
LATE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LABYRINTHODONTIA OWEN, 1860
SUPERORDER: TEMNOSPONDYLIA ZITTEL, 1887
MICROORDER: LIMNARCHIA YATES & WARREN, 2000
ORDER: TREMATOSAURIA PIVETEAU, 1955
Superfamily: METOPOSAUROIDEA Watson, 1919
Family: METOPOSAURIDAE Watson, 1919
Subfamily: METOPOSAURINAE Watson, 1919
Koskinonodon perfectus
= Buettneria perfectaCLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
SUPERORDER: PARASUCHIMORPHA NOVA
ORDER: PARASUCHIA HUXLEY, 1875
Family: ANGISTORHINIDAE Huene, 1936
Subfamily: ANGISTORHININAE Huene, 1936 (sensu Camp, 1930)
Angistorhinus sp
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: PSEUDOSUCHIA ZITTEL 1887-1890
GRANDORDER: RAUISUCHIFORMES PARRISH, 1993
ORDER: RAUISUCHIA Bonaparte, 1984
SUBORDER: RAUISUCHIA BONAPARTE, 1982
Family: DOSWELLIDAE Weems, 1980
Doswellia kaltenbachi
Family: POPOSAURIDAE Nopcsa, 1928
Poposaurus sp
ORDER: AETOSAURIA NICHOLSON & LYDEKKER, 1889
Family: STAGONOLEPIDIDAE Lydekker, 1887
Incertae sedis
Gen. sp indet.
Subfamily: DESMATOSUCHINAE Huene, 1942
*Longosuchus meadei
_____________________________________________________________________________________
TEXAS
HOWARD COUNTY
TMM LOCALITY 31099
TMM QUARRY 2
“PRE’TECOVAS HORIZON”
DOCKUM GROUP
MIDDLE CARNIAN STAGE
LOWER LATE TRIASSIC EPOCH
LATE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LABYRINTHODONTIA OWEN, 1860
SUPERORDER: TEMNOSPONDYLIA ZITTEL, 1887
MICROORDER: LIMNARCHIA YATES & WARREN, 2000
ORDER: TREMATOSAURIA PIVETEAU, 1955
Superfamily: METOPOSAUROIDEA Watson, 1919
Family: METOPOSAURIDAE Watson, 1919
Subfamily: METOPOSAURINAE Watson, 1919
Koskinonodon perfectus
= Buettneria perfecta
_____________________________________________________________________________________
TEXAS
HOWARD COUNTY
TMM LOCALITY 31100
TMM SITE 3
COLORADO CITY FORMATION (“PRE’TECOVAS HORIZON”)
DOCKUM GROUP
MIDDLE CARNIAN STAGE
LOWER LATE TRIASSIC EPOCH
LATE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LABYRINTHODONTIA OWEN, 1860
SUPERORDER: TEMNOSPONDYLIA ZITTEL, 1887
MICROORDER: LIMNARCHIA YATES & WARREN, 2000
ORDER: TREMATOSAURIA PIVETEAU, 1955
Superfamily: METOPOSAUROIDEA Watson, 1919
Family: METOPOSAURIDAE Watson, 1919
Subfamily: METOPOSAURINAE Watson, 1919
Koskinonodon perfectus
= Buettneria perfecta
= *Buettneria howardensisGen. sp indet.
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
SUPERORDER: ORNITHOSUCHIMORPHA NOVA
ORDER: ORNITHOSUCHIA BONAPARTE 1971
Family: ORNITHOSUCHIDAE Huene, 1908
Incertae sedis
Gen. sp indet.
SUPERORDER: PARASUCHIMORPHA NOVA
ORDER: PARASUCHIA HUXLEY, 1875
Family: ANGISTORHINIDAE Huene, 1936
Subfamily: ANGISTORHININAE Huene, 1936 (sensu Camp, 1930)
Brachysuchus megalodon
= Palaeorhinus megalodon
= Phytosaurus megalodonFamily: MYSTRIOSUCHIDAE Huene, 1915
Subfamily: MESORHININAE Huene, 1936
Parasuchus bransoni
= Palaeorhinus bransoni
= Parasuchus (Paleorhinus) bransoniMAGNORDER: SUCHIA KREBS, 1974
Family: SHUVOSAURIDAE Chatterjee, 1993
Shuvosaurus inexpectatus
= Chatterjeea elegansSUPERORDER: PSEUDOSUCHIA ZITTEL 1887-1890
GRANDORDER: RAUISUCHIFORMES PARRISH, 1993
ORDER: RAUISUCHIA Bonaparte, 1984
SUBORDER: RAUISUCHIA BONAPARTE, 1982
Family: TERATOSAURIDAE Cope 1871
Subfamily: TERATOSAURINAE Cope 1871
Tribe: TERATOSAURINI Cope, 1871 (sensu Sennikov, 1995) (emend) from Tribe RAUISUCHINI Huene, 1942
Postosuchus kirkpatricki
Family: POPOSAURIDAE Nopcsa, 1928
Poposaurus gracilis
Poposaurus langstoni
= Lythrosuchus langstoni
Poposaurus spORDER: AETOSAURIA NICHOLSON & LYDEKKER, 1889
New genus of small aetosaur
New genus of a carnivorus aetosaur
Family: STAGONOLEPIDIDAE Lydekker, 1887
Subfamily: DESMATOSUCHINAE Huene, 1942
*Longosuchus meadei
= Typothorax meadei*Lucasuchus hunti
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
Family: LAGERPETONIDAE Arcucci, 1987
*Dromomeron gregorii
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
Incertae sedis
Gen. sp indet.
Family: LEWISUCHIDAE PAUL, 1988 emend Olshevsky, 1991
Gen. sp indet.
SUPERCOHORT: DINOSAURIA OWEN, 1842
SUPERORDER: HERRASAURA NOVA
ORDER: HERRERASAURIA GALTON, 1985
Family: STAURIKOSAURIDAE Galton, 1977
Chindesaurus bryansmalli
_____________________________________________________________________________________
TEXAS
HOWARD COUNTY
QUARRY 2
“PRE-TECOVAS” HORIZON
DOCKUM GROUP
CARNIAN STAGE
LOWER LATE TRIASSIC EPOCH
LATE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
ORDER: TRILOPHOSAURIA ROMER, 1956
Family: TRILOPHOSAURIDAE Gregory, 1945
Trilophosaurus buettneri
*Malerisaurus langstoniSUPERCOHORT: RHYNOCEPHALIA nova
ORDER: RHYNOCEPHALIA BUTHER, 1867
Incertae sedis
Gen. sp indet.
= Malerisaurus langstoni (partim)SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
SUPERORDER: PARASUCHIMORPHA NOVA
Incertae sedis
Gen. sp indet.
= Malerisaurus langstoni (partim)MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: PSEUDOSUCHIA ZITTEL 1887-1890
GRANDORDER: RAUISUCHIFORMES PARRISH, 1993
ORDER: AETOSAURIA NICHOLSON & LYDEKKER, 1889
Incertae sedis
Gen. sp indet.
= Malerisaurus langstoni (partim)
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TEXAS
HOWARD COUNTY
TMM LOCALITY 31185
TMM SITE 31100
QUARRY 3
DOCKUM GROUP
LATEST CARNIAN-EALRY NORIAN STAGE
LOWER LATE TRIASSIC EPOCH
LATE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
PARACROCODYLOMORPHA PARRSH, 1993 sensu WEINBAUM & HUNGERBUHLER, 2007
LORICATA MERREM, 1820 sensu NESBITT, 2011
GRANDORDER: RAUISUCHIFORMES PARRISH, 1993
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
TEXAS
HOWARD COUNTY
TMM LOCALITY 31185
TMM SITE 31100
QUARRY 3
DOCKUM GROUP
MIDDLE CARNIAN STAGE
LOWER LATE TRIASSIC EPOCH
LATE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
ARCHOSAURIFORMES Gauthier, 1984
PROTOPYKNOSIA NESBITT, STOCKER, CHATTERJEE, HORNER & GOODWIN, 2021
*Triopticus primus
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TEXAS
HOWARD COUNTY
TMM LOCALITY 31185
TMM SITE 3
QUARRY 3A
“PRE-TECOVAS HORIZON”
DOCKUM GROUP
MIDDLE CARNIAN STAGE
LOWER LATE TRIASSIC EPOCH
LATE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
SUPERORDER: PARASUCHIMORPHA NOVA
ORDER: PARASUCHIA HUXLEY, 1875
Family: MYSTRIOSUCHIDAE Huene, 1915
Subfamily: MESORHININAE Huene, 1936
Parasuchus bransoni
= Palaeorhinus bransoni
= Parasuchus (Paleorhinus) bransoniMAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: PSEUDOSUCHIA ZITTEL 1887-1890
GRANDORDER: RAUISUCHIFORMES PARRISH, 1993
ORDER: AETOSAURIA NICHOLSON & LYDEKKER, 1889
Family: STAGONOLEPIDIDAE Lydekker, 1887
Subfamily: DESMATOSUCHINAE Huene, 1942
Longosuchus meadei
= Typothorax meadeiLucasuchus hunti
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TEXAS
HOWARD COUNTY
QUARRY 3A
TMM SITE 3
“PRE’TECOVAS HORIZON”
DOCKUM GROUP
MIDDLE CARNIAN STAGE
LOWER LATE TRIASSIC EPOCH
LATE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LABYRINTHODONTIA OWEN, 1860
SUPERORDER: TEMNOSPONDYLIA ZITTEL, 1887
MICROORDER: LIMNARCHIA YATES & WARREN, 2000
ORDER: TREMATOSAURIA PIVETEAU, 1955
Superfamily: METOPOSAUROIDEA Watson, 1919
Family: METOPOSAURIDAE Watson, 1919
Subfamily: METOPOSAURINAE Watson, 1919
Koskinonodon perfectus
= Buettneria perfectaCLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: RHYNOCEPHALIA nova
ORDER: RHYNOCEPHALIA BUTHER, 1867
SUBORDER: RHYNCHOSAUROIDEA GERVAIS, 1889
Family: RHYNCHOSAURIDAE Huxley, 1887
Otischalkia elderae
= Dockumensia beckorum
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TEXAS
HOWARD COUNTY
TMM LOCALITY 41921
TMM SITE 3
“PRE’TECOVAS HORIZON”
DOCKUM GROUP
MIDDLE CARNIAN STAGE
LOWER LATE TRIASSIC EPOCH
LATE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LABYRINTHODONTIA OWEN, 1860
SUPERORDER: TEMNOSPONDYLIA ZITTEL, 1887
MICROORDER: LIMNARCHIA YATES & WARREN, 2000
ORDER: TREMATOSAURIA PIVETEAU, 1955
Superfamily: METOPOSAUROIDEA Watson, 1919
Family: METOPOSAURIDAE Watson, 1919
Subfamily: METOPOSAURINAE Watson, 1919
Koskinonodon perfectus
= Buettneria perfecta
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TEXAS
HOWARD COUNTY
TMM LOCALITY 42060
APPROXIMATELY 1.6 KM SOUTHWEST OF QUARRY 1
“PRE-TECOVAS HORIZON”
DOCKUM GROUP
MIDDLE CARNIAN STAGE
LOWER LATE TRIASSIC EPOCH
LATE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
SUPERORDER: PARASUCHIMORPHA NOVA
ORDER: PARASUCHIA HUXLEY, 1875
Family: ANGISTORHINIDAE Huene, 1936
Subfamily: ANGISTORHININAE Huene, 1936 (sensu Camp, 1930)
Angistorhinus sp
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TEXAS
HOWARD COUNTY
SITE 1 (TMM LOCALITY 31213)
LOS ESTERO MEMBER
SANTA ROSA FORMATION
ADAMANIAN
LOWER CARNIAN STAGE
LATE TRIASSIC EPOCH
LATE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
ORDER: TRILOPHOSAURIA ROMER, 1956
Family: TRILOPHOSAURIDAE Gregory, 1945
Trilophosaurus buettneri
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TEXAS
HOWARD COUNTY
JUST NORTHEAST OF THE CLASSIC OTIS CHALK
LOCALITIES
DOCKUM SITE 7 GENERAL (TMM LOCLAITY
41936)
OTIS CHALK AREA
DOCKUM GROUP
LOWER NORIAN STAGE?
MIDDLE LATE TRIASSIC EPOCH
LATE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: CERATOSAURIA NOVAS, 1992
ORDER: COELOPHYSIA NOVA
SUBORDER: COELOPHYSOIDEA (NOPCSA, 1928) HOLTZ, 1994
Incertae sedis
*Lepidus praecisio