WYOMING

NORTHERN NIOBRARA COUNTY

4 MILES WEST OF KUHMN 40001

JOHNSON RANCH

SHARON SPRINGS MEMBER (WAS REFERRED TO THE REDBIRD MEMBER)

PIERRE FORMATION

LATE CAMPANIAN STAGE

UPPER SENONIAN SUBEPOCH

UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

INFRACLASS: EURYAPSIDA COLBERT, 1945

SUPERORDER: SAUROPTERYGIA OWEN 1860

GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994

ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835

SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835

INFRAORDER: PLESIOSAUROMORPHA NOVA

EUPLESIOSAURIA O’KEEFE, 2001

Superfamily: CRYPTOCLEIDOIDEA Williston, 1925

TRICLEDIA O’KEEFE , 2001

Family: POLYCOTYLIDAE Cope, 1869

CLADE NAME: DOLICHORHYNCHIA CLARK, O'KEEFE & SLACK, 2024

*Martinectes bonneri
= Dolichorhynchops bonneri
= Trinacromerum bonneri

_____________________________________________________________________________________

WYOMING

NIOBRARA COUNTY

8.5 MILES EAT OF REDBIRD

SHARON SPRINGS FORMATION

BACULITES OBTUSUS ZONE

MIDDLE CAMPANIAN STAGE

UPPER SENONIAN SUBEPOCH

UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

INFRACLASS: EURYAPSIDA COLBERT, 1945

SUPERORDER: SAUROPTERYGIA OWEN 1860

GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994

ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835

SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835

INFRAORDER: PLESIOSAUROMORPHA NOVA

EUPLESIOSAURIA O’KEEFE, 2001

Superfamily: CRYPTOCLEIDOIDEA Williston, 1925

TRICLEDIA O’KEEFE , 2001

Family: POLYCOTYLIDAE Cope, 1869

CLADE NAME: DOLICHORHYNCHIA CLARK, O'KEEFE & SLACK, 2024

*Unktaheela spectai

_____________________________________________________________________________________

EASTERN WYOMING

NIOBRARA COUNTY

LANCE FORMATION

LANCIAN

MAASTRICHTIAN STAGE

UPPER SENONIAN SUBEPOCH

UPPER GULF EPOCH

LATEST CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: AVES LINNAEUS, 1758

AVEBREVICAUDA PAUL, 2001

SUBCLASS: CARINATE MERREM, 1813

INFRACLASS: NEORNITHES GADOW, 1893

PARVCLASS: NEOGNATHAE PYCRAFT 1900

GALLOANSERES SIBLEY & AHLQUIST, 1990

Incertae sedis

Gen. sp indet.

_____________________________________________________________________________________

WYOMING

NORTHERN NIOBRARA COUNTY

SOUTHEASTERN POWEDER BASIN

OLD WOMAN ANTICLINE

UPPER MEMBER

PIERRE FORMATION

EARLY MAASTRICHTIAN STAGE

UPPER SENONIAN SUBEPOCH

UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

INFRACLASS: EURYAPSIDA COLBERT, 1945

SUPERORDER: SAUROPTERYGIA OWEN 1860

GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994

ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835

SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835

INFRAORDER: PLESIOSAUROMORPHA NOVA

EUPLESIOSAURIA O’KEEFE, 2001

Superfamily: CRYPTOCLEIDOIDEA Williston, 1925

TRICLEDIA O’KEEFE , 2001

Family: POLYCOTYLIDAE Cope, 1869

*Serpentisuchops pfisterae

_____________________________________________________________________________________

EASTERN WYOMING

NORTHERN NIOBRARA COUNTY

HANSON RANCH (HR)

LANCE FORMATION

LANCIAN

MAASTRICHTIAN STAGE

UPPER SENONIAN SUBEPOCH

UPPER GULF EPOCH

LATEST CRETACEOUS

WYOMING

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: CHELONOMORPHA KUHN, 1960

SUPERCOHORT: TESTUDINES LINNAEUS, 1758

Incertae sedis

Gen. sp indet.

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

Incertae sedis

Gen. sp indet

SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990

COHORT: CROCODYLOTARSI BENTON & CLARK, 1988

MAGNORDER: SUCHIA KREBS, 1974

SUPERORDER: CROCODYLOMORPHA WALKER, 1968

GRANDORDER: CROCODILIFORMES HAY, 1930

MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983

ORDER: EUSUCHIA emend HUXLEY, 1875

INFRAORDER: CROCODYLIA emend GMELIN, 1877

PARVORDER: BREVIROSTRES

Superfamily: ALLIGATORIDEA Buffetaut, 1985

Family: LEIDYOSUCHIDAE Rauhe & Rossmann, 1995

Leidyosuchus sp

GLOBIDONTA BROUCHU, 1999

Family: ALLIGATORIDAE Gray, 1844

Brachychampsa sp

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

Incertae sedis

Richardoestesia sp

Gen. sp indet

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: TETANURAE GAUTHIER, 1986

MICRORDER: COELUROSAURIA HUENE, 1914

ORDER: MANIRAPTORFORMES HOLTZ, 1996

AVETHEROPODA PAUL, 2001

SUBORDER: COELURIA MARSH, 1881

Superfamily: TYRANNOSAUROIDEA Osborn, 1906 (sensu Bonaparte et. al., 1990)

Family: TYRANNOSAURIDAE Osborn, 1906

= Aublysodon sp

Subfamily: TYRANNOSAURINAE Osborn, 1906

Tribe: TYRANNOSAURINI Osborn, 1906 (sensu Olshevsky, Ford & Yamamoto, 1995)

Nanotyrannus sp

Tyrannosaurus rex

SUBORDER: ORNITHOMIMOSAURIA BARSBOLD, 1976

Family: ORNITHOMIMIDAE Marsh, 1890

Struthiomimus sp

SUBORDER: MANIRAPTORA GAUTHIER, 1986

INFRAORDER: PARVES SERENO, 1997

PARVORDER: DEINONYCHOSAURIA COLBERT & D. A. RUSSELL, 1969

Family: DROMAEOSAURIDAE Russell, 1969

Subfamily: DROMAEOSAURINAE Matthew & Brown, 1922

Acheroraptor

Gen. sp indet.

Family: TROODONTIDAE Gilmore, 1924

Troodon sp

COHORT: ORNITHISCHIFORMES COOPER, 1985

MAGNORDER: ORNITHISCHIA SEELEY, 1888

SUPERORDER: NEORNITHISCHIA COOPER, 1985

GRANDORDER: ORNITHOPODA MARSH, 1871

MICRORDER: EUORNITHOPODA SERENO, 1986

ORDER: IGUANODONTIA DOLLO, 1888

Family: THESCELOSAURIDAE Sternberg, 1937

Thescelosaurus sp

SUBORDER: ANKYLOPOLLEXIA SERENO, 1986

INFRAORDER: STYRACOSTERNA SERENO, 1986

PARVORDER: HADROSAURIFORMES SERENO, 1997

SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)

Family: HADROSAURIDAE COPE, 1869

Subfamily: HADROSAURINAE Cope, 1869 (sensu Lambe, 1918)

Tribe: EDMONTOSAURINI Brett-Surman vide Chapman & Brett-Surman, 1990

Anatosaurus sp
= Edmontosaurus sp

MICRORDER: MARGINOCEPHALIA SERENO, 1986

ORDER: PACHYCEPHALOSAURIA MARYANSKA & OSMOLSKA, 1974

SUBORDER: DOMOCEPHALINIA SERENO, 1986

INFRAORDER: PACHYCEPHALOSAURIA (MARYANSKA & OSMOLSKA, 1974) COOPER, 1985

Family: PACHYCEPHALOSAURIDAE Sternberg, 1945

Subfamily: PACHYCEPHALOSAURINAE Sternberg, 1945 (sensu Sereno, 1998)

Pachycephalosaurus sp

ORDER: CERATOPIA MARSH, 1890

SUBORDER: NEOCERATOPIA SERENO, 1986

INFRAORDER: CORONOSAURIA SERENO, 1986

PARVORDER: CERATOPOMORPHA Wolfe & Kirkland, 1998 (emend)

Superfamily: CERATOPOIDEA Marsh, 1888 (sensu Hay, 1902)

Family: CERATOPIDAE Marsh, 1888

Subfamily: CERATOPIINAE Marsh, 1888 (sensu Abel, 1919)

Tribe: TRICERATOPINI nova

Triceratops sp

CLASS: MAMMALIA LINNAEUS, 1758

Incertae sedis

Gen. sp indet.

_____________________________________________________________________________________

NIOBRARA COUNTY

W. OF WARREN

LANCE FORMATION

LANCIAN

MAASTRICHTIAN STAGE

UPPER SENONIAN SUBEPOCH

UPPER GULF EPOCH

LATEST CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: CHELONOMORPHA KUHN, 1960

SUPERCOHORT: TESTUDINES LINNAEUS, 1758

COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988

MAGANORDER: CRYPTODIRA GAFFNEY & MEYLAN, 1988

SUPERORDER: SELMACRYPTODIRA GAFFNEY, HUTCHISON, JENKINS & MEEKR, 1987

GRANDORDER: DAIOCRYPTODIRA GAFFNEY & MEYLAN, 1988

MICRORDER: CENTROCRYPTODIRA GAFFNEY & MEYLAN, 1988

ORDER: POLYCRPTODIRA GAFFNEY & MEYLAN, 1988

SUBORDER: PROCOELOCRYPTODIRA GAFFNEY & MEYLAN, 1988

INFRAORDER: CHELOMACRYPTODIRA GAFFNEY & MEYLAN, 1988

PARVORDER: TRIONYCHIDA GRAY, 1870

Superfamily: TRIONYCHIDEA Bell, 1828 (sensu Gray, 1870)

Family: TRIONYCHIDAE Bell, 1828

Axestemys infernalis

_____________________________________________________________________________________

WYOMING

NIOBARA COUNTY

ZERBST RANCH

LANCE FORMATION

MAASTRICHTIAN STAGE

UPPER SENONIAN SUBEPOCH

UPPER GULF EPOCH

LATE CRETACEOUS

ICHNOSUPERCLASS: AMNIOTAIPIDII NOVA

ICHNOCLASS: REPTILIPEDIA VIALOV, 1966

ICHNOSUBCLASS: DIAPSIDIPIDII NOVA

ICHNOSTEM-GROUP: NEODIAPSIDIPEDII NOVA

ICHNOINFRACLASS: NEODIAPSIDIPEDII NOVA

ICHNOSUBDIVISION: ARCHOSAURIPEDIDII NOVA

ICHNOINFRADIVISION: CROCODYLOTARSIPEDIDII NOVA

ICHNOSUPERORDER: CROCODYLOMORPHIPEDIDA WALKER, 1968

Incertae sedis

Gen. sp indet.

_____________________________________________________________________________________

WYOMING

WYOMING

NIOBRARA COUNTY

MULE CREEK JUNCTION

SHARON SPRINGS MEMBER

LOWER PIERRE SHALE FORMATION

LOWERMOST MIDDLE CAMPANIAN STAGE

SENONIAN SUBEPOCH

UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: LACERTILIA OWEN, 1842

GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961

MICRORDER: SCINCOMORPHA CAMP, 1923

ORDER: ANGUIMORPHA FURBRINGER, 1900

SUBORDER: DIPLOGLOSSA COPE, 1864

INFRAORDER: THECAGLOSSA COPE, 1890

PARVORDER: MONSTERSAURIA NORELL & GAO, 1977

Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)

Family: MOSASAURIDAE Gervais, 1853

Infrafamily: NATANIA

Subfamily: RUSSELLOSAURINAE Bell, 1993

Tribe: PLIOPLATECARPINI Dollo, 1884 (sensu Russell, 1967)

Latoplatecarpus willistoni

 

_____________________________________________________________________________________

WYOMING

NIOBRARA COUNTY

"TWENTY-THREE MILES (23 MILES (37 KM) W OF EDGEMONT AND 1 MILE (1.6 KM) N OF RED BIRD

SHARON SPRINGS MEMBER

LOWER PIERRE SHALE FORMATION

B. OBTUSUS AND B. MCLEARNI AMMONITE ZONES

LOWERMOST MIDDLE CAMPANIAN STAGE

SENONIAN SUBEPOCH

UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1903

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: LACERTILIA OWEN, 1842

GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961

MICRORDER: SCINCOMORPHA CAMP, 1923

ORDER: ANGUIMORPHA FURBRINGER, 1900

SUBORDER: DIPLOGLOSSA COPE, 1864

INFRAORDER: THECAGLOSSA COPE, 1890

PARVORDER: MONSTERSAURIA NORELL & GAO, 1977

Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)

Family: MOSASAURIDAE Gervais, 1853

Infrafamily: NATANIA

Subfamily: RUSSELLOSAURINAE Bell, 1993

Tribe: PLIOPLATECARPINI Dollo, 1884 (sensu Russell, 1967)

Latoplatecarpus willistoni

 

_____________________________________________________________________________________

WYOMING

NIOBARA COUNTY

WHITNEY RANCH

LANCE FORMATION

LANCIAN AGE

UPPER MAASTRICHTIAN STAGE

UPPERMOST SENONIAN SUBEPOCH

UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: TETANURAE GAUTHIER, 1986

MICRORDER: COELUROSAURIA HUENE, 1914

ORDER: MANIRAPTORFORMES HOLTZ, 1996

AVETHEROPODA PAUL, 2001

SUBORDER: COELURIA MARSH, 1881

Superfamily: TYRANNOSAUROIDEA Osborn, 1906 (sensu Bonaparte et. al., 1990)

Family: TYRANNOSAURIDAE Osborn, 1906

Subfamily: TYRANNOSAURINAE Osborn, 1906

Tribe: TYRANNOSAURINI Osborn, 1906 (sensu Olshevsky, Ford & Yamamoto, 1995)

Tyrannosaurus rex

_____________________________________________________________________________________

WYOMING

NIOBARA COUNTY

37 KM NORTH OF LUSK

BUCK CREEK

LANCE FORMATION

LANCIAN AGE

UPPER MAASTRICHTIAN STAGE

UPPERMOST SENONIAN SUBEPOCH

UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: TETANURAE GAUTHIER, 1986

MICRORDER: COELUROSAURIA HUENE, 1914

ORDER: MANIRAPTORFORMES HOLTZ, 1996

AVETHEROPODA PAUL, 2001

SUBORDER: COELURIA MARSH, 1881

Superfamily: TYRANNOSAUROIDEA Osborn, 1906 (sensu Bonaparte et. al., 1990)

Family: TYRANNOSAURIDAE Osborn, 1906

Subfamily: TYRANNOSAURINAE Osborn, 1906

Tribe: TYRANNOSAURINI Osborn, 1906 (sensu Olshevsky, Ford & Yamamoto, 1995)

Tyrannosaurus rex

cf. Tyrannosaurus

_____________________________________________________________________________________

NIOBRARA COUNTY

LUSK

LANCE FORMATION

LANCIAN AGE

UPPER MAASTRICHTIAN STAGE

UPPERMOST SENONIAN SUBEPOCH

UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: ORNITHISCHIFORMES COOPER, 1985

MAGNORDER: ORNITHISCHIA SEELEY, 1888

SUPERORDER: NEORNITHISCHIA COOPER, 1985

GRANDORDER: ORNITHOPODA MARSH, 1871

MICRORDER: EUORNITHOPODA SERENO, 1986

ORDER: IGUANODONTIA DOLLO, 1888

Family: THESCELOSAURIDAE Sternberg, 1937

Thescelosaurus infernalis
= Bugenasaura infernalis

_____________________________________________________________________________________

WYOMING

NORTHERN NIOBRARA COUNTY

4 MILES WEST OF KUHMN 40001

JOHNSON RANCH

SHARON SPRINGS MEMBER (WAS REFERRED TO THE REDBIRD MEMBER)

PIERRE FORMATION

CAMPANIAN STAGE

UPPER SENONIAN SUBEPOCH

UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

INFRACLASS: EURYAPSIDA COLBERT, 1945

SUPERORDER: SAUROPTERYGIA OWEN 1860

GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994

ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835

SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835

INFRAORDER: PLESIOSAUROMORPHA NOVA

EUPLESIOSAURIA O’KEEFE, 2001

Superfamily: CRYPTOCLEIDOIDEA Williston, 1925

TRICLEDIA O’KEEFE , 2001

Family: POLYCOTYLIDAE Cope, 1869

*Dolichorhynchops bonneri
= Trinacromerum bonneri

_____________________________________________________________________________________

WYOMING

NIOBRARA COUNTY

NEAR THE SOUTH DAKOTA BORDER

MULE CREEK

SHARON SPRINGS FORMATION

LOWER PIERRE SHALE GROUP

LOWER CAMPANIAN STAGE

SENONIAN SUBEPOCH

GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUPERCOHORT: AVEMETATARSALIA BENTON, 1999

COHORT: PROLACERTIFORMES CAMP, 1945 NOVA

SUPERORDER: TAPINOPLATIA PETERS, 2000

GRANDORDER: CHARACIOPODA PETERS, 2000

MICRORDER: FENESTRASAURIA PETERS, 2000

ORDER: PTEROSAURIA KAUP, 1834

SUBORDER: EUPTERODACTYLOIDEA BENNETT, 1994

Superfamily: PTERANODONTIDEA Marsh, 1876

Family: PTERANODONTIDAE Marsh, 1876

Pteranodon longiceps

_____________________________________________________________________________________

WYOMING

NIOBRARA COUNTY

GREASEWOOD CREEK LOCALITY

SDSM LOCALITY V685

LANCE FORMATION

LANCIAN AGE

UPPER MAASTRICHTIAN STAGE

UPPERMOST SENONIAN SUBEPOCH

UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: MAMMALIA LINNAEUS, 1758

SUBCLASS: PROTOTHERIA GILL, 1872

INFRACLASS: TRECHNOTHERIA McKENNA, 1975

LEGION: CLADOTHERIA McKENNA, 1975

SUBLEGION: ZATHERIA MCKENNA, 1975

INFRALEGION: TRIBOSPHENIDA (McKENNA, 1975) McKENNA & BELL, 1997

SUPERCOHORT: THERIA (PARKER & HASWELL, 1987) McKENNA & BELL, 1997

COHORT: MARSUPIALIA (ILLIGER, 1811) McKENNA & BELL, 1997

ORDER: STAGODONTIA ARCHER, 1984

Family: PEDIOMYIDAE Simpson, 1927

Leptalestes krejcii
= Pediomys sp. cf. P. krejcii

_____________________________________________________________________________________ WYOMING

NIOBRARA COUNTY

ICHNOSUPERCLASS: AMNIOTAIPIDII NOVA

ICHNOCLASS: REPTILIPEDIA VIALOV, 1966

ICHNOSUBCLASS: DIAPSIDIPIDII NOVA

ICHNOSTEM-GROUP: NEODIAPSIDIPEDII NOVA

ICHNOINFRACLASS: NEODIAPSIDIPEDII NOVA

ICHNOSUBDIVISION: ARCHOSAURIPEDIDII NOVA

ICHNOLEGION: DINOSAUROMORPHIPEDA NOVA

ICHNOSUBLEGION: DINOSAURIFORMEIPEDIDII NOVA

ICHNOSUPERCOHORT: DINOSAUROPEDIDA NOVA

ICHNOCOHORT: THEROPODIPEDIA NOVA

Incerate sedis

Saurexallopus sp

ICHNOSUPERORDER: NEOTHEROPODOPEDIDA NOVA

ICHNOORDER: NEOTETANURIPEDIDII NOVA

ICHNOSUBORDER: AVETHEROPODIPIDA NOVA

ICHNOMICRORDER: COELUROSAURIPIDIDA NOVA

ICHNOORDER: MANIRAPTORFORMIPIDA NOVA

ICHNOSUBORDER: COELURIPIDA NOVA

Ichnosuperfamily: TYRANNOSAUROIPIDEA Nova

Ichnofamily: TYRANNOSAURIPODIDAE emend Vylov, 1988

Tyrannosauripus sp

ICHNOSUBORDER: ORNITHOMIMOSAUROPIDA NOVA

Family: ORNITHIMIMIOPIDIDAE NOVA

Gen. sp indet.

ICHNOCOHORT: ORNITHISCHIFORMEIPIDA NOVA

ICHNOMAGNORDER: ORNITHISCHIPIDA NOVA

ICHNOSUPERORDER: NEORNITHISCHIPIDA NOVA

ICHNOGRANDORDER: ORNITHOPODIPDIA NOVA

ICHNOMICRORDER: EUORNITHOPODIPIDA NOVA

ICHNOORDER: IGUANODONTIPIDIA NOVA

ICHNOSUBORDER: ANKYLOPOLLEXIPEDIDA NOVA

ICHNOINFRAORDER: STYRACOSTERNIPIDA NOVA

ICHNOPARVORDER: HADROSAURIFORMIPEDIDA NOVA

ICHNOSUPERFAMILY: HADROSAUROIDIPEDIA NOVA

Ichnofamily: HADROSAURIOPDIDAE Vylov, 1988

Gen. sp indet.

 

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: MAMMALIA LINNAEUS, 1758

SUBCLASS: PROTOTHERIA GILL, 1872

INFRACLASS: TRECHNOTHERIA McKENNA, 1975

LEGION: CLADOTHERIA McKENNA, 1975

SUBLEGION: ZATHERIA MCKENNA, 1975

INFRALEGION: TRIBOSPHENIDA (McKENNA, 1975) McKENNA & BELL, 1997

SUPERCOHORT: THERIA (PARKER & HASWELL, 1987) McKENNA & BELL, 1997

COHORT: MARSUPIALIA (ILLIGER, 1811) McKENNA & BELL, 1997

MAGNORDER: AMERIDELPHIA (SZALAY, in ARCHER, ED, 1982) McKENNA & BELL, 1997

ORDER: PAUCITUBERCULATA AMEGHINO, 1894

Superfamily: CAROLOAMEGHINOIDEA Ameghino, 1901

Family: GLASBIIDAE Clemens, 1966

*Hatcheritherium alpha

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: AVES LINNAEUS, 1758

AVEBREVICAUDA PAUL, 2001

SUBCLASS: CARINATE MERREM, 1813

INFRACLASS: NEORNITHES GADOW, 1893

PARVCLASS: NEOGNATHAE PYCRAFT 1900

ORDER: ANSERIFORMES WAGLER, 1831

Family: PRESBYORNITHIDAE Wetmore, 1926

Incertae sedis

Undetermined species 3

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

INFRACLASS: EURYAPSIDA COLBERT, 1945

SUPERORDER: SAUROPTERYGIA OWEN 1860

GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994

ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835

SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835

INFRAORDER: PLESIOSAUROMORPHA NOVA

EUPLESIOSAURIA O’KEEFE, 2001

Superfamily: ELASMOSAUROIDEA Cope, 1869

Family: ELASMOSAURIDAE Cope, 1869

Hydralmosaurus serpentinus
= *Styxosaurus browni

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: ORNITHISCHIFORMES COOPER, 1985

MAGNORDER: ORNITHISCHIA SEELEY, 1888

SUPERORDER: NEORNITHISCHIA COOPER, 1985

GRANDORDER: ORNITHOPODA MARSH, 1871

MICRORDER: MARGINOCEPHALIA SERENO, 1986

ORDER: CERATOPIA MARSH, 1890

SUBORDER: NEOCERATOPIA SERENO, 1986

INFRAORDER: CORONOSAURIA SERENO, 1986

PARVORDER: CERATOPOMORPHA Wolfe & Kirkland, 1998 (emend)

Superfamily: CERATOPOIDEA Marsh, 1888 (sensu Hay, 1902)

Family: CERATOPIDAE Marsh, 1888

Subfamily: CERATOPIINAE Marsh, 1888 (sensu Abel, 1919)

Tribe: TRICERATOPINI nova

Triceratops horridus
= *Triceratops sternbergii

NIOBRARA COUNTY

LANCE FORMATION

LANCIAN AGE

UPPER MAASTRICHTIAN STAGE

UPPERMOST SENONIAN SUBEPOCH

UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: AMPHIBIA LINNAEUS, 1758

SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?

ORDER: ALLOCAUDATA FOX & NAYLOR, 1982

Family: ALBANERPETONTIDAE Fox & Naylor, 1982

Albanerpeton nexuosus

SUPERORDER: BATRACHIA BRONGNIART, 1800

ORDER: CAUDATA SCOPOLI, 1777

SUBORDER: PORTEOIDEA (NOBEL 1931) EMEND EDWARDS, 1976

Family: BATRACHOSAUROIDIDAE Auffenberg, 1958

*Opisthotriton kayi

SUBORDER: AMBYSTOMATOIDEA NOBLE, 1931

Family: SCAPHERPETONTIDAE Auffenberg & Goin, 1959

Scapherpeton tectum
Scapherpeton sp

SUBORDER: SALAMANDROIDEA NOBLE, 1931

Family: SIRENIDAE Gray, 1825

*Habrosaurus dilatus
= *Adelphesiren olivae

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: CHELONOMORPHA KUHN, 1960

SUPERCOHORT: TESTUDINES LINNAEUS, 1758

COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988

MAGANORDER: CRYPTODIRA GAFFNEY & MEYLAN, 1988

SUPERORDER: SELMACRYPTODIRA GAFFNEY, HUTCHISON, JENKINS & MEEKR, 1987

GRANDORDER: DAIOCRYPTODIRA GAFFNEY & MEYLAN, 1988

MICRORDER: BAENOIDEA WILLIAMS, 1960

Family: BAENIDAE Cope, 1882

Subfamily: EUBAENINAE Gaffney, 1972

Eubaena cephalica
= Baena cephalica

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: TETANURAE GAUTHIER, 1986

MICRORDER: COELUROSAURIA HUENE, 1914

ORDER: MANIRAPTORFORMES HOLTZ, 1996

AVETHEROPODA PAUL, 2001

SUBORDER: COELURIA MARSH, 1881

Superfamily: TYRANNOSAUROIDEA Osborn, 1906 (sensu Bonaparte et. al., 1990)

Family: TYRANNOSAURIDAE Osborn, 1906

Subfamily: TYRANNOSAURINAE Osborn, 1906

Tribe: TYRANNOSAURINI Osborn, 1906 (sensu Olshevsky, Ford & Yamamoto, 1995)

Tyrannosaurus rex
= *Aublysodon cristatus
= Deinodon cristatus
= Stygivenator cristatus

COHORT: ORNITHISCHIFORMES COOPER, 1985

MAGNORDER: ORNITHISCHIA SEELEY, 1888

SUPERORDER: NEORNITHISCHIA COOPER, 1985

GRANDORDER: ORNITHOPODA MARSH, 1871

MICRORDER: EUORNITHOPODA SERENO, 1986

ORDER: IGUANODONTIA DOLLO, 1888

SUBORDER: ANKYLOPOLLEXIA SERENO, 1986

INFRAORDER: STYRACOSTERNA SERENO, 1986

PARVORDER: HADROSAURIFORMES SERENO, 1997

SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)

Family: HADROSAURIDAE COPE, 1869

Subfamily: HADROSAURINAE Cope, 1869 (sensu Lambe, 1918)

Tribe: EDMONTOSAURINI Brett-Surman vide Chapman & Brett-Surman, 1990

*Anatosaurus longiceps
= Trachodon longiceps
= Edmontosaurus longiceps
= Hadrosaurus longiceps
= Anatotian longiceps
Anatosaurus annectens

MICRORDER: MARGINOCEPHALIA SERENO, 1986

ORDER: PACHYCEPHALOSAURIA MARYANSKA & OSMOLSKA, 1974

SUBORDER: DOMOCEPHALINIA SERENO, 1986

INFRAORDER: PACHYCEPHALOSAURIA (MARYANSKA & OSMOLSKA, 1974) COOPER, 1985

Family: PACHYCEPHALOSAURIDAE Sternberg, 1945

Subfamily: PACHYCEPHALOSAURINAE Sternberg, 1945 (sensu Sereno, 1998)

Pachycephalosaurus wyomingensis
= Troodon wyomingensis
= Stegoceras wyomingense
= Stegoceras wyomingensis

ORDER: CERATOPIA MARSH, 1890

SUBORDER: NEOCERATOPIA SERENO, 1986

INFRAORDER: CORONOSAURIA SERENO, 1986

PARVORDER: CERATOPOMORPHA Wolfe & Kirkland, 1998 (emend)

Superfamily: CERATOPOIDEA Marsh, 1888 (sensu Hay, 1902)

Family: CERATOPIDAE Marsh, 1888

Subfamily: CERATOPIINAE Marsh, 1888 (sensu Abel, 1919)

Tribe: TRICERATOPINI nova

Triceratops horridus
= Triceratops elatus
Triceratops prorsus
Triceratops sp

CLASS: MAMMALIA LINNAEUS, 1758

SUBCLASS: THERIIFORMES (ROWE, 1988) McKENNA & BELL, 1997

INFRACLASS: ALLOTHERIA (MARSH, 1880) McKENNA & BELL, 1997

ORDER: MULTITUBERCULATA COPE, 1884

SUBORDER: CIMOLODONTA McKENNA, 1975

Superfamily: TAENIOLABIDOIDEA Granger & Simpson, 1929 (sensu McKenna & Bell, 1997)

Family: TAENIOLABIDIDAE Granger & Simpson, 1929

Meniscoessus sp

SUBCLASS: PROTOTHERIA GILL, 1872

INFRACLASS: TRECHNOTHERIA McKENNA, 1975

LEGION: CLADOTHERIA McKENNA, 1975

SUBLEGION: DRYOLESTOIDEA BUTLER, 1939

ORDER: DRYOLESTOIDA PROTHERO, 1981

Family: DRYOLESTIDAE Marsh, 1879

*Dryolestes tenax

SUBLEGION: ZATHERIA MCKENNA, 1975

INFRALEGION: TRIBOSPHENIDA (McKENNA, 1975) McKENNA & BELL, 1997

SUPERCOHORT: THERIA (PARKER & HASWELL, 1987) McKENNA & BELL, 1997

COHORT: MARSUPIALIA (ILLIGER, 1811) McKENNA & BELL, 1997

ORDER: STAGODONTIA ARCHER, 1984

Family: STAGODONTIDAE Marsh, 1889

Didelphodon vorax
= *Stagodon validus
= *Stagodon nitor
= *Cimolestes curtus
= Diaphorodon curtus

Eodelphis browni
= Didelphodon petersoni
= Ectoconodon petersoni

Family: PEDIOMYIDAE Simpson, 1927

Pediomys elegans

Gen. sp indet.

MAGNORDER: AMERIDELPHIA (SZALAY, in ARCHER, ED, 1982) McKENNA & BELL, 1997

ORDER: DIDELPHIMORPHIA GILL, 1872

Family: DIDELPHIDAE Gray 1821

Subfamily: ALPHADONTIDAE Marshall, Case & Woodburne, 1990

*Alphadon marshi

COHORT: PLACENTALIA (OWEN, 1837) McKENNA & BELL, 1997

MAGNORDER: EPITHERIA (McKENNA, 1975) McKENNA & BELL, 1997

SUPERORDER: PREPTOTHERIA McKENNA, 1975

GRANDORDER: UNGULATOMORPHA ARCHIBALD, 1996

Family: GYPSONICTIPIDAE Van Valen, 1967

*Gypsonictops hypoconus
= Euangelestes petersoni

NIOBRARA COUNTY (OLD CONVERSE COUNTY)

LANCE FORMATION

LANCIAN AGE

UPPER MAASTRICHTIAN STAGE

UPPERMOST SENONIAN SUBEPOCH

UPPER UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: CHELONOMORPHA KUHN, 1960

SUPERCOHORT: TESTUDINES LINNAEUS, 1758

COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988

MAGANORDER: CRYPTODIRA GAFFNEY & MEYLAN, 1988

SUPERORDER: SELMACRYPTODIRA GAFFNEY, HUTCHISON, JENKINS & MEEKR, 1987

GRANDORDER: DAIOCRYPTODIRA GAFFNEY & MEYLAN, 1988

MICRORDER: BAENOIDEA WILLIAMS, 1960

Family: BAENIDAE Cope, 1882

Subfamily: EUBAENINAE Gaffney, 1972

*Eubaena cephalica
= Baena cephalica

WYOMING

NIOBRARA COUNTY

LANCE FORMATION

LANCIAN

MAASTRICHTIAN STAGE

UPPER SENONIAN SUBEPOCH

UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: CHELONOMORPHA KUHN, 1960

SUPERCOHORT: TESTUDINES LINNAEUS, 1758

COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988

MAGANORDER: CRYPTODIRA GAFFNEY & MEYLAN, 1988

SUPERORDER: SELMACRYPTODIRA GAFFNEY, HUTCHISON, JENKINS & MEEKR, 1987

GRANDORDER: DAIOCRYPTODIRA GAFFNEY & MEYLAN, 1988

MICRORDER: CENTROCRYPTODIRA GAFFNEY & MEYLAN, 1988

ORDER: POLYCRPTODIRA GAFFNEY & MEYLAN, 1988

SUBORDER: PROCOELOCRYPTODIRA GAFFNEY & MEYLAN, 1988

INFRAORDER: CHELOMACRYPTODIRA GAFFNEY & MEYLAN, 1988

PARVORDER: TRIONYCHIDA GRAY, 1870

Superfamily: TRIONYCHIDEA Bell, 1828 (sensu Gray, 1870)

Family: TRIONYCHIDAE Bell, 1828

*Gilmoremys lancensis = Aspiderets lancensis

_____________________________________________________________________________________

 WYOMING

NIOBRARA COUNTY

LATE LANCE FORMATION

LANCIAN AGE

UPPER MAASTRICHTIAN STAGE

UPPERMOST SENONIAN SUBEPOCH

UPPER UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: ORNITHISCHIFORMES COOPER, 1985

MAGNORDER: ORNITHISCHIA SEELEY, 1888

SUPERORDER: NEORNITHISCHIA COOPER, 1985

GRANDORDER: ORNITHOPODA MARSH, 1871

MICRORDER: MARGINOCEPHALIA SERENO, 1986

ORDER: CERATOPIA MARSH, 1890

SUBORDER: NEOCERATOPIA SERENO, 1986

INFRAORDER: CORONOSAURIA SERENO, 1986

PARVORDER: CERATOPOMORPHA Wolfe & Kirkland, 1998 (emend)

Superfamily: CERATOPOIDEA Marsh, 1888 (sensu Hay, 1902)

Family: CERATOPIDAE Marsh, 1888

Subfamily: CERATOPIINAE Marsh, 1888 (sensu Abel, 1919)

Tribe: TRICERATOPINI nova

Triceratops horridus
= *Triceratops sulcatus

_____________________________________________________________________________________

NIOBRARA COUNTY

LANCE CREEK

LANCE FORMATION

LANCIAN AGE

UPPER MAASTRICHTIAN STAGE

UPPERMOST SENONIAN SUBEPOCH

UPPER UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: CHELONOMORPHA KUHN, 1960

SUPERCOHORT: TESTUDINES LINNAEUS, 1758

COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988

MAGANORDER: CRYPTODIRA GAFFNEY & MEYLAN, 1988

SUPERORDER: SELMACRYPTODIRA GAFFNEY, HUTCHISON, JENKINS & MEEKR, 1987

GRANDORDER: DAIOCRYPTODIRA GAFFNEY & MEYLAN, 1988

MICRORDER: BAENOIDEA WILLIAMS, 1960

Family: NEURANKYLIDAE Lambe, 1902

Compsemys victa

Family: BAENIDAE Cope, 1882

Incertae sedis

non Baena, *B. hayi

SUBCLASS: DIAPSIDA OSBORN, 1904

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA DUMERIL & BIBRON, 1839

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA OPPEL, 1811

SUPERORDER: LACERTILIA OWEN, 1842

GRANDORDER: ASCALABOTA GILMORE, 1928

ORDER: IGUANIA LATREILLE, 1825

SUBORDER: DRACOMORPHA NORTHCUTT, 1978

INFRAORDER: TEIUMORPHA ALIFANOV, 2000

Family: TEIIDAE Gray, 1827

Subfamily: CHAMOPSINAE Denton & O’Neill 1995

*Chamops segnis
= *Iguanius teres
= *Lanceosaurus hatcheri
= *Lanceosaurus compressus

GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961

MICRORDER: SCINCOMORPHA CAMP, 1923

ORDER: ANGUIMORPHA FURBRINGER, 1900

SUBORDER: DIPLOGLOSSA COPE, 1864

INFRAORDER: XENOSAUIRA FURBRINGER, 1900

Family: XENOSAURIDAE Cope, 1886

Exostinus lancensis
= *Prionosaurus regularis

INFRAORDER: OPHISAUROMORPHA ALIFANOV, 2000

Family: ANGUIDAE Gray, 1825

Subfamily: GLYPTOSAURINAE Marsh, 1872

Tribe: MELANOSAURINI Sullivan, 1979

Odaxosaurus piger
= Odaxosaurus obliquus

INFRAORDER: THECAGLOSSA COPE, 1890

PARVORDER: MONSTERSAURIA NORELL & GAO, 1977

Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)

Family: VARANIDAE Gray, 1827

Palaeosaniwa canadensis

Family: PARASANIWIDAE Estes, 1964

*Parasaniwa wyomingensis
*Parasaniwa obtusa

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: TETANURAE GAUTHIER, 1986

MICRORDER: COELUROSAURIA HUENE, 1914

ORDER: MANIRAPTORFORMES HOLTZ, 1996

AVETHEROPODA PAUL, 2001

SUBORDER: COELURIA MARSH, 1881

Superfamily: TYRANNOSAUROIDEA Osborn, 1906 (sensu Bonaparte et. al., 1990)

Family: TYRANNOSAURIDAE Osborn, 1906

Subfamily: TYRANNOSAURINAE Osborn, 1906

Tribe: TYRANNOSAURINI Osborn, 1906 (sensu Olshevsky, Ford & Yamamoto, 1995)

Tyrannosaurus rex

COHORT: ORNITHISCHIFORMES COOPER, 1985

MAGNORDER: ORNITHISCHIA SEELEY, 1888

SUPERORDER: NEORNITHISCHIA COOPER, 1985

GRANDORDER: ORNITHOPODA MARSH, 1871

MICRORDER: EUORNITHOPODA SERENO, 1986

ORDER: IGUANODONTIA DOLLO, 1888

Family: THESCELOSAURIDAE Sternberg, 1937

Thescelosaurus neglectus

SUBORDER: ANKYLOPOLLEXIA SERENO, 1986

INFRAORDER: STYRACOSTERNA SERENO, 1986

PARVORDER: HADROSAURIFORMES SERENO, 1997

SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)

Family: HADROSAURIDAE COPE, 1869

Incertae sedis

Gen. sp indet.

Subfamily: HADROSAURINAE Cope, 1869 (sensu Lambe, 1918)

Tribe: EDMONTOSAURINI Brett-Surman vide Chapman & Brett-Surman, 1990

Edmontosaurus sp

SUPERORDER: EURYOPDA SERENO, 1986

GRANDORDER: ARMATOSAURIA ZHAO, 1983 (emend)

ORDER: ANKYLOSAURIA OSBORN, 1923

SUBORDER: SCELIDOSAURIA COOPER, 1985

INFRAORDER: ANKYLOSAURA OSBORN, 1923

Family: ANKYLOSAURIDAE Brown, 1908

Incertae sedis

Gen. sp indet.

 

NIOBRARA COUNTY

LANCE CREEK

QUARRY 1

LANCE FORMATION

LANCIAN AGE

UPPER MAASTRICHTIAN STAGE

UPPERMOST SENONIAN SUBEPOCH

UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA DUMERIL & BIBRON, 1839

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA OPPEL, 1811

SUPERORDER: LACERTILIA OWEN, 1842

GRANDORDER: ASCALABOTA GILMORE, 1928

ORDER: IGUANIA LATREILLE, 1825

SUBORDER: DRACOMORPHA NORTHCUTT, 1978

INFRAORDER: TEIUMORPHA ALIFANOV, 2000

Family: TEIIDAE Gray, 1827

Subfamily: CHAMOPSINAE Denton & O’Neill 1995

Chamops segnis
= *Alethesaurus quadratus

_____________________________________________________________________________________

NIOBRARA COUNTY

BUCK CREEK

LANCE FORMATION

LANCIAN AGE

UPPER MAASTRICHTIAN STAGE

UPPERMOST SENONIAN SUBEPOCH

UPPER UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: ORNITHISCHIFORMES COOPER, 1985

MAGNORDER: ORNITHISCHIA SEELEY, 1888

SUPERORDER: NEORNITHISCHIA COOPER, 1985

GRANDORDER: ORNITHOPODA MARSH, 1871

MICRORDER: MARGINOCEPHALIA SERENO, 1986

ORDER: CERATOPIA MARSH, 1890

SUBORDER: NEOCERATOPIA SERENO, 1986

INFRAORDER: CORONOSAURIA SERENO, 1986

PARVORDER: CERATOPOMORPHA Wolfe & Kirkland, 1998 (emend)

Superfamily: CERATOPOIDEA Marsh, 1888 (sensu Hay, 1902)

Family: CERATOPIDAE Marsh, 1888

Subfamily: CERATOPIINAE Marsh, 1888 (sensu Abel, 1919)

Tribe: TRICERATOPINI nova

Triceratops horridus
= Triceratops sulcatus
= Triceratops elatus
Triceratops prorsus
Triceratops cf. albertensis

_____________________________________________________________________________________

NIOBRARA COUNTY

ABOUT 3 M ABOVE THE MOUTH OF LIGNTHING CREEK

LANCE FORMATION

LANCIAN AGE

UPPER MAASTRICHTIAN STAGE

UPPERMOST SENONIAN SUBEPOCH

UPPER UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: ORNITHISCHIFORMES COOPER, 1985

MAGNORDER: ORNITHISCHIA SEELEY, 1888

SUPERORDER: NEORNITHISCHIA COOPER, 1985

GRANDORDER: ORNITHOPODA MARSH, 1871

MICRORDER: MARGINOCEPHALIA SERENO, 1986

ORDER: CERATOPIA MARSH, 1890

SUBORDER: NEOCERATOPIA SERENO, 1986

INFRAORDER: CORONOSAURIA SERENO, 1986

PARVORDER: CERATOPOMORPHA Wolfe & Kirkland, 1998 (emend)

Superfamily: CERATOPOIDEA Marsh, 1888 (sensu Hay, 1902)

Family: CERATOPIDAE Marsh, 1888

Subfamily: CERATOPIINAE Marsh, 1888 (sensu Abel, 1919)

Tribe: TRICERATOPINI nova

Triceratops prorsus
= *Triceratops brevicornus

_____________________________________________________________________________________

NIOBRARA COUNTY

LIGHTNING CREEK

NEAR THE SUMMIT OF THE BLUFF ON THE NORTH SIDE OF LIGHTNING CREEK

LANCE FORMATION

LANCIAN AGE

UPPER MAASTRICHTIAN STAGE

UPPERMOST SENONIAN SUBEPOCH

UPPER UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: ORNITHISCHIFORMES COOPER, 1985

MAGNORDER: ORNITHISCHIA SEELEY, 1888

SUPERORDER: NEORNITHISCHIA COOPER, 1985

GRANDORDER: ORNITHOPODA MARSH, 1871

MICRORDER: MARGINOCEPHALIA SERENO, 1986

ORDER: CERATOPIA MARSH, 1890

SUBORDER: NEOCERATOPIA SERENO, 1986

INFRAORDER: CORONOSAURIA SERENO, 1986

PARVORDER: CERATOPOMORPHA Wolfe & Kirkland, 1998 (emend)

Superfamily: CERATOPOIDEA Marsh, 1888 (sensu Hay, 1902)

Family: CERATOPIDAE Marsh, 1888

Subfamily: CERATOPIINAE Marsh, 1888 (sensu Abel, 1919)

Tribe: TOROSAURINI Nopcsa, 1915

*Torosaurus latus

_____________________________________________________________________________________

NIOBRARA COUNTY

LIGHTNING CREEK

LANCE FORMATION

LANCIAN AGE

UPPER MAASTRICHTIAN STAGE

UPPERMOST SENONIAN SUBEPOCH

UPPER UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: ORNITHISCHIFORMES COOPER, 1985

MAGNORDER: ORNITHISCHIA SEELEY, 1888

SUPERORDER: NEORNITHISCHIA COOPER, 1985

GRANDORDER: ORNITHOPODA MARSH, 1871

MICRORDER: EUORNITHOPODA SERENO, 1986

ORDER: IGUANODONTIA DOLLO, 1888

SUBORDER: ANKYLOPOLLEXIA SERENO, 1986

INFRAORDER: STYRACOSTERNA SERENO, 1986

PARVORDER: HADROSAURIFORMES SERENO, 1997

SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)

Family: HADROSAURIDAE COPE, 1869

Subfamily: HADROSAURINAE Cope, 1869 (sensu Lambe, 1918)

Tribe: EDMONTOSAURINI Brett-Surman vide Chapman & Brett-Surman, 1990

*Anatosaurus annectens
= Claosaurus annectens
= Anatosaurus annectens
= Thespesius annectens
= Trachodon annectens
= Claosaruus annectans

MICRORDER: MARGINOCEPHALIA SERENO, 1986

ORDER: CERATOPIA MARSH, 1890

SUBORDER: NEOCERATOPIA SERENO, 1986

INFRAORDER: CORONOSAURIA SERENO, 1986

PARVORDER: CERATOPOMORPHA Wolfe & Kirkland, 1998 (emend)

Superfamily: CERATOPOIDEA Marsh, 1888 (sensu Hay, 1902)

Family: CERATOPIDAE Marsh, 1888

Subfamily: CERATOPIINAE Marsh, 1888 (sensu Abel, 1919)

Tribe: TRICERATOPINI nova

Triceratops horridus
= *Triceratops serratus
*Triceratops prorsus
= Triceratops brevicornus

_____________________________________________________________________________________

WYOMING

NIOBRARA COUNTY

NEAR THE TYPE LOCALIYT MIDWAY BETWEEN LIGNTHING AND COW CREEKS

LANCE FORMATION

LANCIAN AGE

UPPER MAASTRICHTIAN STAGE

UPPERMOST SENONIAN SUBEPOCH

UPPER UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: ORNITHISCHIFORMES COOPER, 1985

MAGNORDER: ORNITHISCHIA SEELEY, 1888

SUPERORDER: NEORNITHISCHIA COOPER, 1985

GRANDORDER: ORNITHOPODA MARSH, 1871

MICRORDER: MARGINOCEPHALIA SERENO, 1986

ORDER: CERATOPIA MARSH, 1890

SUBORDER: NEOCERATOPIA SERENO, 1986

INFRAORDER: CORONOSAURIA SERENO, 1986

PARVORDER: CERATOPOMORPHA Wolfe & Kirkland, 1998 (emend)

Superfamily: CERATOPOIDEA Marsh, 1888 (sensu Hay, 1902)

Family: CERATOPIDAE Marsh, 1888

Subfamily: CERATOPIINAE Marsh, 1888 (sensu Abel, 1919)

Tribe: TOROSAURINI Nopcsa, 1915

Torosaurus latus
= Torosaurus gladius

_____________________________________________________________________________________

NIOBRARA COUNTY

25 M ABOVE THE MOUTH OF LANCE CREEK

LANCE FORMATION

LANCIAN AGE

UPPER MAASTRICHTIAN STAGE

UPPERMOST SENONIAN SUBEPOCH

UPPER UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: ORNITHISCHIFORMES COOPER, 1985

MAGNORDER: ORNITHISCHIA SEELEY, 1888

SUPERORDER: NEORNITHISCHIA COOPER, 1985

GRANDORDER: ORNITHOPODA MARSH, 1871

MICRORDER: MARGINOCEPHALIA SERENO, 1986

ORDER: PACHYCEPHALOSAURIA MARYANSKA & OSMOLSKA, 1974

SUBORDER: DOMOCEPHALINIA SERENO, 1986

INFRAORDER: PACHYCEPHALOSAURIA (MARYANSKA & OSMOLSKA, 1974) COOPER, 1985

Family: PACHYCEPHALOSAURIDAE Sternberg, 1945

Subfamily: PACHYCEPHALOSAURINAE Sternberg, 1945 (sensu Sereno, 1998)

Stygimoloch spinifer

_____________________________________________________________________________________

NIOBRARA COUNTY

LOCALITY 1

LANCE FORMATION

LANCIAN AGE

UPPER MAASTRICHTIAN STAGE

UPPERMOST SENONIAN SUBEPOCH

UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: AMPHIBIA LINNAEUS, 1758

SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?

SUPERORDER: BATRACHIA BRONGNIART, 1800

ORDER: CAUDATA SCOPOLI, 1777

SUBORDER: AMBYSTOMATOIDEA NOBLE, 1931

Family: SCAPHERPETONTIDAE Auffenberg & Goin, 1959

Scapherpeton sp

NIOBRARA COUNTY

BUG CREEK

LANCE FORMATION

LANCIAN AGE

UPPER MAASTRICHTIAN STAGE

UPPERMOST SENONIAN SUBEPOCH

UPPER UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: ORNITHISCHIFORMES COOPER, 1985

MAGNORDER: ORNITHISCHIA SEELEY, 1888

SUPERORDER: NEORNITHISCHIA COOPER, 1985

GRANDORDER: ORNITHOPODA MARSH, 1871

MICRORDER: MARGINOCEPHALIA SERENO, 1986

ORDER: CERATOPIA MARSH, 1890

SUBORDER: NEOCERATOPIA SERENO, 1986

INFRAORDER: CORONOSAURIA SERENO, 1986

PARVORDER: CERATOPOMORPHA Wolfe & Kirkland, 1998 (emend)

Superfamily: CERATOPOIDEA Marsh, 1888 (sensu Hay, 1902)

Family: CERATOPIDAE Marsh, 1888

Subfamily: CERATOPIINAE Marsh, 1888 (sensu Abel, 1919)

Tribe: TRICERATOPINI nova

Triceratops horridus
= Triceratops serratus

_____________________________________________________________________________________

WYOMING

NIOBRARA COUNTY

ALKALI CREEK

LANCE FORMATION

LANCIAN AGE

UPPER MAASTRICHTIAN STAGE

UPPERMOST SENONIAN SUBEPOCH

UPPER UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: TETANURAE GAUTHIER, 1986

MICRORDER: COELUROSAURIA HUENE, 1914

ORDER: MANIRAPTORFORMES HOLTZ, 1996

AVETHEROPODA PAUL, 2001

SUBORDER: COELURIA MARSH, 1881

Superfamily: TYRANNOSAUROIDEA Osborn, 1906 (sensu Bonaparte et. al., 1990)

Family: TYRANNOSAURIDAE Osborn, 1906

Subfamily: ALBERTOSAURINAE Currie, Hurum & Sabath, 2003

Albertosaurus sarcophagus
= *Ornithomimus grandis
= Aublysodon grandis
= Deinodon grandis

Subfamily: TYRANNOSAURINAE Osborn, 1906

Tribe: TYRANNOSAURINI Osborn, 1906 (sensu Olshevsky, Ford & Yamamoto, 1995)

Tyrannosaurus sp

SUBORDER: ORNITHOMIMOSAURIA BARSBOLD, 1976

Family: ORNITHOMIMIDAE Marsh, 1890

*Ornithomimus sedens

_____________________________________________________________________________________

NIOBRARA COUNTY

A MAIN DIVIDE BETWEEN BUCK CREEK AND LANCE CREEK

LANCE FORMATION

LANCIAN AGE

UPPER MAASTRICHTIAN STAGE

UPPERMOST SENONIAN SUBEPOCH

UPPER UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: ORNITHISCHIFORMES COOPER, 1985

MAGNORDER: ORNITHISCHIA SEELEY, 1888

SUPERORDER: NEORNITHISCHIA COOPER, 1985

GRANDORDER: ORNITHOPODA MARSH, 1871

MICRORDER: MARGINOCEPHALIA SERENO, 1986

ORDER: CERATOPIA MARSH, 1890

SUBORDER: NEOCERATOPIA SERENO, 1986

INFRAORDER: CORONOSAURIA SERENO, 1986

PARVORDER: CERATOPOMORPHA Wolfe & Kirkland, 1998 (emend)

Superfamily: CERATOPOIDEA Marsh, 1888 (sensu Hay, 1902)

Family: CERATOPIDAE Marsh, 1888

Subfamily: CERATOPIINAE Marsh, 1888 (sensu Abel, 1919)

Tribe: TRICERATOPINI nova

Triceratops horridus
= *Triceratops flabellatus
= Sterrholophus flabellatus

_____________________________________________________________________________________

NIOBRARA COUNTY

9 M SW OF WARREN POST OFFICE ON BUG CREEK

LANCE FORMATION

LANCIAN AGE

UPPER MAASTRICHTIAN STAGE

UPPERMOST SENONIAN SUBEPOCH

UPPER UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: ORNITHISCHIFORMES COOPER, 1985

MAGNORDER: ORNITHISCHIA SEELEY, 1888

SUPERORDER: NEORNITHISCHIA COOPER, 1985

GRANDORDER: ORNITHOPODA MARSH, 1871

MICRORDER: MARGINOCEPHALIA SERENO, 1986

ORDER: PACHYCEPHALOSAURIA MARYANSKA & OSMOLSKA, 1974

SUBORDER: DOMOCEPHALINIA SERENO, 1986

INFRAORDER: PACHYCEPHALOSAURIA (MARYANSKA & OSMOLSKA, 1974) COOPER, 1985

Family: PACHYCEPHALOSAURIDAE Sternberg, 1945

Subfamily: PACHYCEPHALOSAURINAE Sternberg, 1945 (sensu Sereno, 1998)

*Pachycephalosaurus wyomingensis
= Troodon wyomingensis
= Stegoceras wyomingense
= Stegoceras wyomingensis

_____________________________________________________________________________________

NIOBRARA COUNTY

“BEECHER’S QUARRY”

LANCE FORMATION

LANCIAN AGE

UPPER MAASTRICHTIAN STAGE

UPPERMOST SENONIAN SUBEPOCH

UPPER UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: ORNITHISCHIFORMES COOPER, 1985

MAGNORDER: ORNITHISCHIA SEELEY, 1888

SUPERORDER: NEORNITHISCHIA COOPER, 1985

GRANDORDER: ORNITHOPODA MARSH, 1871

MICRORDER: EUORNITHOPODA SERENO, 1986

ORDER: IGUANODONTIA DOLLO, 1888

Family: THESCELOSAURIDAE Sternberg, 1937

Thescelosaurus neglectus

_____________________________________________________________________________________

NIOBRARA COUNTY

2 OR 3 M N OF DOGIE CREEK

LANCE FORMATION

LANCIAN AGE

UPPER MAASTRICHTIAN STAGE

UPPERMOST SENONIAN SUBEPOCH

UPPER UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: ORNITHISCHIFORMES COOPER, 1985

MAGNORDER: ORNITHISCHIA SEELEY, 1888

SUPERORDER: NEORNITHISCHIA COOPER, 1985

GRANDORDER: ORNITHOPODA MARSH, 1871

MICRORDER: EUORNITHOPODA SERENO, 1986

ORDER: IGUANODONTIA DOLLO, 1888

SUBORDER: ANKYLOPOLLEXIA SERENO, 1986

INFRAORDER: STYRACOSTERNA SERENO, 1986

PARVORDER: HADROSAURIFORMES SERENO, 1997

SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)

Family: HADROSAURIDAE COPE, 1869

Subfamily: HADROSAURINAE Cope, 1869 (sensu Lambe, 1918)

Tribe: EDMONTOSAURINI Brett-Surman vide Chapman & Brett-Surman, 1990

Anatosaurus annectens

____________________________________________________________________________________

WYOMING

NIOBRARA COUNTY

DOGIE CREEK

LANCE FORMATION

LANCIAN AGE

UPPER MAASTRICHTIAN STAGE

UPPERMOST SENONIAN SUBEPOCH

UPPER UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: ORNITHISCHIFORMES COOPER, 1985

MAGNORDER: ORNITHISCHIA SEELEY, 1888

SUPERORDER: NEORNITHISCHIA COOPER, 1985

GRANDORDER: ORNITHOPODA MARSH, 1871

MICRORDER: EUORNITHOPODA SERENO, 1986

ORDER: IGUANODONTIA DOLLO, 1888

Family: THESCELOSAURIDAE Sternberg, 1937

*Thescelosaurus neglectus

MICRORDER: MARGINOCEPHALIA SERENO, 1986

ORDER: CERATOPIA MARSH, 1890

SUBORDER: NEOCERATOPIA SERENO, 1986

INFRAORDER: CORONOSAURIA SERENO, 1986

PARVORDER: CERATOPOMORPHA Wolfe & Kirkland, 1998 (emend)

Superfamily: CERATOPOIDEA Marsh, 1888 (sensu Hay, 1902)

Family: CERATOPIDAE Marsh, 1888

Subfamily: CERATOPIINAE Marsh, 1888 (sensu Abel, 1919)

Tribe: TRICERATOPINI nova

Triceratops horridus
= *Triceratops ingens
Triceratops cf. albertensis
Triceratops sp

_____________________________________________________________________________________

NIOBRARA COUNTY

ABOUT 3 M N OF MOUTH OF DOGIE CREEK

LANCE FORMATION

LANCIAN AGE

UPPER MAASTRICHTIAN STAGE

UPPERMOST SENONIAN SUBEPOCH

UPPER UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: ORNITHISCHIFORMES COOPER, 1985

MAGNORDER: ORNITHISCHIA SEELEY, 1888

SUPERORDER: NEORNITHISCHIA COOPER, 1985

GRANDORDER: ORNITHOPODA MARSH, 1871

MICRORDER: EUORNITHOPODA SERENO, 1986

ORDER: IGUANODONTIA DOLLO, 1888

SUBORDER: ANKYLOPOLLEXIA SERENO, 1986

INFRAORDER: STYRACOSTERNA SERENO, 1986

PARVORDER: HADROSAURIFORMES SERENO, 1997

SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)

Family: HADROSAURIDAE COPE, 1869

Incertae sedis

Gen. sp indet.

____________________________________________________________________________________

NIOBRARA COUNTY

DRAGON’S GRAVE

LANCE FORMATION

LANCIAN AGE

UPPER MAASTRICHTIAN STAGE

UPPERMOST SENONIAN SUBEPOCH

UPPER UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

ORDER: Incertae sedis

cf. Ricardoestesia

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: TETANURAE GAUTHIER, 1986

MICRORDER: COELUROSAURIA HUENE, 1914

ORDER: MANIRAPTORFORMES HOLTZ, 1996

AVETHEROPODA PAUL, 2001

SUBORDER: COELURIA MARSH, 1881

Superfamily: TYRANNOSAUROIDEA Osborn, 1906 (sensu Bonaparte et. al., 1990)

Family: TYRANNOSAURIDAE Osborn, 1906

Subfamily: TYRANNOSAURINAE Osborn, 1906

Tribe: TYRANNOSAURINI Osborn, 1906 (sensu Olshevsky, Ford & Yamamoto, 1995)

Nanotyrannus sp

SUBORDER: MANIRAPTORA GAUTHIER, 1986

INFRAORDER: PARVES SERENO, 1997

PARVORDER: DEINONYCHOSAURIA COLBERT & D. A. RUSSELL, 1969

Family: DROMAEOSAURIDAE Russell, 1969

Subfamily: DROMAEOSAURINAE Matthew & Brown, 1922

Dromaeosaurus sp

Subfamily: VELOCIRAPTORINAE Barsbold, 1978

Saurornitholestes sp

Family: TROODONTIDAE Gilmore, 1924

Troodon formosus

Genus nova?

COHORT: ORNITHISCHIFORMES COOPER, 1985

MAGNORDER: ORNITHISCHIA SEELEY, 1888

SUPERORDER: NEORNITHISCHIA COOPER, 1985

GRANDORDER: ORNITHOPODA MARSH, 1871

MICRORDER: EUORNITHOPODA SERENO, 1986

ORDER: IGUANODONTIA DOLLO, 1888

SUBORDER: ANKYLOPOLLEXIA SERENO, 1986

INFRAORDER: STYRACOSTERNA SERENO, 1986

PARVORDER: HADROSAURIFORMES SERENO, 1997

SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)

Family: HADROSAURIDAE COPE, 1869

Subfamily: HADROSAURINAE Cope, 1869 (sensu Lambe, 1918)

Tribe: EDMONTOSAURINI Brett-Surman vide Chapman & Brett-Surman, 1990

Anatosaurus annectens

VETEROVATA

OOSUPERCOHORT: DINOSAURIOVUM VIALOV, 1975

OOCOHORT: THEROPODA NOVA

oofamily: ELONGATOOLITHIDAE Zhao, 1975

Elongatoolithus

Macroolithus
Note: TROODONTIDAE?

____________________________________________________________________________________

NIOBRARA COUNTY

NORTH OF LANCE CREEK

T64W, R38N

LANCE FORMATION

LANCIAN AGE

UPPER MAASTRICHTIAN STAGE

UPPERMOST SENONIAN SUBEPOCH

UPPER UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: ORNITHISCHIFORMES COOPER, 1985

MAGNORDER: ORNITHISCHIA SEELEY, 1888

SUPERORDER: NEORNITHISCHIA COOPER, 1985

GRANDORDER: ORNITHOPODA MARSH, 1871

MICRORDER: EUORNITHOPODA SERENO, 1986

ORDER: IGUANODONTIA DOLLO, 1888

SUBORDER: ANKYLOPOLLEXIA SERENO, 1986

INFRAORDER: STYRACOSTERNA SERENO, 1986

PARVORDER: HADROSAURIFORMES SERENO, 1997

SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)

Family: HADROSAURIDAE COPE, 1869

Subfamily: HADROSAURINAE Cope, 1869 (sensu Lambe, 1918)

Tribe: EDMONTOSAURINI Brett-Surman vide Chapman & Brett-Surman, 1990

Anatosaurus annectens

____________________________________________________________________________________

NIOBRARA COUNTY

GREASEWOOD CREEK LOCALITY

SDSM LOCALITY V865

LANCE FORMATION

LANCIAN AGE

UPPER MAASTRICHTIAN STAGE

UPPERMOST SENONIAN SUBEPOCH

UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: AMPHIBIA LINNAEUS, 1758

SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?

SUPERORDER: BATRACHIA BRONGNIART, 1800

ORDER: CAUDATA SCOPOLI, 1777

SUBORDER: PORTEOIDEA (NOBEL 1931) EMEND EDWARDS, 1976

Family: BATRACHOSAUROIDIDAE Auffenberg, 1958

Opisthotriton kayi

SUBORDER: AMBYSTOMATOIDEA NOBLE, 1931

Family: SCAPHERPETONTIDAE Auffenberg & Goin, 1959

Scapherpeton tectum

SUBORDER: SALAMANDROIDEA NOBLE, 1931

Family: SIRENIDAE Gray, 1825

Habrosaurus dilatus

NIOBRARA COUNTY

GREASEWOOD CREEK LOCALITY

SDSM LOCALITY V852

LANCE FORMATION

LANCIAN AGE

UPPER MAASTRICHTIAN STAGE

UPPERMOST SENONIAN SUBEPOCH

UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: AMPHIBIA LINNAEUS, 1758

SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?

SUPERORDER: BATRACHIA BRONGNIART, 1800

ORDER: CAUDATA SCOPOLI, 1777

SUBORDER: PORTEOIDEA (NOBEL 1931) EMEND EDWARDS, 1976

Family: BATRACHOSAUROIDIDAE Auffenberg, 1958

Opisthotriton kayi

SUBORDER: AMBYSTOMATOIDEA NOBLE, 1931

Family: SCAPHERPETONTIDAE Auffenberg & Goin, 1959

Scapherpeton tectum

SUBORDER: SALAMANDROIDEA NOBLE, 1931

Family: SIRENIDAE Gray, 1825

Habrosaurus dilatus

NIOBRARA COUNTY

LEONARD’S LIZARD SITE

SDSM LOCALITY V853

LANCE FORMATION

LANCIAN AGE

UPPER MAASTRICHTIAN STAGE

UPPERMOST SENONIAN SUBEPOCH

UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: AMPHIBIA LINNAEUS, 1758

SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?

SUPERORDER: BATRACHIA BRONGNIART, 1800

ORDER: CAUDATA SCOPOLI, 1777

SUBORDER: AMBYSTOMATOIDEA NOBLE, 1931

Family: SCAPHERPETONTIDAE Auffenberg & Goin, 1959

Scapherpeton tectum

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUPERCOHORT CHORISTODERA NOVA

ORDER: CHORISTODERA COPE, 1876

Family: CHAMPSOSAURIDAE Cope, 1884

Champsosaurus laramiensis

NIOBRARA COUNTY

PETERSON’S QUARRY

LANCE FORMATION

LANCIAN AGE

UPPER MAASTRICHTIAN STAGE

UPPERMOST SENONIAN SUBEPOCH

UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: AMPHIBIA LINNAEUS, 1758

SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?

SUPERORDER: BATRACHIA BRONGNIART, 1800

ORDER: CAUDATA SCOPOLI, 1777

SUBORDER: AMBYSTOMATOIDEA NOBLE, 1931

Family: SCAPHERPETONTIDAE Auffenberg & Goin, 1959

Scapherpeton sp

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: SERPENTES LINNAEUS, 1759

ORDER: OPHIDIA BRONGNIART, 1800

SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923

Superfamily: ANILIODEA Fitzinger, 1826

Family: ANILIIDAE Fitzenger, 1826

*Coniophis precedens

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: TETANURAE GAUTHIER, 1986

MICRORDER: COELUROSAURIA HUENE, 1914

ORDER: MANIRAPTORFORMES HOLTZ, 1996

AVETHEROPODA PAUL, 2001

SUBORDER: COELURIA MARSH, 1881

Superfamily: TYRANNOSAUROIDEA Osborn, 1906 (sensu Bonaparte et. al., 1990)

Family: TYRANNOSAURIDAE Osborn, 1906

Subfamily: TYRANNOSAURINAE Osborn, 1906

Tribe: TYRANNOSAURINI Osborn, 1906 (sensu Olshevsky, Ford & Yamamoto, 1995)

Tyrannosaurus rex
= Manospondylus gigas
= *Aublysodon amplus
= Deinodon amplus
= Manospondylus amplus
= Tyrannosaurus amplus
= Stygivenator amplus

COHORT: ORNITHISCHIFORMES COOPER, 1985

MAGNORDER: ORNITHISCHIA SEELEY, 1888

SUPERORDER: EURYOPDA SERENO, 1986

GRANDORDER: ARMATOSAURIA ZHAO, 1983 (emend)

ORDER: ANKYLOSAURIA OSBORN, 1923

SUBORDER: SCELIDOSAURIA COOPER, 1985

INFRAORDER: ANKYLOSAURA OSBORN, 1923

Family: NODOSAURIDAE Marsh, 1890

*Palaeoscincus latus

NIOBRARA COUNTY

ON THE NORTH PLATTE RIVER

LANCE FORMATION

LANCIAN AGE

UPPER MAASTRICHTIAN STAGE

UPPERMOST SENONIAN SUBEPOCH

UPPER UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: ORNITHISCHIFORMES COOPER, 1985

MAGNORDER: ORNITHISCHIA SEELEY, 1888

SUPERORDER: NEORNITHISCHIA COOPER, 1985

GRANDORDER: ORNITHOPODA MARSH, 1871

MICRORDER: EUORNITHOPODA SERENO, 1986

ORDER: IGUANODONTIA DOLLO, 1888

SUBORDER: ANKYLOPOLLEXIA SERENO, 1986

INFRAORDER: STYRACOSTERNA SERENO, 1986

PARVORDER: HADROSAURIFORMES SERENO, 1997

SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)

Family: HADROSAURIDAE COPE, 1869

Incertae sedis

Gen. sp indet.

____________________________________________________________________________________

NIOBARA COUNTY

UC LOCALITIY V-5002

LANCE FORMATION

LANCIAN AGE

UPPER MAASTRICHTIAN STAGE

UPPERMOST SENONIAN SUBEPOCH

UPPER UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990

COHORT: CROCODYLOTARSI BENTON & CLARK, 1988

MAGNORDER: SUCHIA KREBS, 1974

SUPERORDER: CROCODYLOMORPHA WALKER, 1968

GRANDORDER: CROCODILIFORMES HAY, 1930

MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983

ORDER: EUSUCHIA emend HUXLEY, 1875

INFRAORDER: CROCODYLIA emend GMELIN, 1877

PARVORDER: BREVIROSTRES

Superfamily: ALLIGATORIDEA Buffetaut, 1985

Family: LEIDYOSUCHIDAE Rauhe & Rossmann, 1995

Leidyosuchus canadensis

GLOBIDONTA BROUCHU, 1999

Family: ALLIGATORIDAE Gray, 1844

Brachychampsa montana

CLASS: MAMMALIA LINNAEUS, 1758

SUBCLASS: THERIIFORMES (ROWE, 1988) McKENNA & BELL, 1997

INFRACLASS: ALLOTHERIA (MARSH, 1880) McKENNA & BELL, 1997

ORDER: MULTITUBERCULATA COPE, 1884

SUBORDER: CIMOLODONTA McKENNA, 1975

Superfamily: PTILODONTOIDEA Cope, 1887 (sensu McKenna & Bell, 1997)

Family: CIMOLODONTIDAE Marsh, 1889

Cimolodon nitidus

Family: PTILODONTIDAE Cope, 1887 (sensu McKenna & Bell, 1997)

Subfamily: NEOPLAGIAULACINAE Ameghino, 1890 (sensu McKenna & Bell, 1997)

Mesodma thompsoni
Mesodma sp

Superfamily: TAENIOLABIDOIDEA Granger & Simpson, 1929 (sensu McKenna & Bell, 1997)

Family: TAENIOLABIDIDAE Granger & Simpson, 1929

Meniscoessus robustus

SUBCLASS: PROTOTHERIA GILL, 1872

INFRACLASS: TRECHNOTHERIA McKENNA, 1975

LEGION: CLADOTHERIA McKENNA, 1975

SUBLEGION: ZATHERIA MCKENNA, 1975

INFRALEGION: TRIBOSPHENIDA (McKENNA, 1975) McKENNA & BELL, 1997

SUPERCOHORT: THERIA (PARKER & HASWELL, 1987) McKENNA & BELL, 1997

COHORT: MARSUPIALIA (ILLIGER, 1811) McKENNA & BELL, 1997

ORDER: STAGODONTIA ARCHER, 1984

Family: STAGODONTIDAE Marsh, 1889

Didelphodon vorax

Family: PEDIOMYIDAE Simpson, 1927

Leptalestes? cooki
= Pediomys cooki

COHORT: PLACENTALIA (OWEN, 1837) McKENNA & BELL, 1997

MAGNORDER: EPITHERIA (McKENNA, 1975) McKENNA & BELL, 1997

SUPERORDER: PREPTOTHERIA McKENNA, 1975

GRANDORDER: UNGULATOMORPHA ARCHIBALD, 1996

Family: GYPSONICTIPIDAE Van Valen, 1967

Gypsonictops hypoconus

NIOBRARA COUNTY

POWDER RIVER BASIN

UC LOCALITY V-5003

LANCE FORMATION

LANCIAN AGE

UPPER MAASTRICHTIAN STAGE

UPPERMOST SENONIAN SUBEPOCH

UPPER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: AMPHIBIA LINNAEUS, 1758

SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?

SUPERORDER: BATRACHIA BRONGNIART, 1800

ORDER: CAUDATA SCOPOLI, 1777

SUBORDER: PORTEOIDEA (NOBEL 1931) EMEND EDWARDS, 1976

Family: BATRACHOSAUROIDIDAE Auffenberg, 1958

Opisthotriton kayi

Prodesmodon copei

SUBORDER: AMBYSTOMATOIDEA NOBLE, 1931

Family: SCAPHERPETONTIDAE Auffenberg & Goin, 1959

Scapherpeton tectum

SUBORDER: SALAMANDROIDEA NOBLE, 1931

Family: SIRENIDAE Gray, 1825

Habrosaurus dilatus

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: CHELONOMORPHA KUHN, 1960

SUPERCOHORT: TESTUDINES LINNAEUS, 1758

COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988

MAGANORDER: CRYPTODIRA GAFFNEY & MEYLAN, 1988

SUPERORDER: SELMACRYPTODIRA GAFFNEY, HUTCHISON, JENKINS & MEEKR, 1987

GRANDORDER: DAIOCRYPTODIRA GAFFNEY & MEYLAN, 1988

MICRORDER: BAENOIDEA WILLIAMS, 1960

Family: BAENIDAE Cope, 1882

Subfamily: BAENINAE Cope, 1882 (sensu Williams, 1950)

Baena sp

Family: NEURANKYLIDAE Lambe, 1902

Compsemys victa

MICRORDER: CENTROCRYPTODIRA GAFFNEY & MEYLAN, 1988

ORDER: POLYCRPTODIRA GAFFNEY & MEYLAN, 1988

SUBORDER: CHELYDROIDEA AGASSIZ, 1857

Family: CHELYDRIDAE Agassiz, 1857

Gen. sp indet.

SUBORDER: PROCOELOCRYPTODIRA GAFFNEY & MEYLAN, 1988

INFRAORDER: CHELOMACRYPTODIRA GAFFNEY & MEYLAN, 1988

PARVORDER: TRIONYCHIDA GRAY, 1870

Superfamily: TRIONYCHIDEA Bell, 1828 (sensu Gray, 1870)

Family: TRIONYCHIDAE Bell, 1828

Aspideretes beecheri

cf Helopanoplia

Ge.n sp indet.

Family: ADOCIDAE Cope, 1870

Adocus sp

Family: NANHSIUNGCHELYIDAE Yeh, 1966

Basilemys sp

Superfamily: TESTUDINOIDEA Baur, 1893

Family: EMYDIDAE Gray, 1825

Subfamily: EMYDINAE Gray, 1825

Gen. sp indet.

SUBCLASS: DIAPSIDA OSBORN, 1904

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA DUMERIL & BIBRON, 1839

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA OPPEL, 1811

SUPERORDER: LACERTILIA OWEN, 1842

GRANDORDER: ASCALABOTA GILMORE, 1928

ORDER: IGUANIA LATREILLE, 1825

SUBORDER: DRACOMORPHA NORTHCUTT, 1978

INFRAORDER: TEIUMORPHA ALIFANOV, 2000

Family: TEIIDAE Gray, 1827

Subfamily: CHAMOPSINAE Denton & O’Neill 1995

Chamops segnis

GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961

MICRORDER: SCINCOMORPHA CAMP, 1923

ORDER: ANGUIMORPHA FURBRINGER, 1900

SUBORDER: DIPLOGLOSSA COPE, 1864

INFRAORDER: OPHISAUROMORPHA ALIFANOV, 2000

Family: ANGUIDAE Gray, 1825

Subfamily: GERRHONOTINAE McDowell & Bogert, 1954

cf. Gerrhonotus

SUBORDER: PLATYNOTA GILMORE, 1928

INFRAORDER: HELODERMOIDA COPE, 1900

Family: HELODERMATIDAE Gray, 1837

Paraderma bogerti

SUBORDER: DIPLOGLOSSA COPE, 1864

INFRAORDER: XENOSAUIRA FURBRINGER, 1900

Family: XENOSAURIDAE Cope, 1886

Exostinus lancensis

INFRAORDER: OPHISAUROMORPHA ALIFANOV, 2000

Family: ANGUIDAE Gray, 1825

Subfamily: GLYPTOSAURINAE Marsh, 1872

Tribe: MELANOSAURINI Sullivan, 1979

Odaxosaurus piger

INFRAORDER: THECAGLOSSA COPE, 1890

PARVORDER: MONSTERSAURIA NORELL & GAO, 1977

Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)

Family: VARANIDAE Gray, 1827

Palaeosaniwa canadensis

Family: PARASANIWIDAE Estes, 1964

Parasaniwa wyomingensis

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990

COHORT: CROCODYLOTARSI BENTON & CLARK, 1988

MAGNORDER: SUCHIA KREBS, 1974

SUPERORDER: CROCODYLOMORPHA WALKER, 1968

GRANDORDER: CROCODILIFORMES HAY, 1930

MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983

ORDER: EUSUCHIA emend HUXLEY, 1875

INFRAORDER: CROCODYLIA emend GMELIN, 1877

PARVORDER: BREVIROSTRES

Superfamily: ALLIGATORIDEA Buffetaut, 1985

Family: LEIDYOSUCHIDAE Rauhe & Rossmann, 199

Leidyosuchus canadensis

GLOBIDONTA BROUCHU, 1999

Family: ALLIGATORIDAE Gray, 1844

Brachychampsa montana

SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

INFRALEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

COHORT: THEROPODA EMEND MARSH, 1881

AVEPODA PAUL, 2002

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: TETANURAE GAUTHIER, 1986

ORDER: NEOTETANURAE SERENO, WILSON, LARSSON, DUTHEIL &

SUBORDER: CARNOSAURIA HUENE, 1920

Superfamily: Incertae sedis

Family: DRYPTOSAURIDAE Marsh, 1890

Dryptosaurus sp

MICRORDER: COELUROSAURIA HUENE, 1914

ORDER: MANIRAPTORFORMES HOLTZ, 1996

AVETHEROPODA PAUL, 2001

AVEPECTORA PAUL, 2001

SUBORDER: MANIRAPTORA GAUTHIER, 1986

INFRAORDER: PARVES SERENO, 1997

PARVORDER: DEINONYCHOSAURIA COLBERT & D. A. RUSSELL, 1969

Incertae sedis

Paronychodon lacustris

Family: DROMAEOSAURIDAE Russell, 1969

Incerate sedis

Gen. sp indet.

COHORT: ORNITHISCHIFORMES COOPER, 1985

MAGNORDER: ORNITHISCHIA SEELEY, 1888

SUPERORDER: NEORNITHISCHIA COOPER, 1985

GRANDORDER: ORNITHOPODA MARSH, 1871

MICRORDER: MARGINOCEPHALIA SERENO, 1986

ORDER: CERATOPIA MARSH, 1890

SUBORDER: NEOCERATOPIA SERENO, 1986

INFRAORDER: CORONOSAURIA SERENO, 1986

PARVORDER: CERATOPOMORPHA Wolfe & Kirkland, 1998 (emend)

Superfamily: CERATOPOIDEA Marsh, 1888 (sensu Hay, 1902)

Family: CERATOPIDAE Marsh, 1888

Subfamily: CERATOPIINAE Marsh, 1888 (sensu Abel, 1919)

Tribe: TRICERATOPINI nova

Triceratops sp

CLASS: AVES LINNAEUS, 1758

AVEBREVICAUDA PAUL, 2001

SUBCLASS: CARINATE MERREM, 1813

INFRACLASS: NEORNITHES GADOW, 1893

PARVCLASS: NEOGNATHAE PYCRAFT 1900

ORDER: CHARADRIIFORMES HUXLEY, 1867

SUBORDER: CHARADIIDINE (HUXLEY, 1867) FURBRINGER, 1888

Family: CIMOLOPTERYGIDAE Brodkorb, 1963

*Cimolopteryx minima

CLASS: MAMMALIA LINNAEUS, 1758

SUBCLASS: THERIIFORMES (ROWE, 1988) McKENNA & BELL, 1997

INFRACLASS: ALLOTHERIA (MARSH, 1880) McKENNA & BELL, 1997

ORDER: MULTITUBERCULATA COPE, 1884

SUBORDER: CIMOLODONTA McKENNA, 1975

Superfamily: PTILODONTOIDEA Cope, 1887 (sensu McKenna & Bell, 1997)

Family: CIMOLODONTIDAE Marsh, 1889

Cimolodon nitidus

Family: PTILODONTIDAE Cope, 1887 (sensu McKenna & Bell, 1997)

Subfamily: NEOPLAGIAULACINAE Ameghino, 1890 (sensu McKenna & Bell, 1997)

Mesodma formosa
Mesodma sp

Superfamily: Incertae sedis

Family: CIMOLOMYIDAE Marsh, 1889

Cimolomys gracilis

Family: TAENIOLABIDIDAE Granger & Simpson, 1929

Meniscoessus robustus

SUBCLASS: PROTOTHERIA GILL, 1872

INFRACLASS: TRECHNOTHERIA McKENNA, 1975

LEGION: CLADOTHERIA McKENNA, 1975

SUBLEGION: ZATHERIA MCKENNA, 1975

INFRALEGION: TRIBOSPHENIDA (McKENNA, 1975) McKENNA & BELL, 1997

SUPERCOHORT: THERIA (PARKER & HASWELL, 1987) McKENNA & BELL, 1997

COHORT: MARSUPIALIA (ILLIGER, 1811) McKENNA & BELL, 1997

ORDER: STAGODONTIA ARCHER, 1984

Family: STAGODONTIDAE Marsh, 1889

Didelphodon vorax

Family: PEDIOMYIDAE Simpson, 1927

Pediomys elegans

Protolambda hatcheri
= Pediomys hatcheri
Protolambda florencae
= Pediomys florencae

MAGNORDER: AMERIDELPHIA (SZALAY, in ARCHER, ED, 1982) McKENNA & BELL, 1997

ORDER: DIDELPHIMORPHIA GILL, 1872

Family: DIDELPHIDAE Gray 1821

Turgidodon rhaister
= Alaphadon rhaister

COHORT: PLACENTALIA (OWEN, 1837) McKENNA & BELL, 1997

MAGNORDER: EPITHERIA (McKENNA, 1975) McKENNA & BELL, 1997

SUPERORDER: PREPTOTHERIA McKENNA, 1975

GRANDORDER: FERAE LINNAEUS, 1758

ORDER: CIMOLESTA McKENNA, 1975

SUBORDER: DIDELPHODONTA McKENNA, 1975

Family: CIMOLESTIDAE Marsh, 1889

Altacreodus magnus = Cimolestes magnus

Cimolestes incisus

ORDER: LIPOTYPHLA HAECKEL, 1866

SUBORDER: SORICOMORPHA GREGORY, 1910

Family: GEOLABIDIDAE McKenna, 1960

*Batodon tenuis

GRANDORDER: UNGULATOMORPHA ARCHIBALD, 1996

Family: GYPSONICTIPIDAE Van Valen, 1967

Gypsonictops hypoconus