Species: primus FOX, 1971
Etymology:

Holotype: UA 5528

Locality: UA-MR-6, Verdigris Coulee, approximately 18 miles east of the village of Milk River, Warner County, Warner County, Alberta Province, Canada.

Horizon: Upper Milk River Formation.

Biostratigraphy:

Age: Aquilian age, Lower Campanian Stage, Aquilian in age, Senonian subepoch, Upper Gulf epoch, Late Cretaceous.

Material: RUM3.

Referred material:

Localities: UA-MR-4, 6, Verdigris Coulee, approximately 18 miles east of the village of Milk River, Warner County, Warner County, Alberta Province, Canada.

Horizon: Upper Milk River Formation.

Biostratigraphy:

Age: Aquilian age, Lower Campanian Stage, Aquilian in age, Senonian subepoch, Upper Gulf epoch, Late Cretaceous.

Material:

UA 5532: Broken RUM3.

UA 5547: Broken RUM3.

UA 5547: LUM4.

JOHNSON, 1994

UALVP 29599, 29600, 29607: M1.

UALVP 29601, 29603, 29616, 29617: M2.

UALVP 29606, 29610, 29611, 29612: M3.

 

DAVIS, CIFELLI, & COHEN, 2016

Locality: OMNH V1409, Toole County, Montana.

Horizon: Deadhorse Coulee Member, Eagle Formation.

Biostratigraphy:

Age: Santonian Stage, Senonian  Subepoch, Gulf Epoch, Late Cretaceous.

Material:

OMNH 66378: Right M1 missing protoconal region.

OMNH 66377: Left M2 missing protoconal region.

OMNH 66391: Fragmentary left M4.

_____________________________________________________________________________________

Species: secundus JOHANSON, 1994
Etymology:

Holotype: UALVP 29534

Locality: Verdigris Coulee, near Milk River, Warner County, Alberta Province, Canada.

Horizon: Upper Milk River Formation.

Biostratigraphy:

Age: Aquilian age, Lower Campanian Stage, Aquilian in age, Senonian subepoch, Upper Gulf epoch, Late Cretaceous.

Material: M3.

Referred material:

UALVP 29590: M2.

UALVP 29528: M3.