Genus: Machimosaurus MEYER, 1837 emen. 1838
Etymology: Greek, machimo, "pugnacious" (having a combative nature, presumably refering to the robust dentition), and Greek, saurus Latinzed sauros, "lizard."
= Ephoenosaurus (ANONYMOUS, 1839)
Etymology:
= Madrimosaurus MEYER, 1837 (sic)

Species: hugii MEYER, 1837
Etymology: In honor of Franz Joseph Hugi (1791-1855), the Swiss geologist and naturalist.

Lectotype: NMS 8342 (old Palaeontol. Samml. Mus. Solothurn Nr. 96)

Locality: Kreuzen Quarry, St. Verena, near Solothurn Canton, Switzerland.

Horizon: "Retschenbank" der Schildkrotenschichten (= Solothurn Turtle Limestone, Reuchenette Formation).

Biostratigraphy:

Age: Uppermost Kimmeridgian Stage, Middle Malm Epoch, Late Jurassic.

Material: Teeth.

Referred material:

HUENE, 1926

Locality: Solothurn near St. Verena, Solothurn Canton, Switzerland.

Horizon: Portland.

Biostratigraphy:

Age: Middle Tithonian Stage, Upper Malm Epoch, Late Jurassic.

Material:

Number: Not given: Teeth, fragmentary skull, fragmentary snout, fragment of skull, splenial, vertebrae, ulna, metacarpal, right femur, fibula, metatarsals, radius.

 

KARL, GRÖNING, BRAUCKMANN, SCHWARZ & KNÖTSCHIKE, 2010

Locality: Langenberg Quarry, a working quarry of the "Rohstoffbetriebe Oker GmbH and Co." for cement chinese rolex submariner and road gravel, southern slope of the Langenberg, 5 km east of Goslar near the Harz Mountians, Oker, near Goslar, Lower Saschen (Saxony) State, Germany.

Horizon:

Biostratigraphy:

Age: Middle Kimmeridgian Stage, Middle Malm Epoch, Late Jurassic.

Material:

DFMMh 270, 329, 268, 271, 313, 331, 332, 333, 366, 372, 473, 506, 614, 620, 623, 624, 625: Fragmentary teeth.

DFMMh 330, 541: Teeth.

 

KREBS, 1967

Locality: Guimarota lignite mine (Mina de Lignito Guimarota), 1.5 km SSE of Leiria, Leiria District, Portugal.

Horizon:

Biostratigraphy:

Age: Upper Oxfordian Stage, Lower Malm Epoch, Late Jurassic.

Material:

MG-8730-1, MG-8730-3, and unumbered elements: Mid-section of snout, post orbital, a piece of frontal, posterior section of skull, and a dorsal vertebra.

 

LYDEKKER, 1888

Locality: Soleure (or Solothurn), Solothurn State, Switzerland.

Horizon: Portland.

Biostratigraphy:

Age: Middle Tithonian Stage, Upper Malm Epoch, Late Jurassic.

Material:

NHMUK (old BMNH) R233: Cast of type.

NHMUK (old BMNH) 33239, 43638: 3 fragmentary teeth.

NHMUK (old BMNH) R5: 5 fragmentary teeth.

NHMUK (old BMNH): R232: Isolated tooth crown.

YOUNG, HUA, STEEL, FOFFA, BRUSATTE, THURING, MATEUS, RUIZ-OMENACA, HAVLIK, LEPAGE, & de ANDRADE, 2014

NMS 7012: Incomplete snout.

NMS 7015: Incomplete snout, maxilla-palatine fragment.

NMS 7021: Incomplete mandible.

NMS 7029: Temporal and orbital region of a skull.

 

MADZIA, SZCZYGIELSKI & WOLNIEWICZ, 2021

Locality: Czarnoglowy (Zarnglaff), Krzyzanowice, Poland.

Horizon:

Biostratigraphy:

Age: Kimmeridgian stage, Malm Epoch, Late Jurassic.

Material:

GG303-34, GG303-35, GG303-36, GG303-40: Teeth mandible fragment and vertebrae.

 

Locality: Wrzosowo (Fritzow), Poland.

Horizon:

Biostratigraphy:

Age: Kimmerdigian Stage, Malm Epoch, Late Jurassic.

Material:

GG303-38, GG303-39: Teeth.

Note: Originally listed as Ichthyosaurus? sp

 

MARTIN & VINCENT, 2013

Locality: Am Hörnle Quarry, Neufen, Baden-Wurttemberg State, Germany.

Horizon:

Biostratigraphy: Ataxioceras hypselocyclum Zone.

Age: Lower Kimmeridgian Stage, Middle Malm Epoch, Late Jurassic.

Material:

SMNS 91415: Nearly complete skull.

 

MAUBEUGE, 1968

Locality: Near Verdun, Meuse Department, France.

Horizon:

Biostratigraphy:

Age:

Material:

Number: Not given:: Incomplete mandible.

 

MEYER & HUNT, 1998

Locality: From an abandoned quarry close to the village of Oberbuchsiten, Solothurn Canton, Switzerland.

Horizon: Wettingen Member, Villigen Formation.

Biostratigraphy:

Age: Late Oxfordian Stage, Lower Malm Epoch, Late Jurassic.

Material:

Number: Not given:

 

SAUVAGE, 1900

Locality: Fumel, Lot-et-Garonne how to tell a real rolex from a fake Department, France.

Horizon:

Biostratigraphy:

Age: Late Jurassic.

Material:

Number: Not given:

 

SAUVAGE, 1874

Locality: Boulogne-sur-Mer, Pas-De-Calais Department, France.

Horizon:

Biostratigraphy:

Age: Kimmeridgian Stage, Middle Malm Epoch, Late Jurassic.

Material:

Number: Not given: Teeth.

 


SAUVAGE, 1902

Locality: Fumel, Lot-et-Garonne Department, France.

Horizon:

Biostratigraphy:

Age: Late Jurassic.

Material:

Number: Not given: Teeth, fragmentary maxillae, dorsal vertebrae.

 

 

SAUVAGE, 1987/98

Locality: Portugal.

Horizon: Lusitanian.

Biostratigraphy:

Age: Early Jurassic.

Material:

Number: Not given: Teeth and upper and lower jaw fragments.

 

SELENKA, 1867

Locality: Hannover, Saschen (Saxony) State, Germany.

Horizon:

Biostratigraphy:

Age: Kimmeridgian Stage, Middle Malm Epoch, Late Jurassic.

Material:

Number: Not given: Teeth, ribs, vertebrae and pelvic bones.

 

YOUNG, HUA, STEEL, FOFFA, BRUSATTE, THURING, MATEUS, RUIZ-OMENACA, HAVLIK, LEPAGE, & de ANDRADE, 2014

Locality: Porto das Barcas, Lourinha, Estremadura Province, Portugal.

Horizon: Praia Azul Member, Lourinha Formation.

Biostratigraphy:

Age: Upper Kimmeridgian Stage, Middle Malm Epoch, Late Jurassic.

Material:

ML491, ML495, ML959, ML1955: T eeth.

 

Locality: Peralta, Lourinha, Estremadura Province, Portugal.

Horizon: Praia Azul Member, Lourinha Formation.

Biostratigraphy:

Age: Upper Kimmeridgian Stage, Middle Malm Epoch, Late Jurassic.

Material:

ML647: Isolated teeth.

 

Locality: Zimbral, Lourinha, Estremadura Province, Portugal.

Horizon: Praia Azul Member, Lourinha Formation.

Biostratigraphy:

Age: Upper Kimmeridgian Stage, Middle Malm Epoch, Late Jurassic.

Material:

ML657, ML658: Teeth.

 

Locality: Beach near Lourinha, Estremadura Province, Portugal.

Horizon: Praia Azul Member, Lourinha Formation.

Biostratigraphy:

Age: Upper Kimmeridgian Stage, Middle Malm Epoch, Late Jurassic.

Material:

 

Locality: Playa de La Grega, Colunga, Asturias Province, Spain.

Horizon: Terenes Formation.

Biostatigraphy:

Age: Kimmeridgian Stage, Middle Malm Epoch, Late Jurassic.

Material:

MUJA-1008, MUJA-1922: Isolated teeth.

 

Locality: La Escalera, Villaviciosa, Asturias Province, Spain.

Horizon: Lastres Formation.

Biostratigraphy:

Age: Kimmeridgian Stage, Middle Malm Epoch, Late Jurassic.

Material:

MJUA-1298: Tooth crown.

 

Locality: Cantera Carcalin near Bunol, Valencia Province, Spain.

Horizon:

Biostratigraphy:

Age: Tithonian Stage, Upper Malm Epoch, Late Jurassic.

Material:

MCNV-CC-4: Isolated tooth crown.

 

= Ephoenosaurus solodurensis (ANONYMOUS, 1839) (nomen nudum)
Etymology:

Holotype:

Locality:

Horizon:

Biostratigraphy:

Age:

Material:

 

= Ephoenosaurus gracilis (ANONYOMOUS, 1839) (nomen nudum)
Etymology:

Holotype:

Locality:

Horizon:

Biostratigraphy:

Age:

Material:

 

= Machimosaurus bathonicus SAUVAGE, 1874
Etymology:

Holotype:

Locality: Fuller's Earth of Boulogne-sur-Mer, Pas-De-Calais Department, France.

Horizon:

Biostratigraphy:

Age:

Material: Teeth.

 

= Machimosaurus interruptus SAUVAGE, 1874
Etymology:

Holotype:

Locality: Boulogne-sur-Mer, Pas-De-Calais Department, France.

Horizon:

Biostratigraphy:

Age: Upper Kimmeridgian Stage and Lower Portlandian, Middle Tithonian Stage, Upper Malm Epoch, Late Jurassic.

Material: Teeth.

Referred material:

SAUVAGE, 1900

Locality: Indre Department, France.

Horizon:

Biostratigraphy:

Age: Early Jurassic.

Material:

Number: Not given:

 

= Machimosaurus ferox SAUVAGE, 1874
Etymology:

Holotype:

Locality: Chatillon, France.

Horizon:

Biostratigraphy:

Age: Kimmeridgian Stage, Middle Malm Epoch, Late Jurassic.

Material: Tooth.

 

= Machimosaurus rigauxi SAUVAGE, 1874
Etymology:

Holotype:

Locality: Boulonnais, Pas-de-Calais Department, France.

Horizon: Forest Marble.

Biostratigraphy:

Age: Upper Bathonian Stage, Upper Middle Epoch, Early Cretaceous.

Material: Tooth.

 

= Pliosaurus trochanterius OWEN, 1869
Etymology:
= Metorhynchus trochanterius (OWEN, 1869)

Holotype:

Locality: Dorset County, England, Southern United Kingdom.

Horizon:

Biostratigraphy:

Age: Kimmeridgian Stage, Middle Malm Epoch, Late Jurassic.

Material: Occipital region of a cranium and an associated mandible.

 

_____________________________________________________________________________________

Species: mosae SAUVAGE & LIENARD, 1879 emend HUA, VASSE, BUFFETAUT, MARTIN, MAZIN & VADET, 1993
Etymology: In reference to teh French river Meuse, near which the holotype was discovered; 'Punacious lizared of the Meuse'.
= Machimosaurus mosae mosae SAUVAGE & LIENARD, 1879 emend HUA, VASSE, BUFFETAUT, MARTIN, MAZIN & VADET, 1993

Holotype: Disappeard during the First World War and is presumed to have been destroyed (Young, et al,. 2014)

Locality: Issoncourt, near Verdun, Department de la Meuse, Lorraine, France.

Horizon: 'Marnes superieures de la Meuse'

Biostratigraphy: A. autissiodorensis Sub-Boreal ammonite zone.

Age: Kimmeridgian Stage, Middle Malm Epoch, Late Jurassic.

Material: Incomplete skull, mandible, 22 vertebrae, part of the pelvis, numerous ribs, limb bones and 22 osteoderms.

Neotype: Museum d’Histore Naturelle de Boulonge-sur-Mer, No. 1100

Locality: Beach near Ambleteuse, Boulonnais, Pas-de-Calais Department, France.

Horizon: Argiles de Chatillon Formation.

Biostratigraphy: Aulocostephanus autissiodorensis zone.

Age: Kimmeridgian Stage, Middle Malm Epoch, Late Jurassic.

Material: Skull and skeleton.

Referred material:

SAUVAGE & LIENARD, 1879

Locality: Issoncourt near Verdun, Tarn-et-Garonne Department, France.

Horizon:

Biostratigraphy:

Age: Kimmeridgian Stage, Middle Malm Epoch, Late Jurassic.

Material:

Number: Not given: Lower jaw, the greater part of a skull, 22 vertebrae, part of the pelvis, numerous ribs, limb bones and 22 scutes.

Note: Now lost.

 


Locality: Kimmeridge Bay, Gaulter Gap-Broad Bench, Dorset County, England, Southern United Kingdom.

Horizon: Kimmeridge Clay.

Biostratigraphy:

Age: Early Kimmeridgian Stage, Middle Malm Epoch Late Jurassic.

Material:

BMNH R1089: Fragmentary skull and mandible.

_____________________________________________________________________________________

Species: buffetauti YOUNG, HUA, STEEL, FOFFA, BRUSATTE, THÜRING, MATEUS, RUIZ-OMENACA, HAVLIK, LEPAGE & DE ANDRADE, 2014
Etymology: In honor of Eric Buffetaut (b. 1950), whose research was greatly elucidated thalattosuchian and crocodylomorph evolution; 'Buffetaut's pugnacious lizard'.
= Steneosaurus burdigalensis CHANEL, 1905 (nomen nudum)
= Steneosaurus burgobressiensis CHANEL, 1905 (nomen nudum)
= Steneosaurus burgensis CHANEL, 1905 (nomen nudum)
= Steneosaurus montmerli CHANEL, 1905 (nomen nudum)
= Steneosaurus chanuti CHANEL, 1905 (nomen nudum)
= Steneosaurus burgensis chanuti CHANEL, 1905 (nomen nudum)

Holotype: SMNS 91415

Locality: Am Hörnle Quarry, Neuffen, Baden-Württemberg State, Germany.

Horizon: Lacunosamergel Formation.

Biostratigraphy: Ataxioceras hypselocyclum Sub-Mediterranean ammointe zone (Weißer Jura gamma 2)

Age: Lower Kimmeridigian Stage, Middle Malm Epoch, Late Jurassic.

Material: Complete skull and mandible, with partial postcranial skeleton.

Referred material:

 

Locality: Cricqueboeuf, Normandy, Northern France.

Horizon: Calcaires Coquilliers Formation.

Biostratigraphy: P. baylei Sub-Boreal ammonite zone.

Age: Middle Kimmeridigan Stage, Middle Malm Epoch, Late Jurassic.

Material:

MPV V.1600.Bo, V.1601.Bo: Anterior half of rostrum (premaxilla, maxilla, and dentary) in occlusion and a maxilla-nasal fragment.

 

Locality: Langenberg near Oker, Lower Saxony State, Germany.

Horizon: Langenberg Formation.

Biostratigraphy:

Age: Kimmeridgian Stage, Middle Malm Epoch, Late Jurassic.

Material:

DFMMh FV 330, FV 541: Isoalted tooth crowns.

 

Locality: Montmerle, Bourgen-Bresse, Ain Department, France.

Horizon: Calcaires a pteroceras Formation.

Biostratigraphy:

Age: Lower Kimmeridgian Stage, iwc imitation Middle Malm Epoch, Late Jurassic.

Material:

Musee de Brou, number not known: A complete skull and mandible in articulation.

 

= Machimosaurus cf. buffetauti MADZIA, SZCZYGIELSKI & WOLNIEWICZ, 2021

Locality: Czarnoglowy (Zarnglaff), Krzyzanowice, Poland.

Horizon:

Biostratigraphy:

Age: Kimmeridgian stage, Malm Epoch, Late Jurassic.

Material:

GPIT/RE/328, GPI/RE/9280, GPIT/RE/9281: Teeth.


_____________________________________________________________________________________

Species: nowackianus (HUENE, 1938) BARDET, & HUA,1996
Etymology:
= Simolestes nowakianus HUENE, 1938
= Simolestes novakianus BARDET, 1994 (sic)

Holotype: GPIT, Tubingen University.

Locality: Feyambiro near Harrar, Harrar Province, Ethiopia.

Horizon:

Biostratigraphy:

Age: Oxfordian or Kimmeridgian Stage, Lower-Middle Malm Epoch, Late Jurassic.

Material: Premaxillae and broken teeth.

_____________________________________________________________________________________

Species: rex FANTI, MIYASHITA, CANTELLI, MNASRI, DRIDI, CONTESSI, & CAU, 2015
Etymology: Latin, rex, "king," in reference to its majestic size among known Machimosaurus and all thalattosuchians.

Holotype: ONM NG NG 1-25, 80, 81, 83-87

Locality: Touil el Mhahir, Tatouine Governorate, Tunisia.

Horizon: Souiret Sand Member, Douiret Formation.

Biostratigraphy:

Age: Hauterivian/early Barremian Stage, Upper Neocomian Subepoch, Middle Early Cretaceous Epoch, Early Cretaceous.

Material: Fragmentary skull, teeth, fragmentary vertebrae, and osteoderms.