Species: clarki BUCKLEY, BROCHU, KRAUSE & POL, 2000
Etymology: In honor of James M. Clark in recognition of his contributions to the crocodyliform systematics.

Holotype: UA 8679

Locality: Field locality MAD98-17, southeast of the village of Berivotra, Mahajanga Basin, Mahanjanga Province, northwestern Madagascar.

Horizon: Maevarnoa Formation.

Biostratigraphy:

Age: Maastrichtian (perhaps late Maastrichtian?) Stage, Uppermost Senonian Subepoch, Uppermost Gulf Epoch, Late Cretaceous.

Material: Skull, dentary, and anterior portion of postcranial skeleton including cervical and anterior dorsal vertebrae, cervical and anterior dorsal osteoderms, and complete pectorial girdle and forelimbs.

Simosuchus clarki (after Krause et al. 2019).

Referred material:

GAO, 1994

Locality: Mahajanga Basin, Northwestern Madagascar.

Horizon: “Maevarano formation”.

Biostratigraphy:

Age: Late Cretaceous.

Material:

Number: Not given: Osteoderms.
Note: Orginally believed to belong to the lizard family: CORDYLIDAE? But shown to belong to Simosuchus clarki.

 

KRAUSE, SERTICH, ROGERS, KAST, RASOAMIARAMANANA, & BUCKLEY, 2010 (published in 2011)

Locality: MAD 93-06, near the village of Berivotra, Mahajanga Basin, Mahajana Province, northeastern Madagascar.

Horizon: Anebalemba Member, Maevarano Formation.

Biostratigraphy:

Age: Maastrichtian Stage, Uppermost Senonian Subepoch, Uppermost Gulf Epoch, Late Cretaceous.

Material:

UA 9755: Isolated tooth.

Locality: MAD 93-14, near the village of Berivotra, Mahajanga Basin, Mahajana Province, northeastern Madagascar.

Horizon: Anebalemba Member, Maevarano Formation.

Biostratigraphy:

Age: Maastrichtian Stage, Uppermost Senonian Subepoch, Uppermost Gulf Epoch, Late Cretaceous.

Material:

UA 9756: Isolated tooth.

 

Locality: MAD 93-18, near the village of Berivotra, Mahajanga Basin, Mahajana Province, northeastern Madagascar.

Horizon: Anebalemba Member, Maevarano Formation.

Biostratigraphy:

Age: Maastrichtian Stage, Uppermost Senonian Subepoch, Uppermost Gulf Epoch, Late Cretaceous.

Material:

FMNH PR 2824: Nearly complete fifth" cervical vertebra.

UA 9753: Nearly complete right femur.

 

Locality: MAD 93-33, near the village of Berivotra, Mahajanga Basin, Mahajana Province, northeastern Madagascar.

Horizon: Anebalemba Member, Maevarano Formation.

Biostratigraphy:

Age: Maastrichtian Stage, Uppermost Senonian Subepoch, Uppermost Gulf Epoch, Late Cretaceous.

Material:

FMNH PR 2591, 2592: Isolated teeth.

 

Locality: MAD 93-34, near the village of Berivotra, Mahajanga Basin, Mahajana Province, northeastern Madagascar.

Horizon: Anebalemba Member, Maevarano Formation.

Biostratigraphy:

Age: Maastrichtian Stage, Uppermost Senonian Subepoch, Uppermost Gulf Epoch, Late Cretaceous.

Material:

UA 9757: Isolated tooth.

 

Locality: MAD 93-35, near the village of Berivotra, Mahajanga Basin, Mahajana Province, northeastern Madagascar.

Horizon: Anebalemba Member, Maevarano Formation.

Biostratigraphy:

Age: Maastrichtian Stage, Uppermost Senonian Subepoch, Uppermost Gulf Epoch, Late Cretaceous.

Material:

FMNH PR 2600: Isolated tooth.

 

Locality: MAD 93-73, near the village of Berivotra, Mahajanga Basin, Mahajana Province, northeastern Madagascar.

Horizon: Anebalemba Member, Maevarano Formation.

Biostratigraphy:

Age: Maastrichtian Stage, Uppermost Senonian Subepoch, Uppermost Gulf Epoch, Late Cretaceous.

Material:

UA 9758: Isolated tooth.

 

Locality: MAD 95-11, near the village of Berivotra, Mahajanga Basin, Mahajana Province, northeastern Madagascar.

Horizon: Anebalemba Member, Maevarano Formation.

Biostratigraphy:

Age: Maastrichtian Stage, Uppermost Senonian Subepoch, Uppermost Gulf Epoch, Late Cretaceous.

Material:

FMNH PR 2593: Isolated tooth.

 

Locality: MAD 96-01, near the village of Berivotra, Mahajanga Basin, Mahajana Province, northeastern Madagascar.

Horizon: Anebalemba Member, Maevarano Formation.

Biostratigraphy:

Age: Maastrichtian Stage, Uppermost Senonian Subepoch, Uppermost Gulf Epoch, Late Cretaceous.

Material:

UA 9759, 9760: Isolated teeth.

 

Locality: MAD 98-25, near the village of Berivotra, Mahajanga Basin, Mahajana Province, northeastern Madagascar.

Horizon: Anebalemba Member, Maevarano Formation.

Biostratigraphy:

Age: Maastrichtian Stage, Uppermost Senonian Subepoch, Uppermost Gulf Epoch, Late Cretaceous.

Material:

FMNH PR 2594: Isolated tooth.

 

Locality: MAD 03-03, near the village of Berivotra, Mahajanga Basin, Mahajana Province, northeastern Madagascar.

Horizon: Anebalemba Member, Maevarano Formation.

Biostratigraphy:

Age: Maastrichtian Stage, Uppermost Senonian Subepoch, Uppermost Gulf Epoch, Late Cretaceous.

Material:

UA 9762: Partial frontal and 36 partial osteoderms.

 

Locality: MAD 03-09, near the village of Berivotra, Mahajanga Basin, Mahajana Province, northeastern Madagascar.

Horizon: Anebalemba Member, Maevarano Formation.

Biostratigraphy:

Age: Maastrichtian Stage, Uppermost Senonian Subepoch, Uppermost Gulf Epoch, Late Cretaceous.

Material:

UA 9761: Isolated tooth.

Locality: MAD 05-36, near the village of Berivotra, Mahajanga Basin, Mahajana Province, northeastern Madagascar.

Horizon: Anebalemba Member, Maevarano Formation.

Biostratigraphy:

Age: Maastrichtian Stage, Uppermost Senonian Subepoch, Uppermost Gulf Epoch, Late Cretaceous.

Material:

FMNH PR 2597: Partial skeleton including articulated but somewhat distorted skull and lower jaw, parts of all the cervical vertebrae and at least six anterior-most dorsal vertebrae, approximately 13 dorsal ribs (both left and right) in addition to numerous associated rib fragments, partial right and left scapulae, right coracoid, partial left coracoid, right humerus, proximal right radius and ulna, partial left radius, left radiale, fragmentary appendicular elements, numerous associated osteoderms and osteoderm fragments.

 

Locality: MAD 05-37, near the village of Berivotra, Mahajanga Basin, Mahajana Province, northeastern Madagascar.

Horizon: Anebalemba Member, Maevarano Formation.

Biostratigraphy:

Age: Maastrichtian Stage, Uppermost Senonian Subepoch, Uppermost Gulf Epoch, Late Cretaceous.

Material:

FMNH PR 2598: Skeleton including portions of the skull and lower jaw (fragmentary right quadratojugal, angular, and surangular), ceentrum of second? dorsal vertebra and nearly complete caudal vertebral series (with several chevron bones and articulated, infused ribs on the most proximal vertebrae, fragmentary right coracoid, proximal right humerus, proximal left ulnare, partial right manus including MC-I-MC-IV with five associated phalanges, right and proximal left femora, right tibia, right calcaneum and astragalus, fragments of three metatarsals and six pedal phalanges, rib fragments, numerous articulated and associated appendicular (femora), dorsal and ventral truck and caudal ostoderms.

Note: Association of some elements not indirect articulation is equivocal because FMNH PR 2599 was found near by.

FMNH PR 2599: Right articulated calcaneum and astragalus and left cancaneum.

Note: Association of these elements is equivocal but is based on the size and state of preservation, they were found very near to FMNH PR 2598. But that specimen has a right calcaneum and astragalus in close association with the rest of the skeleton and appears to represent a slightly larger individual.

 

Locality: MAD 05-42, near the village of Berivotra, Mahajanga Basin, Mahajana Province, northeastern Madagascar.

Horizon: Anebalemba Member, Maevarano Formation.

Biostratigraphy:

Age: Maastrichtian Stage, Uppermost Senonian Subepoch, Uppermost Gulf Epoch, Late Cretaceous.

Material:

UA 9854: Isolated tooth.

 

Locality: MAD 03-17, Masiakakoho Study Area, Mahajanga Basin, Mahajana Province, northeastern Madagascar.

Horizon: Anebalemba Member, Maevarano Formation.

Biostratigraphy:

Age: Maastrichtian Stage, Uppermost Senonian Subepoch, Uppermost Gulf Epoch, Late Cretaceous.

Material:

UA 9776: Left distal tibia, five phalanges, partial centrum of first caudal vertebra, and numerous osteoderms and osteoderm fragments.

FMNH PR 2596: Nearly complete, articulated but poorly preserved skeleton including entire crushed skull and lower jaw, all 8 cervical vertebrae, and undetermined number of dorsal vertebrae (because of poor preservation and ostodermal cover), but at least the posterior -most 7, both sacral vertebrae, 12 caudal vertebrae, several fragmentary ribs, nearly complete but poorly preserved left forelimb preserving all elements except for distal carpal 4, pisiform, digit V, phalanges I-1 and I-2, II-2, II-3, III-4, and IV-2 adn IV-3, articulated distal right humerus and proximal radius and ulna, broken and distorted portions of both ilia, both ischia, and both pubes, proximal right femur, nearly complete but poorly preserved left hind limb preserving all elements except for distal tarsals, MT V, and phalanges III-4 and IV-2-IV-4, associated appendicular dorsal and ventral trunk, and caudal ostoderms.

UA 9754: Right ectoptrygoid, 2 complete osteoderms, and numerous osteoderm fragments.

 

Locality: MAD 05-59, Masiakakoho Study Area, Mahajanga Basin, Mahajana Province, northeastern Madagascar.

Horizon: Anebalemba Member, Maevarano Formation.

Biostratigraphy:

Age: Maastrichtian Stage, Uppermost Senonian Subepoch, Uppermost Gulf Epoch, Late Cretaceous.

Material:

FMNH PR 2595: Isolated tooth.