FURCULA in non-avian theropods


LEGION: ARCHOSAUROMORPHA HUENE, 1946

LEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

SUBLEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

PLESION: DINOSAURIA GARDNER, 1982

ORDER: COELOPHYSIA NOVA

SUBORDER: COELOPHYSOIDEA (NOPCSA, 1928) HOLTZ, 1994

Superfamily: PODOKESAUROIDEA Madsen & Welles, 2000

Incertae sedis

Genus: Coelophysis COPE, 1889
C. bauri (COPE, 1887) COPE, 1889

Genus: Segisaurus CAMP, 1936
S. halli CAMP, 1936

Subfamily: COELOPHYSINAE Nopcsa, 1928

Genus: Megapnosaurus IVIE, SLIPINSKI & WEGRZYNOWICZ, 2001
M. rhodesiensis (RAATH, 1969) IVIE, SLIPINSKI & WEGRZYNOWICZ, 2001 (Type)

Genus: Syntarsus RAATH, 1969
S. kayentakatae ROWE, 1989

COHORT: THEROPODA EMEND MARSH, 1881

Incertae sedis

Gen sp indet.

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: CERATOSAURIA NOVAS, 1992

ORDER: CERATOSAURIA MARSH, 1884

Superfamily: ABELISAUROIDEA Novas, 1989

Family: ABELISAURIDAE Bonaparte & Novas, 1985

Subfamily: CARNOTAURINAE Sereno, 1998

Genus: Carnotaurus BONAPARTE, 1985
C. sastrei BONPARTE, 1985

ORDER: SPINOSAURIA OLSHEVSKY, 1991

Family: BARYONYCHIDAE Charig & Milner, 1986

Genus: Baryonyx CHARIG & MILNER, 1986
B. tenerensis (SERENO, BECK, DUTHEIL, GADO, LARSSON, LYON, MARCOT, RAUHUT, SADLEIR, SIDOR VARRICCHIO, WILSON & WILSON, 1998) SUES, FREY, MARTILL & SCOTT, 2002

ORDER: NEOTETANURAE SERENO, WILSON, LARSSON, DUTHEIL & SUES, 1994

SUBORDER: AVETHEROPODA PAUL, 1988

Superfamily: ALLOSAUROIDEA (Marsh, 1878) Currie & Zhao, 1994

Family?

Genus: Aerosteon SERENO, MARTINEZ, WILSON, VARRICCHIO, ALCOBER & LARSSON, 2008
A. riocoloradensis SERENO, MARTINEZ, WILSON, VARRICCHIO, ALCOBER & LARSSON, 2008 (Type)

Family: CARCHARODONTOSAURIDAE Stromer, 1931

Subfamily: GIGANOTOSAURIDAE Coria & Saigado, 1995 emmend.
= Family: GIGANOTOSAURIDAE Coria & Saigado, 1995 emmend.

Genus: Mapusaurus CORIA & CURRIE, 2006
M. roseae CORIA & CURRIE, 2006 (Type)

Family: ALLOSAURIDAE Marsh, 1878

Genus: Allosaurus MARSH, 1877
A. fragilis MARSH, 1877

Genus: Nova DALLA VECCHIA, 2001

MICRORDER: COELUROSAURIA HUENE, 1914

ORDER: SEGNOSAURIA BARSBOLD & PERLE, 1980

Incertae sedis

Genus: Falcarius KIRKLAND, ZANNO, SAMPSON, CLARK & DEBILEUX, 2005
F. utahensis KIRKLAND, ZANNO, SAMPSON, CLARK & DEBILEUX, 2005

Genus: Neimongosaurus ZHANG, XU, ZHAO, SERENO, KUANG & TAN, 2001.
N. yangi ZHANG, XU, ZHAO, SERENO, KUANG & TAN, 2001

Family: SEGNOSAURIDAE Perle, 1979

Genus: Beipiaosaurus XU, TANG & WANG, 1999
B. inexpectus XU, TANG & WANG, 1999

ORDER: MANIRAPTORFORMES HOLTZ, 1996

SUBORDER: COELURIA MARSH, 1881

Family: COMPSOGNATHIDAE Cope, 1875

Genus: Huaxiagnathus HWANG, NORELL & JI & GAO, 2004
H. orientalis HWANG, NORELL & JI & GAO, 2004 (Type)

Genus: Scipionyx SASSO & SIGNORE, 1998
S. samniticus SASSO & SIGNORE, 1998 (Type)

Superfamily: TYRANNOSAUROIDEA (Osborn, 1905) Bonaparte ET AL, 1990

Family: TYRANNOSAURIDAE Osborn, 1906

Subfamily: TYRANNOSAURINAE Osborn, 1906

Tribe: ALBERTOSAURINI Olshevsky, Ford & Yamamoto, 1995

Genus: Albertosaurus OSBORN, 1905
A. sarcophagus OSBORN, 1905

Genus: Gorgosaurus LAMBE, 1914
G. libratus LAMBE, 1914
G. sp

Tribe: TYRANNOSAURINI Olshevsky, Ford & Yamamoto, 1995

Genus: Daspletosaurus RUSSELL, 1970
D. torosus RUSSELL, 1970
D. sp

Genus: Tyrannosaurus OSBORN, 1905
T. rex OSBORN, 1905

INFRAORDER: OVIRAPTOROSAURIA BARSBOLD, 1976

Incertae sedis

Genus: Protarchaeopteryx JI & JI, 1997
P. robusta JI & JI, 1997 (Type)

Family: CAUDIPTERIDAE Zhou & Wang, 2000
= AVIFILOPLUMA GAUTHIER & DE QUEIROZ, 2001 (
partim)

Genus: Caudipteryx JI, CURRIE, NORELL & SHU-AN, 1998
C. zoui JI, CURRIE, NORELL & SHU-AN, 1998 (Type)

Family: OVIRAPTORIDAE Barsbold, 1976

Subfamily: OVIRAPTORINAE Barsbold, 1981

Genus: Citipati CLARK, NORELL & BARSBOLD, 2001
C. osmolskae CLARK, NORELL & BARSBOLD, 2001 (Type)

Genus: Heyuannai LU, (2002) 2003
H. huangi LU, (2002) 2003

Genus: Oviraptor OSBORN, 1924
O. philoceratops OSBORN, 1924

Subfamily: INGENIINAE Barsbold, 1981

Genus: Ingenia BARSBOLD, 1981
I. yanshini BARSBOLD, 1981 (Type)

Genus: Khaan CLARK, NORELL & BARSBOLD, 2001
K. mckennae CLARK, NORELL & BARSBOLD, 2001 (Type)

SUBORDER: MANIRAPTORA GAUTHIER, 1986

INFRAORDER: PARVES SERENO, 1997

PARVORDER: DEINONYCHOSAURIA COLBERT & D. A. RUSSELL, 1969

Family: DROMAEOSAURIDAE D. A. Russell, 1069

Subfamily: MICROPTORINI Senter, Barsbold, Brooks & Burnham, 2004

Genus: Bambiraptor BURNHAM, DERSTLER, CURRIE, BAKKER, ZHOU & OSTROM, 2000
B. feinbergorum emend BURNHAM, DERSTLER, CURRIE, BAKKER, ZHOU & OSTROM, 2000 (Type)

Genus: Microraptor XU, ZHOU & WANG, 2000
M. zhaoianus XU, ZHOU & WANG, 2000 (Type)

Genus: Sinornithosaurus XU, WANG & WU, 1999
S. millenii XU, WANG & WU, 1999 (Type)

Subfamily: VELOCIRPATORINAE Barsbold, 1978

Genus: Velociraptor OSBORN, 1924
V. mongoliensis OSBORN, 1924

Subfamily: UNENLAGIINAE Bonparte, 1999

Genus: Buitreraptor MAKOVICKY, APESTEGUIA & AGNOLIN, 2005
B. gonzalezorum MAKOVICKY, APESTEGUIA & AGNOLIN, 2005 (Type)

Family: TROODONTIDAE Gilmore, 1924

Genus: Mei XU & NORELL, 2004
M. long XU & NORELL, 2004 (Type)

Genus: Sinornithoides RUSSELL & DONG, 1993
S. youngi RUSSELL & DONG, 1993 (Type)

Genus: Troodon LEIDY, 1856
T. formosus LEIDY, 1856

COHORT: ORNITHISCHIFORMES COOPER, 1985

SUPERORDER: PROSAUROPODAMORPHA Nova

ORDER: PROSAUROPODA HUENE, 1920 (emend)

SUBORDER: PLATEOSAURIA TORNIER, 1913

Family: MASSOSPONDYLIDAE Huene, 1914

Note the following dinosaurs have been attributed with furcula.

Massospondylus COOPER, 1981

Family: PLATEOSAURIDAE Marsh, 1895

Plateosaurus

Note: HUENE, 1926 found a furcula.

SUPERORDER: SAUROPODOMORPHA HUENE, 1932

GRANDORDER: BRONTOSAURIA OLSHEVSKY, 1991 (emend)

MICRORDER: SAUROPODA MARSH, 1878

ORDER: EUSAUROPODA

SUBORDER: NEOSAUROPODA BONAPARTE, 1986

INFRAORDER: HOMALOSAUROPIA JANENSCH, 1929

PARVORDER: DIPLODOCIMORPHA CALVO & SALGADO, 1995

Superfamily: DIPLODOCOIDEA Upchurch, 1995

Family: DIPLOCODIDAE Marsh, 1884

Note: Possible furcula have been identified in Diplodocus, HATCHER, 1901, 1903, HOLLAND 1906.

MAGNORDER: ORNITHISCHIA SEELEY, 1888

SUPERORDER: NEORNITHISCHIA COOPER, 1985

GRANDORDER: ORNITHOPODA MARSH, 1871

MICRORDER: MARGINOCEPHALIA SERENO, 1986

ORDER: CERATOPIA MARSH, 1890

SUBORDER: NEOCERATOPIA SERENO, 1986

Family: PSITTACOSAURIDAE Osborn, 1923

Genus: Psittacosaurus OSBORN, 1923
P. mongoliensis OSBORN, 1923
P. sinensis YOUNG, 1958

INFRAORDER: CORONOSAURIA SERENO, 1986

Family: PROTOCERATOPIDAE Granger & Gregory, 1923

Subfamily: PROTOCERATOPINAE Nesov, Kaznyshkina & Cherepanov, 1989

Genus: Protoceratops GRANGER & GREGORY, 1923
P. andrewsi GRANGER & GREGORY, 1923

Subfamily: LEPTOCERATOPINAE Nopcsa, 1923
= Subfamily: LEPTOCERATOPSINAE Nopcsa, 1923

Genus: Leptoceratops BROWN, 1914
L. gracilis BROWN, 1914

Bibliography