Species: ingens (MARSH, 1872) MARSH, 1876
Etymology:

Locality: North side of the Smoky (Hill) River, 18 miles (=30.0) km) east of Fort Wallace, 5 miles west of Russell Springs, Sec. 13 and 24, T12S, R36W, Logan County, Kansas.

Horizon: Smoky Hill Member, Niobrara Formation.

Biostratigraphy:

Age: Santonian or Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Referred material:

YPM 1175: Mandublar ramus, scapula, coracoid, humerus, ulna, several wing bones, including an ulna, a radius, a humerus, a 4th metacarpal, 4d, 4v, femur, a metatarsal and a tooth?
Note: The specimens were found at 4 different locations and were very probably the remains of 4
Note: Mandublar ramus has associated hyperplasia on the external surface of the left side due to necroses, right mandibular ramus has bony overgrowth (BENNETT, 2003).

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Species: longiceps MARSH, 1876
Etymology:

Referred material:

BENNETT, 2003

Locality:

Horizon: Niobrara Formation.

Biostratigraphy:

Age: Coniacian Stage, Lower Senonian Subepoch, Lower Gulf Epoch, Late Cretaceous.

FMNH UC 609: Femur and pedal phalanges.
Note: First pedal phalanx has healed fracture.

FMNH uncatalogued: 4th wing phalange.
Note: Healed fracture.

KUVP 49398: Wing phalange 2-4.
Note: Pathological area due to necroses.

KUVP 60617: Tibia.
Note: A large bony overgrowth on the anterolateral surface.

UM 12399: Mandible.
Note: Necrotic pit on the inferior margin of the left mandibular ramus.

YPM 1192: Tibia.
Note: A large bony overgrowth on the anterolateral surface.

YPM 2310: Ulna, metacarpal 4d, 4v.
Note: Metacarpal with a small oval area of pathological bone due to necroses.

YPM 2487: Mandibular ramus.
Note: Small overgrowth on the lingual side of the left mandibular ramus.

YPM 2591: 3rd and 4th phalanges of the 5th digit.
Note: 4th wing phalanx with a pathological lump due to necroses.

YPM 2727: Fragmentary specimen.
Note: Large area of pathological bone on the ventral side of a right first or second wing phalanx due to necroses.

YPM 2686: Badly deformed articular end.
Note: Osteorthritis.

YPM 2711: Pathology on the distal syncarpal.
Note: Osteorthritis.

 

Locality: Western part of Butte Creek, Logan County, Kansas.

Horizon: Niobrara Formation.

Biostratigraphy:

Age: Coniacian Stage, Lower Senonian Subepoch, Lower Gulf Epoch, Late Cretaceous.

Material:

LACM 50926: Fragmentary skull, ight radius, ulna, left scapula, coracoid humerus, radius, metacarpal 4, both metacarpal 4d, both wing phalange 1-2, wing phalange 3, femora and right tibia.
Note: Mandibular ramus is badly deformed due to necroses.

 

= Pteranodon (Longicepia) marshi MILLER, 1972
Etymology:

Holotype: YPM 2594

Locality: Near the Smoky (Hill) River, Wallace County, Kansas.

Horizon: Upper Smoky Hill Member, Niobrara Formation.

Biostratigraphy:

Age: Santonian or Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material: Skull, distal end of the quadrate bone of a male.
Note: Right marginal ridge of mandible shows extensive necrosis.