NAVALON, MARUGAN-LOBON, CHIAPPE, SANZ, & BUSCALIONI, 2015

Locality: Las Hoyas fossil site, La Cierva Township, 20 km east of Cuenca, in the Castellan Branch of the Iberian Range, Cuenca Province, Spain.

Horizon: Calizas de La Huerguina Formation.

Biostratigraphy:

Age: Upper Hauterivian Stage, Late Neocomian subepoch-Lower Barremian Stage, Early Gallic subepoch, Early Cretaceous Epoch, Early Cretaceous.

Material:

MCCMLH31444: Fragmentary wing; ulna, radius, metacarpals, completemanual digits, all surrounded by abundtant remains of epidermal and dermal connective tissues in close association with plumage, 8-9 fragmentary large and strongly asymmetrical primaries, 10-12 secondaries, and the remins of plumulaceous coverts are preserved as carbonized keratinous structures.