Species: serpentinus (COPE, 1871) WELLES, 1943
Etymology:
Referred material:
= Styxosaurus browni WELLES, 1952
Etymology:
Referred material:BROWN, 1904
Locality: Head of Hat Creek Basin, 18 miles, southwest of Edgemont, Fall River County, South Dakota.
Horizon: Sharon Springs Member, Pierre Shale.
Biostratigraphy:
Age: Upper Lower to Lower Middle Campanian Stage, Senonian Subepoch, Upper Gulf Epoch, Late Cretaceous
Note: Now believed to be from the Younger Pierre Shale (MARTIN, SCHUMACHER PARRIS & GRANDSTAFF, 1998)Material:
AMNH 5803: Fragmentary skeleton with gastroliths.
Note: Pteranodon sp found in the stomach region along with a teleost fish, Scaphites and gastroliths (CICIMURRI & EVERHART, 2001).