Genus: Cotylorhynchus STOVALL, 1937
Etymology:

Species: romeri STOVALL, 1937
Etymology: In honor of Alfred S. Romer.

Holotype: OUSM 4-0-S1

Locality: Ross Farm, 4 3/4 miles west of Navina, NW 1/4, SEC. 4, T15N, R4W, Logan County, Oklahoma.

Horizon: Hennessey Formation, (Arroyo equivalent), Low Clear Fork Group.

Biostratigraphy:

Age: Leonardian Stage, Lower Rotliegendes Epoch, Early Permian.

Material: Right side of a skull, part of an interclavicle and a right and left manus.

Referred material:

Locality: Pierce Farm, SE 1/4, NE 1/4, Sec. 17, T9N, R2W, Cleveland County, Oklahoma.

Horizon: Hennessey Formation, Early Permian.

Material:

OUSM 4-0-S2: Caudal vertebrae, pelvis and hind limb. (Mus. no. 1249)

OUSM 4-0-S8: Anterior half of skeleton, first 19 vertebrae, ribs, humeri and scapula.

OUSM 4-44-S1: 3 ribs.

 

Locality: Near Norman, Cleveland County, Oklahoma.

Horizon: Hennessey Formation, (Arroyo equivalent), Low Clear Fork Group.

Biostratigraphy:

Age: Leonardian Stage, Lower Rotliegendes Epoch, Early Permian.

Material:

OUSM 4-0-S3: Partial skeleton of a juvenile.

OUSM 4-0-S13: Partial skeleton.

OUSM 4-0-S16: Foot, radius, humerus, tibia, fibula, femur, vertebrae and pelvis.

OUSM 4-1-S2: Part of skull and lower jaw.

OUSM 4-1-S10: Skull and jaws.

OUSM 4-1-S1: Lumbar vertebrae.

OUSM 4-12-S1: Scapula, humerus, radius, phalanges, and foot.

OUSM 4-21 to 24-S1: Numbers given to scraps that probably pertain to other numbered specimens.

OUSM 4-32-S1: Fragments.

Mus. No. 1250, (OUSM No?): Skeleton.

FMNH PR 272, USNM 21317, AMNH 7517, MCZ 3416: Skeletons lacking skulls.

 


Locality: Polk Farm, NW 1/4, Sec. 16, T9N, R2W, Cleveland County, Oklahoma.

Horizon: Hennessey Formation, (Arroyo equivalent), Low Clear Fork Group.

Biostratigraphy:

Age: Leonardian Stage, Lower Rotliegendes Epoch, Early Permian.

Material:

OUSM 4-0-S4: 4 1/2 caudal vertebrae and pelvis.

OUSM 4-0-S6: Foot.

OUSM 4-0-S21 (Duplicate no): Ribs, humerus, right pes and manus, and part of a pelvis.

OUSM 4-0-S23: Skull.

OUSM 4-10-S2: Caudal vertebrae.

OUSM 4-10-S2: Vertebral column (these two maybe the same).

 


Locality: Boggs Farm, NE 1/4, SEC. 16, T9N, R2W, Cleveland County, Oklahoma.

Horizon: Hennessey Formation, (Arroyo equivalent), Low Clear Fork Group.

Biostratigraphy:

Age: Leonardian Stage, Lower Rotliegendes Epoch, Early Permian.

Material:

OUSM 4-0-S6: Fragmentary foot.

OUSM 4-0-10: Complete skeleton, left femur, tibia, pes missing.

OUSM 4-0-11: Foot, tarsals, metatarsals, phalanges, 10 ribs, fragments of vertebrae, femur, and pelvis.

 


Locality: SW 1/4, Sec. 7, T9N, R2W, Cleveland County Oklahoma.

Horizon: Hennessey Formation, (Arroyo equivalent), Low Clear Fork Group.

Biostratigraphy:

Age: Leonardian Stage, Lower Rotliegendes Epoch, Early Permian.

Material:

OUSM 4-0-S9: Miscellaneous fragments, ribs, etc.

 


Locality: Burton Farm, SW 1/4, Sec. 24, T8N, R2W, Cleveland County, Oklahoma.

Horizon: Hennessey Formation, (Arroyo equivalent), Low Clear Fork Group.

Biostratigraphy:

Age: Leonardian Stage, Lower Rotliegendes Epoch, Early Permian.

Material:

OUSM 4-0-S14: Ribs, and part of foot.

OUSM 4-0-S17: Fragments, mainly ribs.

 


Locality: Burgess Farm, SE 1/4, Sec. 14, T8N, R2W, Cleveland County, Oklahoma.

Horizon: Hennessey Formation, (Arroyo equivalent), Low Clear Fork Group.

Biostratigraphy:

Age: Leonardian Stage, Lower Rotliegendes Epoch, Early Permian.

Material:

OUSM 4-0-S18: Partial skeleton, no skull and hind limbs.

 

 

Locality: South of Cedar Lane Golf Course, SE 1/4, Sec. 15, T8N, R2W, Cleveland County, Oklahoma.

Horizon: Hennessey Formation, (Arroyo equivalent), Low Clear Fork Group.

Biostratigraphy:

Age: Leonardian Stage, Lower Rotliegendes Epoch, Early Permian.

Material:

OUSM 4-0-S20: Part of skull, shoulder girdle, left fore limb, and foot.

 

 

Locality: South Oklahoma City, Cleveland County, Oklahoma.

Horizon: Hennessey Formation, (Arroyo equivalent), Low Clear Fork Group.

Biostratigraphy:

Age: Leonardian Stage, Lower Rotliegendes Epoch, Early Permian.

Material:

OUSM 4-0-S21: Incomplete skeleton.

 


Locality: 3 miles north of Norman on Highway 77, Cleveland County, Oklahoma.

Horizon: Hennessey Formation, (Arroyo equivalent), Low Clear Fork Group.

Biostratigraphy:

Age: Leonardian Stage, Lower Rotliegendes Epoch, Early Permian.

Material:

OUSM 4-0-S24: Partial skeleton.

 


Locality: Richardson Farm, 2 1/2 east on highway 9, South of road, Sec. 34, T9N, R2W, Cleveland County, Oklahoma.

Horizon: Hennessey Formation, (Arroyo equivalent), Low Clear Fork Group.

Biostratigraphy:

Age: Leonardian Stage, Lower Rotliegendes Epoch, Early Permian.

Material:

OUSM 4-0-S26: Partial skeleton, no skull, end of tail, fragment of lower jaw.

 


Locality: Noble, 2 1/2 miles east of Highway 77, sec. 25, T8N, R2W, Cleveland County, Oklahoma.

Horizon: Hennessey Formation, (Arroyo equivalent), Low Clear Fork Group.

Biostratigraphy:

Age: Leonardian Stage, Lower Rotliegendes Epoch, Early Permian.

Material:

OUSM 4-0-S28: Fragments.

 


Locality: NE 1/4, Sec. 15, T8N, R2W, Cleveland County, Oklahoma.

Horizon: Hennessey Formation, (Arroyo equivalent), Low Clear Fork Group.

Biostratigraphy:

Age: Leonardian Stage, Lower Rotliegendes Epoch, Early Permian.

Material:

OUSM 4-1-S3: Indeterminate bones.

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Species: hancocki OLSON & BEERBOWER, 1953
Etymology:

Holotype: FMNH (CMNH) UR 154

Locality: From fine green sandstone near the top of upper third of section, sublocality HA, Pease River, Hardeman County, Texas.

Horizon: Upper part of San Angelo Formation.

Biostratigraphy:

Age: Early Late Permian.

Material: Right humerus and proximal end of right radius.

Referred material:

Locality: Locality KN, MacFayden Ranch south, little Croton Creek, Knox County, Texas.

Horizon: Middle San Angelo Formation.

Biostratigraphy:

Age: Early Late Permian.

Material:

FMNH (CMNH) UR 144: 4 ribs.

 

Locality: Locality KV, MacFayden Ranch north, Knox County, Texas.

Horizon: San Angelo Formation.

Biostratigraphy:

Age: Early Late Permian.

Material:

FMNH (CMNH) UR 249: Humerus and ulna.

FMNH (CMNH) UR 250: 6 vertebrae and ribs.

 


Locality: Locality KR, Alexander Ranch, Knox County, Texas.

Horizon: San Angelo Formation.

Biostratigraphy:

Age: Early Late Permian.

Material:

FMNH (CMNH) UR 266: Parts of ribs, distal end of humerus.

 

 

Locality: Locality KX, Driver Ranch, Knox County, Texas.

Horizon: San Angelo Formation.

Biostratigraphy:

Age: Early Late Permian.

Material:

FMNH (CMNH) UR 479: 3rd or 4th presacral vertebra, 4 dorsal vertebrae and 9 caudals, many parts of ribs, partial femur, clavicle and scapulocoracoid.

 


Locality: Locality KY, Driver Ranch, Knox County, Texas.

Horizon: San Angelo Formation.

Biostratigraphy:

Age: Early Late Permian.

Material:

FMNH (CMNH) UR 480: Ischium, parts of pelvis, ribs, fragments of vertebrae from a young individual.

FMNH (CMNH) UR 481: Distal end of humerus, lumbar vertebrae and fragments.

FMNH (CMNH) UR 563: Humerus, ribs and fragmentary pelvis.

 


Locality: Locality KAB, Driver Ranch, Knox County, Texas.

Horizon: San Angelo Formation.

Biostratigraphy:

Age: Early Late Permian.

Material:

FMNH (CMNH) UR 483: Several partial caudal vertebrae.

 

 

Locality: Locality KAD, Driver Ranch, Knox County, Texas.

Horizon: San Angelo Formation.

Biostratigraphy:

Age: Early Late Permian.

Material:

FMNH (CMNH) UR 582: Fragmentary pelvis.

 


Locality: Locality KAC, Kahn Quarry, Driver Ranch, Knox County Texas.

Horizon: San Angelo Formation.

Biostratigraphy:

Age: Early Late Permian.

Material:

FMNH (CMNH) UR 487: 2 scapulocoracoids.

FMNH (CMNH) UR 488: 2 humeri and 1 femur.

FMNH (CMNH) UR 489, UR 490: Fragmentary skulls.

FMNH (CMNH) UR 491, UR 492, UR 493, UR 494: Fragmentary lower jaws with teeth.

FMNH (CMNH) UR 498: Fragmentary skull, maxillary, postnasal bar, part of the naris dentition.

FMNH (CMNH) UR 500: Fragmentary lower jaw.

FMNH (CMNH) UR 504: Caudal vertebrae.

FMNH (CMNH) UR 506: Shaft of humerus.

FMNH (CMNH) UR 564: 2 dorsal vertebrae.

FMNH (CMNH) UR 565: 5 vertebrae and limb bones.

FMNH (CMNH) UR 567: Right ulna.

FMNH (CMNH) UR 568: Caudal vertebrae.

FMNH (CMNH) UR 569: Dorsal vertebra.

FMNH (CMNH) UR 571: 3rd or 4th presacral vertebra, 1 caudal vertebra, ulna, fragmentary femur, and ribs.

FMNH (CMNH) UR 580: Sacral and parts of 4 posterior presacral vertebrae, and sacral ribs.

FMNH (CMNH) UR 581: Nearly complete skeleton lacking some cervical vertebrae, 1 scapulocoracoid and some limb bones.

FMNH (CMNH) UR 585, UR 623: Ulna.

FMNH (CMNH) UR 586: Tibia, and fragmentary tibia.

FMNH (CMNH) UR 621: Partial skeleton and part of skull, lacking girdles, some limb bones and some vertebrae.

FMNH (CMNH) UR 622: Partial skeleton and part of brain-case and palate of skull, various vertebrae, ribs, limb bones, clavicle, and foot bones.

FMNH (CMNH) UR 624: Fragments of skull and dentition.

FMNH (CMNH) UR 625: Ungual phalanges.

FMNH (CMNH) UR 626: Caudal vertebrae and limb bone.

FMNH (CMNH) UR 703: Part skeleton of very large individual, vertebrae, caudal, sacral, lumbar, dorsal vertebrae, pelvis, femur, radius, ulna, ribs.

FMNH (CMNH) UR 704: Terminal phalanx.

FMNH (CMNH) UR 705, UR 706: Dorsal vertebrae.

FMNH (CMNH) UR 707: Tibia.

FMNH (CMNH) UR 708: Interclavicale.

FMNH (CMNH) UR 709, UR 710: Vertebrae.

FMNH (CMNH) UR 718" Vertebrae and toe bones.

FMNH (CMNH) UR 719: Cervical vertebra and ribs.

FMNH (CMNH) UR 720: Vertebrae, ribs, and clavicle.

FMNH (CMNH) UR 821: Vertebrae and ribs.

FMNH (CMNH) UR 822: Humerus and scapulocoracoid of an immature individual.

FMNH (CMNH) UR 823: Humerus of a juvenile.

 


OLSON, 1968

Locality: Kahn Quarry, Knox County, Texas.

Horizon: San Angelo Formation.

Biostratigraphy:

Age: Early Late Permian.

Material:

FMNH UR 875: Pelvis.

FMNH UR 877: A clavicle, rib, and cervical vertebrae.

FMNH UR 878: Femur and fibula.

FMNH UR 879: Humerus, clavicle, rib radius, 8 vertebrae, and a chevron.

FMNH UR 881: Tibia.

FMNH UR 892: Scapulocoracoid, and 2 vertebrae.

FMNH UR 893: Tibia and sacral vertebrae.

FMNH UR 894: Partial pelvis, 2 sacral and 1 caudal.

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Species: bransoni OLSON & BARGHUSEN, 1962
Etymology:

Holotype: FMNH UR 835

Locality: Site KF-1, Kingfisher County, Oklahoma.

Horizon: Middle part of Flowerpot Formation.

Biostratigraphy:

Age: Early Late Permian.

Material: Left side of pelvis, left femur and partial sacral rib.

Referred material:

FMNH (CMNH) UR 836: Right tibia, right fibula, tarsus, metatarsus, phalanges except for unguals.

FMNH (CMNH) UR 837: Left radius and ulna and part of carpus.

FMNH (CMNH) UR 838: Flattened left astragalus, (Reference tentative.)

FMNH (CMNH) UR 839: Immature left tibia.

FMNH (CMNH) UR 840: Poorly preserved left fibula of somewhat immature individual.

FMNH (CMNH) UR 841: Part of left maxilla with 2 cheek teeth.

FMNH (CMNH) UR 842: 2 fragments of an ungual phalanx.

FMNH (CMNH) UR 843: Ungual phalanx.

 


OLSON, 1965

Locality: Omega quarry, site KF-1, NW 1/4, NW 1/4, NW 1/4, sec. 19, T17N, R9W, Kingfisher County, Oklahoma.

Horizon: Chickasha Formation.

Biostratigraphy:

Age: Guadalupian subepoch, Lower Zechstein Epoch, Early Late Permian.

Material:

FMNH (CMNH) UR 905: Part of foot.

FMNH (CMNH) UR 910: Cervical ribs.

FMNH (CMNH) UR 912: Clavicle.

FMNH (CMNH) UR 913: Chevron.

FMNH (CMNH) UR 915: Series of vertebrae.

FMNH (CMNH) UR 918: Scapulocoracoid.

FMNH (CMNH) UR 919: Scapulocoracoid and humerus.

FMNH (CMNH) UR 923: Sacral vertebrae.

FMNH (CMNH) UR 937: Caudal vertebrae.

 


Locality: BC-6, NW 1/4 sec. 35, Sec. 34, S 1/2 Sec. 27, T18N, R11W, Blaine County, Oklahoma.

Horizon: Chickasha Formation.

Biostratigraphy:

Age: Guadalupian subepoch, Lower Zechstein Epoch, Early Late Permian.

Material:

FMNH (CMNH) UR 972: Caudal vertebrae.

 


Locality: BC-2, SW 1/4, SW 1/4, sec. 31, T18N, R10W, Blaine County, Oklahoma.

Horizon: Chickasha Formation.

Biostratigraphy:

Age: Guadalupian subepoch, Lower Zechstein Epoch, Early Late Permian.

Material:

FMNH (CMNH) UR 983: Dorsal vertebrae.

 


Locality: BC-7, SW 1/4, NW 1/4, sec. 34, NE 1/4, sec. 33, S 1/2 sec. 38, T18N, R11W, Blaine County, Oklahoma.

Horizon: Chickasha Formation.

Biostratigraphy:

Age: Guadalupian subepoch, Lower Zechstein Epoch, Early Late Permian.

Material:

FMNH (CMNH) UR 984: Part of humerus.

FMNH (CMNH) UR 988: Base of pelvis.