Species: haplocerus (COPE, 1892) CASE, 1920 (nomen dubium)
Etymology:
= Episcoposaurus haplocerus COPE, 1892
Holotype: ANSP 14688
Locality: Near windmill in top pasture 3 miles (5 km) north of Dickens, Dickens County, Texas.
Horizon: Tecovas Member, Dockum Formation.
Biostratigraphy:
Age: Carnian Stage, Late Triassic Epoch, Late Triassic.
Material: A sacral and 2 caudal vertebrae, right scapula, ribs and about 30 dermal scutes.
Lectotype:
ANSP 14688: Conjoined cervical right lateral and paramedian scutes.
Paralectotypes:
ANSP 14688 (remander): Pelvic girdle and many paramedian and lateral scutes.
Topotypes:
TMM 18569: Dorsal vertebra and 2 lateral scutes.
Referred material:
HUNT & LUCAS, 1995
Locality: Canada Colorado, NMMNH locality 149, NW 1/4, SW 1/4, NW 1/4, SE 1/4, Sec. 2, T14N, R10E, 4 km east of Lamy, Santa Fe County, New Mexico.
Horizon: Los Esteros Member, Santa Rosa Formation.
Biostratigraphy:
Age: Adamanian, Latest Carnian Stage, Late Triassic Epoch, Late Triassic.
Material:
Number: Not given: Several partial skeletons and many isolated elements.
NMMNH P-3023, 25651: large pectoral horns.
Locality: NMMNH locality 1271, NW 1/4, NW 1/4, SE 1/4, sec. 34, T15N, R10E, Santa Fe County, New Mexico.
Horizon: Los Esteros Member, Santa Rosa Formation.
Biostratigraphy:
Age: Adamanian, Latest Carnian Stage, Late Triassic Epoch, Late Triassic.
Material:
Number: Not given:
PARKER & IRIMIS, 2005
Locality: PFV 202, Blue Forest Area, Petrified Forest National Monument, Apache County, Arizona.
Horizon: Blue Mesa Member, Petrified Forest Member, Chinle Group.
Biostratigraphy:
Age: Latest Carnian Stage, Late Triassic Epoch, Late Triassic.
Material:
PEFO 23338: Fragmentary paramedian plate.
Locality: PFV 212, Blue Forest Area, Petrified Forest National Monument, Apache County, Arizona.
Horizon: Blue Mesa Member, Petrified Forest Member, Chinle Group.
Biostratigraphy:
Age: Latest Carnian Stage, Late Triassic Epoch, Late Triassic.
Material:
PEFO 26668: Fragmentary paramedian plate.
Locality: PFV 198, Blue Forest Area, Petrified Forest National Monument, Apache County, Arizona.
Horizon: Blue Mesa Member, Petrified Forest Member, Chinle Group.
Biostratigraphy:
Age: Latest Carnian Stage, Late Triassic Epoch, Late Triassic.
Material:
PEFO 31177: Fragmentary lateral plate.
___________________________________________________________________________________
Species: spurensis CASE, 1920
Etymology:
Holotype: UMMP No. 7476
Locality: Near the east bank of Blanco (White) or Catfish River, about 1/2 mile (0.8 km) east of the crossing of the old mail road from Spur to Crosbyton, in Crosby County, Texas.
Horizon: Dockum Formation.
Biostratigraphy:
Age: Adamanian, Early to Middle Noriain Stage, Late Triassic Epoch, Late Triassic.
Material: Skull with premaxilla missing, endocranial cast, cervical, dorsal, sacral and anterior vertebrae, and dorsal armor.
Old reconstruction.
Referred material:
UMMP 7504: Bones of second individual found with the above individual.
UMMP 7331: Lateral plates.
Locality: In the breaks of Sand Creek, Crosby County, Texas.Horizon: Dockum Formation.
Biostratigraphy:
Age: Carnian-?Rhaetian Stage, Late Triassic Epoch, Late Triassic.
Material:
UMMP 7322: Right ilium.
Locality: Near the head of Holmes Creek, Crosby County, Texas.Horizon: Dockum Formation.
Biostratigraphy:
Age: Carnian-?Rhaetian Stage, Late Triassic Epoch, Late Triassic.
Material:
UMMP 7470: Ilium, pubes and some of the ischia.
Locality: Saurian Valley, PEFO 085002 (UCMP V82251, PF97) (part of UCMP V7034), NW 1/4 of SE 1/4, Sec. 15, and NE 1/4 of SW 1/4, Sec. 15. T 19N, R 23E, Lithodendron Wash Area, 6 to 8 m below? Sonsela Sandstone, Devil’s Playground Area, Petrified Forest, Apache County, Arizona.Horizon: Lower Petrified Forest Formation, Chinle Group.
Biostratigraphy:
Age: Latest Carnian Stage, Late Triassic Epoch, Late Triassic.
Material:
UCMP 126950: 3 pieces of paramedian plates.
Locality: Findaly's Fourtune. (MNA Loc. 302-111), NE 1/4 of NE 1/4, Sec. 15, T 19N, R 23 E, Lithodendron Wash Area, Petrified Forest, Apache County, Arizona.Horizon: Chinle Group.
Biostratigraphy:
Age: Carnian Stage, Late Triassic Epoch, Late Triassic.
Material:
MNA PL. 3172: A partial paramedian scute.
Locality: Jasper Forest South, PEFO 085070 (UCMP V82041, MNA 302:42, SMU 232, PF167), on line between SE 1/4 and NE 1/4 of SE 1/4, SEC 8, T 17N, R 24E, Jasper Forest Area, Petrified Forest, Apache County, Arizona.Horizon: Lower Petrified Forest Formation, Chinle Group.
Biostratigraphy:
Age: Latest Carnian Stage, Late Triassic Epoch, Late Triassic.
Material:
UCMP 126885: 2 portions of a poorly preserved pectoral horn.
Locality: Delany Tank Northeast, (MNA Loc. 302, no specific locality number), Sec. 2, T 17N, R 23E, Delany Tank Area, Petrified Forest, Apache County, Arizona.Horizon: Lower Petrified Forest Formation, Chinle Group..
Biostratigraphy:
Age: Latest Carnian Stage, Late Triassic Epoch, Late Triassic.
Material:
MNA Pl. 697: Left lateral scute, cervical region.
Locality: Crystal Forest, PEFO 085077 (UCMP V82238, PF172) (part of ucmp v7037), log level, just below Sonseal Sandstone, NE 1/4 of NW 1/4, Sec 22, T 17N, R 14E, Crystal Forest Area, Petrified Forest, Apache County, Arizona.Horizon: Uppermost Blue Mesa Member, Petrified Forest Formation, Chinle Group.
Biostratigraphy:
Age: Late Carnian Stage, Lower Late Triassic Epoch, Late Triassic.
Material:
UCMP 126863, 126838, 126837, 126901, 126905: 5 lots consisting of 20 fragments of paramedian scutes.
Locality: Longs Logs, (UCMP Loc. V82024), Near center of SE 1/4 Sec. 1, T 16N, R 23 E, Longs Logs Area, Petrified Forest, Apache County, Arizona.
Horizon: Lower Petrified Forest Formation, Chinle Group.
Biostratigraphy:
Age: Latest Carnian Stage, Late Triassic Epoch, Late Triassic.
Material:
UCMP 126976: Fragment of paramedian scute.
Locality: Ward’s Bonebed (UCMP V4121, V4150, V3309, V7317, etc), 5 km northeast to 10 km southeast of Cameron, Near Tanner's Crossing, Cameron Area, Petrified Forest, Coconino County, Arizona.Horizon: Lower Petrified Forest Formation, Chinle Group.
Biostratigraphy:
Age: Latest Carnian Stage, Late Triassic Epoch, Late Triassic.
Material:
Number: Not given: Incomplete pectoral spike.
Locality: Rincon Basin E (UCMP), 16 km northeast of Winslow on eastern margin of Rincon Basin, Winslow Area, Petrified Forest, Navajo County, Arizona.Horizon: Lower Petrified Forest Formation, Chinle Group.
Biostratigraphy:
Age: Latest Carnian Stage, Late Triassic Epoch, Late Triassic.
Material:Number: Not given: A small paramedian scute.
Locality: Near head of Home Creek/Swenson Ranch area, Crosby County, Texas.Horizon: Dockum Formation, Carnian-?Rhaetian, Late Triassic.
Material:
UCMP 85103: Pectorical spike.
UCMP 85106: Pectorical spike.
UT 31172-14: Pectorical Spike.
Locality: Cedar Mountain Area, (TMM Locality 31173, UCMP), 13 to 27 km southeast of Crosbyton, Crosby County, Texas.Horizon: Tecovas Member, Dockum Formation.
Biostratigraphy:Age: Carnian Stage, Late Triassic Epoch, Late Triassic.
Material:
UCMP 85112: Scute.
UCMP 102345: Scute.
UCMP 102343: Scute.
UT Collection: Scute.
Locality: North of Cedar Mountain, Dirty Tank, Crosby County, Texas.Horizon: Tecovas Member, Dockum Formation.
Biostratigraphy:Age: Carnian Stage, Late Triassic Epoch, Late Triassic.
Material:
UCMP 102343: Paramedian scute.
Locality: Palo Dura Canyon, Randall County, Texas.
Horizon: Dockum Formation.
Biostratigraphy:
Age: Carnian-?Rhaetian Stage, Late Triassic Epoch, Late Triassic.
Material:
Number: Not given: Pectoral spikes.
Locality: Herring Ranch/Rotten Hill Locality, Sierrita de la Cruz Creek, Potter County, Texas.
Horizon: Tecovas Member, Dockum Formation.
Biostratigraphy:Age: Carnian Stage, Late Triassic Epoch, Late Triassic.
Material:
UMMP V13026, V13637, V13727: Armor plates.
UMMP 13026: Lateral spikes.
UMMP 13727: Paramedian plate.
HECKERT, RINEHART, LUCAS, DOWNS, ESTEP, HARRIS, RESER, & SNYDER, 1999, ZEIGLER, HECKERT & LUCAS, 2000Locality: NMMNH Locality L-3845, Snyder Quarry, near Ghost Ranch, Rio Arriba County, in north-central New Mexico.
Horizon: Petrified Forest Formation, Chinle Group.
Biostratigraphy:
Age: Revueltian, Early Norian Substage, Early Norian Epoch, Middle Late Triassic Epoch, Late Triassic.
Material:
NMMNH P-29045: Right lateral fourth cervical scute.
Number: Not given: Numerous scutes.
HUNT & LUCAS 1989
Locality: Lamy Amphibian Quarry, SW 1/4, SW 1/4, NE 1/4, NE 1/4, Sec. 29, T 12N, R 11E, Santa Fe County, New Mexico.
Horizon: Lower Petrified Forest Formation, Chinle Group..
Biostratigraphy:
Age: Latest Carnian Stage, Late Triassic Epoch, Late Triassic.
Material:
Number: Not given:
LONG & MURRY, 1995
Locality: Placerias quarry (UCMP A269, MNA 207C), Big Hollow Wash near Romero Springs, a small valley 6 1/2 miles (approximately 10.4 km) S 65"W of St. Johns, Apache County, Arizona.
Note: Located 0.3 miles northwest of Romero Springs at approximately Long. 109'28'W and lat. 34"27'N on the USGS St. Johns quadrangle of 1921.Horizon: Bluewater Creek Formation, Lower Chinle Group.
Biostratigraphy:
Age: Adamanian, Latest Carnian Stage, Late Triassic Epoch, Late Triassic.
Material:
UCMP collection by 620 specimens form Placerias Quarry. Specimens include.UCMP 78748: Right cervical parameidan plates.
UCMP 78744, 78745, 78746: Cervical parameidan plates.
UCMP 78747: Anterior left dorsal paramedian plate.
UCMP 27408: Well-preserved occiput and basicranial portion of skull.
UCMP 27407: Posterior portion of skull.
UCMP 27410: Poorly-preserved skull.
UCMP 27406: Posterior portion of skull, probably same individual as UCMP 28345 and 27409.
UCMP 27419, 27417, 27423, 27415: Basicrania.
UCMP 27420: Basicranium with short basipterygoid process.
UCMP 27418: Basicranium with large basipterygoid process.
UCMP 27409: Parietals.
UCMP 27433: Right parietal and opisthoitic.
UCMP 27894, 27476: Left paritals.
UCMP 27477: Left parietal and fragment of right frontal.
UCMP 27482: Right parietal.
UCMP 27346, 27342: Right frontals.
UCMP 27341: Well-preserved left frontal.
UCMP 27345: Left frontal.
UCMP 27343: Left frontal with portion of left parietal.
UCMP 27344: Left frontal, articulates well with UCMP 27408.
UCMP 27468: Complete left quadrate.
UCMP 27574: Distal extremity of right quadrate.
UCMP 27471, 27466, 27467, 27473: Distal extremities of left quadrates.
UCMP 78695: Well-preserved left maxilla.
UCMP 78696, 78699: Left maxillae.
UCMP 27576: Articular/surangular.
UCMP 27575: Left articular/surangular.
UCMP 27576: Left articular/surangular.
UCMP 27961: Right scapulocoracoid missing scapular blade.
UCMP 32199: Left sacpulocoracoid missing scapular blade and most coracoid belongs to same individual as UCMP 27981.
UCMP 27981: Left scapular blade.
UCMP 37216: Right scapulocoracoid missing part of scapular blade and most of coracoid.
UCMP 32214: Left scapular blade.
UCMP 32211: Badly crushed right scapulocoraocoid, same individual as UCMP 32180 and 28349.
UCMP 32145, 32200: Right coracoids.
UCMP 32207: Left scapula.
UCMP 32215, 32158: Left scapulocoracoids.
UCMP 27964, 27965, 27968: Right scapulocoracoids.
UCMP 32181: Left humerus missing proximal extremity.
UCMP 32175: Right humerus missing proximal extremity.
UCMP 32168: Complete right humerus.
UCMP 32190: Proximal half of left humerus.
UCMP 32179: Complete left humerus with crushed distal extremity.
UCMP 32172: Distal extremity of right humerus.
UCMP 28351: Left humerus, probably same individual as UCMP 37216.
UCMP 32177: Right humerus shaft.
UCMP 32184: Right humerus, probably same individual as UCMP 27961.
UCMP 32188: Proximal extremity of humerus.
UCMP 32189: Proximal extremity of left humerus.
UCMP 32178: Proximal extremity of right humerus.
UCMP 32182: Proximal fragment of right humerus.
UCMP 32188: Distal extremity of right humerus.
UCMP 32170: Distal extremity of right humerus.
UCMP 32180: Proximal extremity of right humerus, same individual as UCMP 23849.
UCMP 32174: Distal extremity of left humerus.
UCMP 28349: Distal extremity of right humerus.
UCMP 25989: Well-preserved right ilium.
UCMP 25987: Badly crushed right ilium, same individual as UCMP 25988:
UCMP 25988: Badly crushed left ilium.
UCMP 25954: Acetabular region of right ilium.
UCMP 25070: Posterior blade of right ilium.
UCMP 25003: Fragmentary posterior blade of left ilium.
UCMP 32166: Right pubis, probably same individual as UCMP 32166.
UCMP 25955, 32147, 32162, 32165: Left pubes.
UCMP 25951: Well-preserved left pubis, probably same individual as UCMP 25495.
UCMP 32138, 25964: Right pubes.
UCMP 25945: Well-preserved right pubis.
UCMP 25975, 32157: Left ischia.
UCMP 27988: Well-preserved right femur, only lacking lateral condyle.
UCMP 25926: Proximal half of left femur, same individual as UCMP 25904.
UCMP 25904: Distal half of left femur, probably same individual as UCMP 27988.
UCMP 25910: Left femur, probably same individual as UCMP 25911.
UCMP 25916, 25911: Left femora.
UCMP 25874: Right tibia with distorted shaft, probably same individual as UCMP 25877.
UCMP 25877: Left tibia with some reconstruction.
UCMP 25886: Right tibia with crushed proximal extremity.
UCMP 25889: Proximal extremity of left tibia, same individual as UCMP 25895.
UCMP 25895: Distal extremity of left tibia.
UCMP 25882: Proximal extremity of left tibia.
UCMP 25883: Distal extremity of left tibia.
UCMP 25891: Distal extremity of right tibia.
UCMP 25878: Badly crushed proximal extremity of left tibia of a very large individual, probably same individual as UCMP 25881 and 25852.
UCMP 25881: Distal extremity of left tibia.
UCMP 25850: Well-preserved left fibula.
UCMP 25867: Complete left fibula.
UCMP 32382: Complete right fibula.
UCMP 25859: Proximal extremity of left fibula.
UCMP 25861: Proximal extremity of left fibula.
UCMP25863: Proximal extremity of right fibula.
UCMP 25852: Distal extremity of left fibula.
UCMP 27992: Proximal extremity of left fibula.
UCMP 25857: Distal extremity of left fibula.
UCMP 34438, 34490, 34489: Right astragali.
UCMP 34492: Right astragalus, probably same individual as UCMP 25883.
?UCMP 34486: Small right astragalus.
UCMP 34494: Left atragalus, probably same individual as UCMP 25878.
UCMP 34493, 34491: Left astagali.
UCMP 34484: Left calcaneum missing “lip”.
UCMP 34481, 34480: Right calcanei.
UCMP 34483: Well-preserved right calcaneum.
UCMP 34478: Right calcaneum, same individual as UCMP 34490 and 25883.
The MNA collection includes the following specimens.
MNA Pl. 2831: Fifth cervical paramedian plate.
MNA Pl. 2934: Cervical paramedian plate.
MNA Pl. 2933: Posterior cervical or anterior dorsal paramedian plate.
?MNA Pl. 2858: Left paramedian plate.
MNA Pl. 2773, 2774, 2783, 2791, 2807, 2809, 2812, 2821, 2883, 2890, 2900, 2901, 2907, 2909, 2910, 2912, 2914, 2915, 2932, 2979, 2982: Paramedian plates.
MNA Pl. 2884: Paramedian plate fragments.
MNA Pl. 2974, 3004: Cervical lateral plates.
MNA Pl. 2754, 2755, 2756, 2757, 2758, 2759, 3742, 2813, 2827, 2850, 2874, 2889, 2891, 2913, 2938, 2940, 2958: Lateral plates.
MNA Pl. 2792: Lateral plate fragment.
MNA Pl. 2892: Cervical plate.
MNA Pl. 2842, 2944: Caudal lateral plates.
MNA Pl. 2885, 2925, 2967: Caudal plates.
MNA Pl. 2888: Plate.
MNA Pl. 2793: Plate fragment.
MNA Pl. 2717: Right cervical spine, fourth or fifth series.
MNA Pl. 2845: Spike from fifth lateral plate.
MNA V3633: Braincase.
MNA V3658: Left dentary.
MNA V3657: Dentary.
MNA Pl. 3034: Right articular and surangular.
MNA Pl. 2936: Axis vertebra.
MNA Pl. 2761, 2893, 2894, 2895, 2911, 2926, 2972, 2973, 2978, 2987, 2988, 2994: Cervical vertebrae.
?MNA Pl. 2762: Sacral rib.
MNA Pl. 2719, 2948: Anterial caudal vertebrae.
MNA V3745: Mid-caudal vertebra.
MNA Pl. 2796, 2816, 2963, 2964, 3036: Distal caudal vertebrae.
MNA Pl. 2899: Scapula.
MNA Pl. 2786: Right humerus.MNA Pl. 2785, 2831: Right pubes.
MNA Pl. 2760: Proximal portion of left tibia.
?MNA Pl. 2772: Left fibula.
MNA Pl. 2983: Phalange.
Locality: Downs Quarry (MNA Loc. 207N), approximately 72 m east of Placerias Quarry, Big Hollow Wash near Romero Spring, approximately 10.4 km southwest of St. Johns, Apache County, Arizona.Horizon: Bluewater Creek Formation, Lower Chinle Group.
Biostratigraphy:
Age: Adamanian, Latest Carnian Stage, Late Triassic Epoch, Late Triassic.
Material:
?MNA Pl. 2710: Cervical vertebra.
MNA Pl. 3096: Lateral plate.
MNA Pl. 3110: Lateral scute.
MNA Pl. 3127: Dermal plate.
Locality: Big Hollow Wash, (UCMP V80003), St. Johns area, Apache County, Arizona.Horizon: Lower Petrified Forest Formation, Chinle Group.
Biostratigraphy:
Age: Latest Carnian Stage, Late Triassic Epoch, Late Triassic.
Material:
UCMP: Fragments of armor, limb bones, and vertebrae.
Locality: Tepees East, PEFO 085021 (UCMP V82257, PF118), NW 1/4 of NW 1/4, Sec. 24, T 18N, R 24E, in variegated siltstone bed, Blue Mesa W Area, Petrified Forest, Apache County, Arizona.Horizon: Lower Petrified Forest Formation, Chinle Group.
Biostratigraphy:
Age: Latest Carnian Stage, Late Triassic Epoch, Late Triassic.
Material:
UCMP: Caudal centrum.
Locality: Paulcell 6 (PF 145), Blue Mesa E Area, Petrified Forest, Apache County, Arizona.Horizon: Lower Petrified Forest Formation, Chinle Group.
Biostratigraphy:
Age: Latest Carnian Stage, Late Triassic Epoch, Late Triassic.
Material:
UCMP: Caudal vertebra.
Locality: Agate Bridge N (UCMP V82339, PF 161), NE 1/4 of SE 1/4 and SE 1/4 of NE 1/4, SEC. 4, T 17N, R 24E, near base of black/grey siltstone, approximately 6 m below Sonsela Sandstone, resting on a loose sandstone, Lot’s Wife Area, Petrified Forest, Apache County, Arizona.Horizon: Lower Petrified Forest Formation, Chinle Group.
Biostratigraphy:
Age: Latest Carnian Stage, Late Triassic Epoch, Late Triassic.
Material:
UCMP: Plate fragments.
Locality: Battleship Northwest, (UCMP Loc. V8261), on line between NE 1/4 of SE 1/4, Sec. 17, and NW 1/4 of SW 1/4, Sec. 16, T 17N, R 24E, Battleship Northwest Area, Petrified Forest, Apache County, Arizona.Horizon: Lower Petrified Forest Formation, Chinle Group.
Biostratigraphy:
Age: Latest Carnian Stage, Late Triassic Epoch, Late Triassic.
Material:
UCMP: Lateral horn and caudal centrum.
Locality: Allentown Claypit (MNA), Allentown Area, Petrified Forest, Apache County, Arizona.
Horizon: Lower Petrified Forest Formation, Chinle Group.
Biostratigraphy:
Age: Latest Carnian Stage, Late Triassic Epoch, Late Triassic.
Material:
MNA Pl. 716: Enormous pectoral horn.
MNA P;. 708: Lateral spine.
Locality: St. Michaels S (UA, MNA 227), MNA is approximately 2.8 km south and 5.2 km west of St. Michaels, UA is approximately 3 km south of St. Michaels, St. Michaels Area, Petrified Forest, Apache County, Arizona.
Horizon: Lower Petrified Forest Formation, Chinle Group.
Biostratigraphy:
Age: Latest Carnian Stage, Late Triassic Epoch, Late Triassic.
Material:
MNA Pl. 683: Complete right pectoral spike.
Note: Brady, 1958.MNA Pl. 686: Paramedian scute.
MNA Pl. 687: Left anterior proximal plate articulated with left lateral plate.
Note: Brady, 1958.MNA Pl. 700: Lateral plate.
MNA Pl. 701: Natural mold of base of pectoral spike.
Locality: Lupton W (MNA), Lupton Area, approximately 13 km west of Lupton, Petrified Forest, Apache County, Arizona.
Horizon: Lower Petrified Forest Formation, Chinle Group..
Biostratigraphy:
Age: Latest Carnian Stage, Late Triassic Epoch, Late Triassic.
Material:
MNA: Base of pectoral horn.
Note: BRADY, 1954.
Locality: Window Rock N (MNA), Window Rock Area, Petrified Forest, Apache County, Arizona.
Horizon: Lower Petrified Forest Formation, Chinle Group.
Biostratigraphy:
Age: Latest Carnian Stage, Late Triassic Epoch, Late Triassic.
Material:
MNA Pl. 684: Portions of 3 paramedian plates.
Locality: Fossil Garden, PEFO 085003 (UCMP V82252, PF98) (part of UCMP V7034), 6 to 8 m below? Sonsela Sandstone, Devil’s Playground Area, Petrified Forest, Apache County, Arizona.
Horizon: Lower Petrified Forest Formation, Chinle Group.
Biostratigraphy:
Age: Latest Carnian Stage, Late Triassic Epoch, Late Triassic.
Material:
UCMP 126955: Paramedian knob.
Locality: Saurian Valley N 1, PEFO 085004 (UCMP V85225, PF 99) (part of UCMP V7034), fauna at top of Sonsela Sandstone, Devil’s Playground area, Petrified Forest, Apache County, Arizona.Horizon: Lower Petrified Forest Formation, Chinle Group.
Biostratigraphy:
Age: Latest Carnian Stage, Late Triassic Epoch, Late Triassic.
Material:
UCMP: Fragment of paramedian plate.
Locality: Ward’s Bonebed (UCMP V4121, V4150, V3309, V7317, etc), 5 km northeast to 10 km southeast of Cameron, Near Tanner's Crossing, Cameron Area, Petrified Forest, Coconino County, Arizona.
Horizon: Lower Petrified Forest Formation, Chinle Group.
Biostratigraphy:
Age: Latest Carnian Stage, Late Triassic Epoch, Late Triassic.
Material:
AMNH 3060: Tibia and fibula found with enormous phytosaur (“Machaeroprosopus” gregori of Colbert, 1947 = Smilosuchus gregori).
AMNH 19344: Lateral horns with conjoined fragment of paramedian plate.
CM 9042: Paramedian plate.
UA: Pectoral spike.
Locality: Jim Camp Wash 3 (PF 84), Jim Camp Wash E Area, Petrified Forest, Apache County, Arizona.
Horizon: Lower Petrified Forest Formation, Chinle Group.
Biostratigraphy:
Age: Latest Carnian Stage, Late Triassic Epoch, Late Triassic.
Material:
PEFO 4856: Distal extremity of pectoral spike.
Locality: Canada Colorado, NMMNH locality 149, NW 1/4, SW 1/4, NW 1/4, SE 1/4, Sec. 2, T14N, R10E, 4 km east of Lamy, Santa Fe County, New Mexico.Horizon: Los Esteros Member, Santa Rosa Formation.
Biostratigraphy:
Age: Adamanian, Latest Carnian Stage, Late Triassic Epoch, Late Triassic.
Material:
NMMNH P18163: Very large pectoral horn.
NMMNH P3023: Fifth cervical horn.
NMMNH P3020: Cervical vertebra.
Locality: ?Revuelto Creek, Locality L00171, New Mexico.Horizon: Bull Canyon Formation.
Biostratigraphy:
Age: Revueltian, Early Norian Substage, Early Norian Epoch, Middle Late Triassic Epoch, Late Triassic.
Material:
NMMNH P4895: Pectoral spike.
NMMNH P4894: Small lateral spike.
Locality: Not given, New Mexico.Horizon: Bull Canyon Formation.
Biostratigraphy:
Age: Revueltian, Early Norian Substage, Early Norian Epoch, Middle Late Triassic Epoch, Late Triassic.
Material:
UCM 47225: Left pectoral spike.
2 uncatalogued pectoral spikes.
Locality: Near Otis Chalk, Howard County, Texas.Horizon: “Pre-Tecovas Horizon”, Dockum Group.
Biostratigraphy:
Age: Middle Carnian Stage, Lower Late Triassic Epoch, Late Triassic.
Material:
UMMP 11688: Pectoral spike.
Locality: TMM Site 1 (TMM Locality 31213), Borden County, Texas.Horizon: “Pre-Tecovas Horizon”, Dockum Group.
Biostratigraphy:
Age: Middle Carnian Stage, Lower Late Triassic Epoch, Late Triassic.
Material:
TMM 31213-1: Left anterior dorsal paramedian plate, right anterior dorsal paramedian plate fused to right lateral plate (horn forms knob-like process), complete posterior cervical left paramedian plate.
TMM 31213-34: Anterior caudal centrum.
Note: This site also yielded a large Paleorhinus skull (LONG & BALLEW, 1985).
Locality: Rotten Hill Bonebed (PPHM WT), Herring Ranch Locality, Rotten Hill, Potter County, Texas.Horizon: Lower part of the Tecovas Formation, Chinle Group (or Tecovas Member, Dockum Formation).
Biostratigraphy:Age: Adamanian age, Late Carnian Stage, Late Triassic Epoch, Late Triassic.
Material:
PPHM WT 3036.1: Lateral plate.
PPHW WT 2961: 1 anterior presacral lateral plate and one mid to posterior presacral lateral plate.
PPHW WT, 2939, 3216, 2203.1: Pectoral spikes.
?PPHW WT 3122.2: Sacral vertebra with ribs, mid-cervical vertebra.
PPHW WT 3231: Mid-caudal vertebra.
PPHW WT 3122.3: Cervical vertebra.
PPHW WT 2930.1: Caudal vertebra.
Locality: Near head of Home Creek/Swenson Ranch area, Crosby County, Texas.
Horizon: Tecovas Member, Dockum Formation.
Biostratigraphy:Age: Carnian Stage, Late Triassic Epoch, Late Triassic.
Material:
TTUP 9172: Pelvis of a large individual at Kalgary.
Locality: TMM Site 1 (TMM 31172), Crosby County, Texas.Horizon: Tecovas Member, Dockum Formation.
Biostratigraphy:Age: Carnian Stage, Late Triassic Epoch, Late Triassic.
Material:
TMM 31172-10: Right femur missing proximal and distal extremities.
TMM 31172-14: Nearly complete pectoral spike, other armor, and incomplete dorsal centrum.
TMM 31172-24: Incomplete pectoral spike.
Locality: TMM Site 2 (TMM 31173), Crosby County, Texas.Horizon: Tecovas Member, Dockum Formation.
Biostratigraphy:Age: Carnian Stage, Late Triassic Epoch, Late Triassic.
Material:
TMM 31173-54: Armor plates including fused paramedian and lateral plates of cervical series.
TMM 31173-43: Armor plates including numerous cervical spines, cervical paramedian plates and paramedian plates of anterior dorsal region.
TMM 31173-24: Numerous poorly-preserved paramedian plate (Mixed with phytosaur plates.).
TMM 31173-12: Pectoral spikes representing several individuals.
TMM 31173-1: Enormous right pectoral spike.
TMM 31173-2: Pectoral spike missing tip.
TMM 31173-59, -56: Complete pectoral spikes.
TMM 31173-17: Proximal portion of 3 pectoral spikes, one smaller nearly complete spike.
TMM 31173-8: Proximal portion pectoral spike.
TMM 31173-19: Pectoral spike.
TMM 31173- 10: 8 anterior presacral lateral plates.
TMM 31173-131: Eminence from lateral plate.
TMM 31173-130: Mid-cervical vertebra.
TMM 31173-131: Eminence from lateral plate.
TMM 31173-65: 6 cervical vertebrae.
TMM 31173-38: 4 cervical vertebrae.
TMM 31173-128: Mid-dorsal vertebra preserving much of neural arch.
TMM 31173-51: Right ilium missing acetabulum.
TMM 31173-4: Left femur missing distal extremity.
Locality: Red Tank/Walker’s Tank, Crosby County, Texas.Horizon: Tecovas Member, Dockum Formation.
Biostratigraphy:Age: Carnian Stage, Late Triassic Epoch, Late Triassic.
Material:
UMMP 16149: Left scapula.
UMMP 16147: 3 cervical vertebrae.
TTUP 9223: Paramedian plate.
Locality: Espuela Ranch, Dickens County (TMM Locality 1010), Dickens County, Texas.Horizon: Tecovas Member, Dockum Formation.
Biostratigraphy:Age: Carnian Stage, Late Triassic Epoch, Late Triassic.
Material:
TMM 1010-6: Right pectoral spike.
PARKER, 2008
Locality: MNA Locality 1476, badlands approximately 5 km northeast of Many Farms, Apache County, Arizona.
Horizon: Blue Mesa Member, Lower Petrified Forest Formation, Chinle Group.
Biostratigraphy:
Age: Latest Carnian Stage, Late Triassic Epoch, Late Triassic.
Material:
MNA V9300: An almost complete vertebral column, a complete pelvis, and a large amount of the armor carapace.
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Species: smalli PARKER, 2005
Etymology: In honor of Bryan Small for his detailed work on aetosaurs, especially
at teh Post Quarry Desmatosuchus specimens.
= Species nova PARKER, 2003
= Desmatosuchus supurensis CASE, 1920
Holotype: TTU P 9024
Locality: Miller Quarry (Post Quarry MOTT VPL 3624, TUPP, DMNH Locality 1160), 2 km due south of Cooper Creek on the R. C. Miller's Ranch, 8 km southeast of Post, Lat. 33°31’17”N, Long. 101°18’54"W, 9 miles southeast of Post Garza County, West Texas.
Horizon: Upper Part, Lower Cooper Canyon Member, Dockum Formation (Tecovas Member, Dockum Formation).
Biostratigraphy:
Age: Latest Adamanian, Middle Norian Stage, Late Triassic Epoch, Late Triassic.
Material: Skull, mandible, vertebrae, pelvis, femora, lateral shoulder spine, nearly complete cervical armor, and assorted dorsal armor.
After Reyes, et al., 2021.
Paratypes:
TTU P 9023: Skull, mandibles, scapulocoracoid, humerus, dorsal vertebra, lateral shoulder horn, assorted dermal armor.
TTU P 9170: Right humerus, and ulna.
TTU P 9025: Partial preserved skull and dermal armor.
TTU P 9027: Pelvis.
Referred material:
DMNH 8655: Cervical lateral spine.
DMNH 8656: Lateral spine.
DMNH 8657: Partial laeral spine.
DMNH 9893: Paramedian plate fragment.
DMNH 9906: First caudal vertebra missing neural spine.
DMNH 9909: Incomplete large lateral horn.
DMNH 9910: Nearly complete, strongly curved pectoral spike.
DMNH 9913: Mid-caudal vertebra.
DMNH 9939: Sacrum with complete first sacral rib, second rib unattached from enormous individual.
DMNH 9940: 2 well-preserved paramedian plates and portions of others, one base of lateral plate.
DMNH 9941: Nearly complete anterior caudal paramedian plate.
MNA V697: Partial right dorsal lateral plate.
Referred material:?
TTU P9169: 1 isolated paramedian plate.
TTU P9171: Juvenile ulna.
TTU P 9204: Vertebrae, numerous assorted dermal armor, ribs and unprepared material.
TTU P 9207: Pelvis.
TTU P 9229: Vertebrae, radius, numerous assorted dermal armor.
DMNH 9893: Various armor plate fragments.
DMNH 9915: Lateral plate fragment with boss.
DMNH 9998: Incomplete posterior cervical lateral spike.
DMNH 9890: Anterior caudal vertebra, in matrix, poorly preserved.