Species: lancicollis NESOV, 1984
Etymology: Latin, lancea, "lance or spear" and Latin, collum, "neck."
= Azhdarcho imparidens NESOV, 1982 (nomen nudum)

Holotype: CCMGE 1/11915 (Old TsNIGRmuzey, TsNIGRI, LU-N 1/11915)

Locality: Dzharakhuduk (= Dzhyrakhudunk, Itemir, Beleuta), Navoi District of Bukhara Region, Uzbekistan.

Horizon: Lower and Middle part of the Beleuta Svita (Taykarshinskaya unit).

Biostratigraphy:

Age: Upper Turonian Stage, Uppermost Gallic Subepoch to Coniacian Stage, Lower Senonian Subepoch, Lower Gulf Epoch, Late Cretaceous.

Material: Anterior part of a cervical vertebra.

Referred material:

TsNIGRmuzey 1/44: Distal end of a right radius.

TsNIGRmuzey 3/11915: Atlas and epistropheus.

TsNIGRmuzey 4/11915: Posterior cervical vertebra.

TsNIGRmuzey 5/11915, TsNIGRmuzey 6/11915: Anterior cervical vertebra.

TsNIGRmuzey 7/11915: Notarium.

TsNIGRmuzey 9/11915: Part of femur.

TsNIGRmuzey 10/11915: ?Distal part of ulna.

TsNIGRmuzey 9/11915: ?Proximal part of 2nd phalange of 4th digit.

TsNIGRmuzey 18/11915: Anterior dentary.

TsNIGRmuzey 17/11915: Anterior part of dentary.

A) Anterior; B) Left side of TsNIGRmuzey 3/11915: Atlas and epistropheus C) Dorsal; D) Posterior of TsNIGRmuzey 4/11915: Posterior cervical vertebra; E) Inside view of TsNIGRmuzey 18/11915: Anterior dentary; F) Dorsal; G) Ventral of TsNIGRmuzey 6/11915: Anterior cervical vertebra; H) Dorsal; I) Dorsal of TsNIGRmuzey 5/11915: Anterior cervical vertebra; J) Inner view; K) Lateral and somewhat to the rear of TsNIGRmuzey 17/11915: Anterior part of dentary; L) Anterior; M) Dorsal of TsNIGRmuzey 7/11915: Notarium; N) Anterior; O) Posterior of TsNIGRmuzey 9/11915: Part of femur; P) Articular surface; Q) ?Side facing radius of TsNIGRmuzey 10/11915: ?Distal part of ulna; R) TsNIGRmuzey 9/11915: ?Proximal part of 2nd phalange of 4th digit.

AVERIANOV, 2010

Locality: Locality CBI-4, Dzhyrakuuk, Kizylkum Desert, Uzbekistan.

Horizon: Bissekty Formation.

Biostratigraphy:

Age: Upper Turonian Stage, Uppermost Gallic Subepoch, Lower Gulf Epoch, Late Cretaceous.

Material:

CCMGE 6/11915: Anterior fragment of cervical VI.

ZIN PH 11/44: Left humerus distal fragment.

ZIN PH 13/44: Left humeral head.

ZIN PH 25/44: Left proximal humerus fragment lacking humeral head.

ZIN PH 27/44: Left proximal humerus fragment lacking humeral head.

ZIN PH 29/44: Left humeral head.

ZIN PH 31/44: Left metacarpal IV distal fragment with epiphysis.

ZIN PH 34/44: Right metacarpal IV distal fragment with epiphysis.

ZIN PH 39/44: Left proximal fragments of phalanx IV-1.

ZIN PH 90/44: Right humeral head.

ZIN PH 102/44: Right distal syncarpal.

ZIN PH 105/44: Atlas and axis.

ZIN PH 118/44: Unidentified rostrum fragment.

ZIN PH 134/44: Anterior fragment of cervical III.

ZIN PH 152/44: Posterior fragment of cervical VI.

ZINPH 154/44: Dorsal centrum.

ZIN PH 156/44: Dorsal centrum.

ZIN PH 158/44: Dorsal centrum.

ZIN PH 161/44: Proximal end of a dorsal rib.

ZIN PH 177/44: Right fragmentary juvenle coracoids unfused with scapula.

ZIN PH 184/44: Right quadrate condyle fragments.

ZIN PH 202/44: Left proximal fragment of phalanx IV-2.

ZIN PH 221/44: Proximal wing phalange.

ZIN PH 226/44: Distal wing phalange.

Locality: Locality CBI-4a, Dzhyrakuuk, Kizylkum Desert, Uzbekistan.

Horizon: Bissekty Formation.

Biostratigraphy:

Age: Upper Turonian Stage, Uppermost Gallic Subepoch, Lower Gulf Epoch, Late Cretaceous.

Material:

ZIN PH 237/44: Distal wing phalange.

 

Locality: Locality CBI-4e, Dzhyrakuuk, Kizylkum Desert, Uzbekistan.

Horizon: Bissekty Formation.

Biostratigraphy:

Age: Coniacian Stage, Lower Senonian Subepoch, Lower Gulf Epoch, Late Cretaceous.

Material:

ZIN PH 130/44: Centrum of cervical IX.

ZIN PH 147/44: Cervical VI.

ZIN PH 197/44: Right femur proximal fragment.

 

Locality: Locality CBI-4g, Dzhyrakuuk, Kizylkum Desert, Uzbekistan.

Horizon: Bissekty Formation.

Biostratigraphy:

Age: Upper Turonian Stage, Uppermost Gallic Subepoch, Lower Gulf Epoch, Late Cretaceous.

Material:

ZIN PH 9/44: Right humerus distal fragment.

 

Locality: Locality CBI-4v, Dzhyrakuuk, Kizylkum Desert, Uzbekistan.

Horizon: Bissekty Formation.

Biostratigraphy:

Age: Upper Turonian Stage, Uppermost Gallic Subepoch, Lower Gulf Epoch, Late Cretaceous.

Material:

ZIN PH 15/44: Left distal end of an ulna.

ZIN PH 110/44: Centrum of atlas axis.

ZIN PH 135/44: Posterior fragment of cervical III.

 

Locality: Locality CBI-5, Dzhyrakuuk, Kizylkum Desert, Uzbekistan.

Horizon: Bissekty Formation.

Biostratigraphy:

Age: Upper Turonian Stage, Uppermost Gallic Subepoch, Lower Gulf Epoch, Late Cretaceous.

Material:

ZIN PH 12/44: Left humerus lacking distal end.

ZIN PH 28/44: Right radius distal fragment.

ZIN PH 89/44: Left proximal fragment of a humerus.

ZIN PH 140/44: Posterior fragment cervical V.

ZIN PH 170/44: Anterior fragment of a sternum.

ZIN PH 224/44: Proximal fragment of a wing phalange.

 

Locality: Locality CBI-5a, Dzhyrakuuk, Kizylkum Desert, Uzbekistan.

Horizon: Bissekty Formation.

Biostratigraphy:

Age: Upper Turonian Stage, Uppermost Gallic Subepoch, Lower Gulf Epoch, Late Cretaceous.

Material:

CCMGE 10/11915: Right radius distal fragment.

ZIN PH 16/44: Right femur proximal fragment.

ZIN PH 18/44: Left femur proximal fragment.

ZIN PH 21/44: Right femur distal fragment.

ZIN PH 22/44: Right femur distal fragment.

ZIN PH 100/44: Right distal syncarpal.

ZIN PH 101/44: Right distal syncarpal.

ZIN PH 116/44: Anterior dentary fragment.

ZIN PH 125/44: Centrum of cervical IX.

ZIN PH 128/44: Centrum of cervical IX.

ZIN PH 149/44: Anterior fragment of cervical VI.

ZIN PH 151/44: Posterior fragment of cervical VI.

ZIN PH 171/44: Anterior fragment of a sternum.

ZIN PH 173/44: Fragments with glenoid fossa of a scapulocoracoid.

ZIN PH 185/44: Left quadrate condyle fragments.

ZIN PH 187/44: Left metacarpal IV distal shaft fragments.

ZIN PH 190/44: Left tibiotarsus distal fragment.

ZIN PH 200/44: Proximal fragment of right second or third metatarsal.

ZIN PH 205/44: Left proximal fragments of phalanx IV-2.

ZIN PH 210/44: Right proixmal fragment of phalanx IV-2.

ZIN PH 213/44: Left proximal fragments of phalanx IV-3.

ZIN PH 220/44: Proximal phalange

 

Locality: Locality CBI-7a, Dzhyrakuuk, Kizylkum Desert, Uzbekistan.

Horizon: Bissekty Formation.

Biostratigraphy:

Age: Upper Turonian Stage, Uppermost Gallic Subepoch, Lower Gulf Epoch, Late Cretaceous.

Material:

CCMGE 8/11915: Right radius proximal fragment.

 

Locality: Locality CBI-14, Dzhyrakuuk, Kizylkum Desert, Uzbekistan.

Horizon: Bissekty Formation.

Biostratigraphy:

Age: Upper Turonian Stage, Uppermost Gallic Subepoch, Lower Gulf Epoch, Late Cretaceous.

Material:

CCMGE 3/11915: Centrum of atlas axis.

CCMGE 6/11915: Anterior fragment of cervical VI.

ZIN PH 5/44: Notarium fragments.

ZIN PH 6/44: Notarium fragments.

ZIN PH 7/44: 2 fused dorsal vertebrae.

ZIN PH 17/44: Left femur shaft.

ZIN PH 19/44: Right femur proximal fragment.

ZIN PH 21/44: Left femur distal fragment.

ZIN PH 26/44: Left proximal fragment.

ZIN PH 30/44: Left metacarpal IV distal fragment with epiphysis.

ZIN PH 32/44: Left metacarpal IV distal fragment with epiphysis.

ZIN PH 33/44: Left metacarpal IV distal shaft fragments.

ZIN PH 35/44: Left metacarpal IV distal shaft fragments.

ZIN PH 37/44: Left proximal fragments of phalanx IV-1.

ZIN PH 59/44: Fragment of fused premaxillae and nasals.

ZIN PH 85/44, 114/44: Anterior premaxillary fragments.

ZIN PH 86/44: Left ulna lacking distal end and with mostly destroyed porximal end.

ZIN PH 88/44: Right radius proximal fragment.

ZIN PH 93/44: Right juvenile humerus.

ZIN PH 94/44: Right proximal syncarpal.

ZIN PH 97/44: Right proximal syncarpal.

ZIN PH 99/44: Right distal syncarpal.

ZIN PH 103/44: Left distal syncarpal.

ZIN PH 104/44: Left fragment distal syncarpal.

ZIN PH 106/44: Atlas axis neural arch.

ZIN PH 107/44: Atlas axis neural arch.

ZIN PH 111/44: Centrum of atlas axis.

ZIN PH 113/44: Unidentified rostrum fragment.

ZIN PH 115/44: Anterior dentary fragment.

ZIN PH 119/44: Unidentified rostrum fragment.

ZIN PH 120/44: Left mandibular fragment with glenoid fossa.

ZIN PH 124/44: Centrum of cervical IX.

ZIN PH 129/44: Centrum of cervical IX.

ZIN PH 131/44: Cervical III.

ZIN PH 132/44: Cervical III.

ZIN PH 133/44: Anterior cervical III.

ZIN PH 141/44: Posterior fragment of cervical V of a juveinle

ZIN PH 145/44: Anterior fragment of cervical V.

ZIN PH 148/44: Cervical VI.

ZIN PH 155/44: Dorasal centrum.

ZIN PH 159/44; Dorsal centrum.

ZIN PH 160/44: Dorsal centrum.

ZIN PH 162/44: Distal end of rib.

ZIN PH 163/44: Distal end of rib.

ZIN PH 164/44: Distal end of rib.

ZIN PH 165/44: Distal end of rib.

ZIN PH 172/44: Juvenile neural arch of cervical III.

ZIN PH 174/44: Right scapulocoracoid.

ZIN PH 175/44: Right scapulocoracoid.

ZIN PH 178/44: Right fragmentary juvenle coracoids unfused with scapula.

ZIN PH 179/44: Left distal coracoid fragment.

ZIN PH 180/44: Right distal coracoid fragment.

ZIN PH 181/44: Right distal coracoid fragment.

ZIN PH 182/44: Left fragmentary juvenile coracoids unfused with scapula.

ZIN PH 183/44: Preaxial carpal.

ZIN PH 186/44: Left quadrate condyle fragments.

ZIN PH 191/44: Right tibiotarsus distal fragment.

ZIN PH 192/44: Right femur proximal fragment.

ZIN PH 193/44: Right femur proximal fragment.

ZIN PH 194/44: Right femur proximal fragment.

ZIN PH 195/44: Left femur proximal fragment.

ZIN PH 199/44: Right proximal radius fragment.

ZIN PH 202/44: Left proximal fragments of phalanx IV-1.

ZIN PH 203/44: Left proximal fragments of phalanx IV-2.

ZIN PH 206/44: Left proximal fragment of phalanx IV-2.

ZIN PH 207/44: Left proximal fragment of phalanx IV-2.

ZIN PH 208/44: Left proximal fragment of phalanx IV-2.

ZIN PH 209/44: Left proximal fragment of phalanx IV-2.

ZIN PH 211/44: Right proximal fragment of phalanx IV-2.

ZIN PH 212/44: Right proximal fragment of phalanx IV-2.

ZIN PH 214/44: Left proximal fragments of phalanx IV-3.

ZIN PH 215/44: Right proximal fragments of phalanx IV-3.

ZIN PH 216/44-219/44: Fragments of phalanx IV-4.

ZIN PH 222/44: Proximal fragment of a wing phalange.

ZIN PH 223/44: Proximal fragment of a wing phalange.

ZIN PH 227/44: Distal wing phalange.

ZIN PH 228/44: Distal wing phalange.

ZIN PH 229/44: Distal wing phalange.

ZIN PH 230/44: Proximal wing phalange.

ZIN PH 231/44: Distal wing phalange.

ZIN PH 232/44: Distal wing phalange.

ZIN PH 234/44: Distal wing phalange.

ZIN PH 235/44: Distal wing phalange.

ZIN PH 240/44: Proximal pedal phalanx.

ZIN PO 5216: Unidentified rostrum fragment.

ZIN PO 5217: Unidentified rostrum fragment.

ZIN PO 6442: Distal pedal phalanx.

ZIN PO 6472: Left juvenile humerus lacking proximal and distal ends.

 

Locality: Locality CBI-14v, Dzhyrakuuk, Kizylkum Desert, Uzbekistan.

Horizon: Bissekty Formation.

Biostratigraphy:

Age: Upper Turonian Stage, Uppermost Gallic Subepoch, Lower Gulf Epoch, Late Cretaceous.

Material:

ZINPH 1/44: Right radius distal fragment.

 

Locality: Locality CBI-16, Dzhyrakuuk, Kizylkum Desert, Uzbekistan.

Horizon: Bissekty Formation.

Biostratigraphy:

Age: Upper Turonian Stage, Uppermost Gallic Subepoch, Lower Gulf Epoch, Late Cretaceous.

Material:

ZIN PH 112/44: Fragment of fused maxillae and premaxillae.

ZIN PH 153/44: Notarium fragments.

Locality: Locality CBI-17, Dzhyrakuuk, Kizylkum Desert, Uzbekistan.

Horizon: Bissekty Formation.

Biostratigraphy:

Age: Upper Turonian Stage, Uppermost Gallic Subepoch, Lower Gulf Epoch, Late Cretaceous.

Material:

CCMGE 4/11915: Posterior fragment of cervical V.

CCMGE 9/11915: Left femur proximal fragment.

CCMGE 17/11915: Unidentified rostrum fragment.

CCMGE 52/11915: Distal pedal phalanx.

ZIN PH 96/44: Right proximal syncarpal.

ZIN PH 109/44: Centrum of atlas axis.

ZIN PH 127/44: Centrum of cervical IX.

ZIN PH 137/44: Centrum of Cervical VIII.

ZIN PH 167/44: Distal end of a rib.

ZIN PH 188/44: Right metacarpal IV distal shaft fragments.

ZIN PH 189/44: Right metacarpal IV distal shaft fragments.

ZIN PH 196/44: Left humeurs proximal fragment.

ZIN PH 225/44: Proximal fragment of a wing phalange.

ZIN PH 233/44: Distal wing phalange.

ZIN PO 3471: Anterior dentary fragment.

 

Locality: Locality CBI-17a, Dzhyrakuuk, Kizylkum Desert, Uzbekistan.

Horizon: Bissekty Formation.

Biostratigraphy:

Age: Upper Turonian Stage, Uppermost Gallic Subepoch, Lower Gulf Epoch, Late Cretaceous.

Material:

CCMGE 18/11915: Unidentified rostrum fragment.

 

Locality: Locality CBI-41, Dzhyrakuuk, Kizylkum Desert, Uzbekistan.

Horizon: Bissekty Formation.

Biostratigraphy:

Age: Upper Turonian Stage, Uppermost Gallic Subepoch, Lower Gulf Epoch, Late Cretaceous.

Material:

CCMGE 8/12454: Right humerus distal fragment.

ZIN PH 10/44: Right humerus distal fragment.

ZIN PH 14/44: Right distal ulna end.

ZIN PH 24/44: Right humerus proximal fragment lacking humeral head.

ZIN PH 139/44: Posterior fragment of cervical V.

ZIN PH 142/44: Posterior fragment of cervical V.

ZIN PH 143/44: Posterior fragment of cervical V.

 

Locality: Locality CBI-50, Dzhyrakuuk, Kizylkum Desert, Uzbekistan.

Horizon: Bissekty Formation.

Biostratigraphy:

Age: Upper Turonian Stage, Uppermost Gallic Subepoch, Lower Gulf Epoch, Late Cretaceous.

Material:

ZIN PH 117/44: Unidentified rostrum fragment.

 

Locality: Locality CBI-52, Dzhyrakuuk, Kizylkum Desert, Uzbekistan.

Horizon: Bissekty Formation.

Biostratigraphy:

Age: Upper Turonian Stage, Uppermost Gallic Subepoch, Lower Gulf Epoch, Late Cretaceous.

Material:

ZIN PH 38/44: Right proximal fragments of phalanx IV-1.

 

Locality: Locality CBI-56, Dzhyrakuuk, Kizylkum Desert, Uzbekistan.

Horizon: Bissekty Formation.

Biostratigraphy:

Age: Upper Turonian Stage, Uppermost Gallic Subepoch, Lower Gulf Epoch, Late Cretaceous.

Material:

ZIN PH 8/44: Left humerus distal fragment.

ZIN PH 121/44: Posterior fragment of cervical V of a juvenile.

 

 

Locality: Locality CBI-, Dzhyrakuuk, Kizylkum Desert, Uzbekistan.

Horizon: Bissekty Formation.

Biostratigraphy:

Age: Upper Turonian Stage, Uppermost Gallic Subepoch, Lower Gulf Epoch, Late Cretaceous..

Material:

ZIN PH 23/44: Left femur distal fragment.

ZIN PH 36/44: Left proximal fragment of phalanx IV-1.

ZIN PH 41/44: Left proximal ulna fragment.

ZIN PH 87/44: Right radius distal fragment.

ZIN PH 93/44: Right juvenile humerus.

ZIN PH 95/44: RIght proximal syncarpal.

ZIN PH 108/44: Neural arch.

ZIN PH 122/44: Centrum of cervical IX.

ZIN PH 123/44: Centrum of cervical IX.

ZIN PH 126/44: Centrum of cervical IX.

ZIN PH 136/44: Neural arch fragment of cervical III.

ZIN PH 144/44: Cervical IV.

ZIN PH 146/44: Posterior fragment of cervical VI.

ZIN PH 150/44: Anterior fragment of cervical VI.

ZIN PH 166/44, 168/44, 169/44: Distal end of ribs.

ZIN PH 176/44: Left fragmentary juvenile coracoids unfused with scapula.

ZIN PH 197/44: Right femur proximal fragment.

ZIN PH 201/44: Left proximal fragments of phalanx IV-1.

ZIN PH 236/44: Distal wing phalange.

ZIN PH 239/44: Distal wing phalange.

ZIN PH 238/44: Proximal wing phalange.

ZIN PH 241/44: Distal pedal phalanx.

 

Locality: Locality VBI-14, Dzhyrakuuk, Kizylkum Desert, Uzbekistan.

Horizon: Bissekty Formation.

Biostratigraphy:

Age: Upper Turonian Stage, Uppermost Gallic Subepoch, Lower Gulf Epoch, Late Cretaceous.

Material:

CCMG 7/11915: Notarium fragments.

 

NESOV, 1995

Locality: Dzhyrakuduk locality, site CBI-17, north of Bukhara in central Kyzylkum Desert, Uzbekistan.

Horizon: Lower part of Bissekty Formation.

Biostratigraphy:

Age: Upper Turonian Stage, Uppermost Gallic Subepoch, Lower Gulf Epoch, Late Cretaceous.

Material:

N 8/11915: Proximal part of phalanx 2 of digit IV.