Species: wellnoferi BUFFETAUT, LEPAGE & LEPAGE, 1998
Etymology: In honor of Dr. Peter Wellnhofer (Munich), a great expert on pterosaurs,
and a great friend of the senior author.
Holotype: Musee Geologique Cantonal de Lausanne, number 59’583
Locality: Cliffs of Ecqueville, Octeville, Seine-Maritime Department, Haute-Normandie, France.
Horizon: Argiles d’Ecqueville.
Biostratigraphy:
Age: Upper Kimmeridgian Stage, Middle Malm Epoch, Late Jurassic.
Material: Associated upper and lower jaws on a block of clay.