Species: gracilis (MARSH, 1876a) MARSH, 1876b
Etymology: Latin, gracilis, "slender."
= Pteranodon gracilis MARSH, 1876a
= Nyctodactylus gracilis (MARSH, 1876b) MARSH, 1881
= Pteranodon (Nyctosaurus) gracilis (MARSH, 1876)
= Nyctosaurus leptodactylus WILLISTON, 1903
Etymology:

Holotype: YPM 1178

Locality: Huckberry, Gove County, Kansas.

Horizon: Upper Smoky Hill Member, Niobrara Formation.

Biostratigraphy:

Age: Santonian or Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material: Right and left scapulae and coracoids, 2 cervical vertebrae, 5 free dorsal vertebrae, another ?dorsal vertebra, ?8th cervical vertebra, left ulna and radius, right and left carpals, right and left 4th metacarpals, right and left 1st phalanges of the 4th digit of an immature individual.

Nyctosaurus gracilis Holotype: YPM 1178, after Schoch (1984): A-B) pectoral region; A) dorsal; B) ventral views; C) right first phalanx of digit IV; D) right carpals and metacarpal IV; E) left first phalanx of digit IV, and F) left ulna, radius, carpals and metacarpal IV.

Skull of Nyctosaurus gracilis; A) dorsal; B) ventral, and C) new reconstruction of lateral view.

Nyctosaurus gracilis referred material: A) Field Museum of Natural History Specimen, after Case, (1982); B, C) new reconstruction: B) dorsal and C) lateral views.

Referred material:

SMM 7968: Humerus, radius, ulna, metacarpal IV, phalanx 1, phalanx 4 and femur.

CMNH: Humerus, radius, ulna, metacarpal VI, phalanx 1and 2, femur and tibia.

WILLISTON, 1902, 1903

Number:? Skull and skeleton.

 

BENNETT, 1991

Locality: Sec. 30, T15S, R26W, Gove County, Kansas.

Horizon: Lower Smoky Hill Member, Niobrara Formation.

Biostratigraphy:

Age: Santonian or Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

National Science Museum in Tokyo: Incomplete skeleton lacking skull.

 

BENNETT, 1993

Locality:

Horizon:

Biostratigraphy:

Age:

Material:

FHSM 2148: Notarium, synsacrum, scapulacoracoid, humerus, ulna, carpal, wing-metacarpal, wing-phalanage 1, wing-phalanx and pelvis of an immature individual.

KUVP 49401: Wing phalanx of an immature individual.

KUVP 85394: Wing metacarpal, wing phalange 1 of an immature individual.

KUVP 85463: Wing phalange 1, and wing phalanx of an immature individual.

USNM 93000: Skull, atlas-axis, notarium, synsacrum, humerus, radius, ulna, carpal, wing metacarpal, wing phalange 1, wing phalanx, pelvis, femur and tibia.

YPM 2303: Skull, radius, ulna, wing metacarpal, wing phalange 1 and wing phalanx of an immature individual.

YPM 2334: Skull, mandible, scapulacoracoid, humerus, radius, ulna, femur and tibia of an immature individual.

YPM 2341: Radius, ulna, wing phalanx of an immature individual.

YPM 2527: Mandible, notarium, humerus, radius, ulna, carpal, and wing phalanx of an immature individual.

Nyctosaurus referred specimens.

LAMBE, 1914

Locality: Sampson’s pasture, Hackberry Creek, 13 miles south of Quinter, Gove County, Kansas.

Horizon: Niobrara Formation.

Biostratigraphy:

Age: Late Cretaceous.

Material:

CMN (old NMC) 8168: Humerus, radius, ulna, carpal, wing metacarpal, wing phalange 1, wing phalanx of an immature individual.

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Species: nanus (MARSH, 1881) EATON, 1910
Etymology: Latin, nanus, “dwarf.”
= Pteranodon nanus MARSH, 1881

Holotype: YPM 1182

Locality: Between Graham and Rooks counties, Kansas.

Horizon: Lower Smoky Hill Member, Niobrara Formation.

Biostratigraphy:

Age: Late Coniacian-Early Santonian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material: Left scapula, left coracoid, left humerus, dorsal vertebra with ankylosed rib, and other bone fragments.

Nyctosaurus nanus Holotype: YPM 1182, after Schoch, (1984): A, B) trunk vertebra with ankylosed rib; A) dorsal; B) ventral view; C-D) left scapula and coracoid; C) anterior; D) posterior views; E-F) left humerus; E) external and F) internal views.

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Species: lamegoi PRICE, 1953
Etymology: In honor of the geologist Alberto Ribeiro Lamego.
= Pteranodon (Nyctosaurus) lamegoi (PRICE, 1953)

Holotype: DGM. No. 238-R.

Locality: Fazeda Congo, Rio Gramame, Paraiba State, Brazil.

Horizon: Gramame Itamaraca Formation.

Biostratigraphy:

Age: Maastrichtian Stage, Upper Senonian Subepoch, Upper Gulf Epoch, Late Cretaceous.

Material: Humerus, missing the distal and proximal end.

Nyctosaurus lamegoi Holotype: DGM. No. 238-R, after Price (1953): A) anterior; B) posterior; and C) reconstruction.
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Species: bonneri (MILLER, 1972) WELLNHOFER, 1978
Etymology: In honor of O. W. Bonner, who first described the specimen in his thesis.
= Pteranodon (Nyctosaurus) bonneri MILLER, 1972
= Nyctosaurus sternbergi BONNER, 1964 (chironym) /HARKSEN, 1966 (as Pteranodon sternbergi)

Holotype: Sternberg Memorial Museum No. 11311

Locality: Elkader, Logan county, Kansas.

Horizon: Upper Smoky Hill Member, Niobrara Formation.

Biostratigraphy:

Age: Santonian or Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material: Skull, dentary missing anterior portion, atlas-axis, 8 cervical, 8 dorsal, a false sacral, and 3 sacral vertebrae, ischia, pubi, ilia, left wing is complete, right humerus, radius-ulna, 4th metacarpal, 4th phalanx, sternum, coracoids, scapulae, femora, tibia, ribs, notarium fragments, pteroids, and some metatarsal bones.

Skull of Nyctosaurus bonneri Holotype: Sternberg Memorial Museum No. 11311. after Miller (1971); A) as preserved; B) new reconstruction.

Skeleton of Nyctosaurus bonneri Holotype: Sternberg Memorial Museum No. 11311. after Miller (1971); A) as preserved; B-C) new reconstruction; B) dorsal, and C) lateral views.

Referred material:

YPM 2395: Humerus, radius, ulna, metacarpal 1.