Genus: Nyctosaurus MARSH, 1876b
Etymology: Greek, nyktis (nyx), “night”, and Greek, sauros, 'lizard'.
= Nyctodactylus MARSH, 1881
Etymology: Greek, nyktis (nyx), “night”, and Greek, dactylus, ' finger'.
= Nytosaurus MOLNAR, 1980 (sic)

Species: gracilis (MARSH, 1876a) MARSH, 1876b
Etymology: Latin, gracilis, "slender."
= Pteranodon gracilis MARSH, 1876a
= Nyctodactylus gracilis (MARSH, 1876b) MARSH, 1881
= Pteranodon (Nyctosaurus) gracilis (MARSH, 1876)
= Nyctosaurus leptodactylus WILLISTON, 1903
Etymology:

Holotype: YPM 1178

Locality: Huckberry, Gove County, Kansas.

Horizon: Upper Smoky Hill Member, Niobrara Formation.

Biostratigraphy:

Age: Santonian or Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material: Right and left scapulae and coracoids, 2 cervical vertebrae, 5 free dorsal vertebrae, another ?dorsal vertebra, ?8th cervical vertebra, left ulna and radius, right and left carpals, right and left 4th
metacarpals, right and left 1st phalanges of the 4th digit of an immature individual.

Referred material:

SMM 7968: Humerus, radius, ulna, metacarpal IV, phalanx 1, phalanx 4 and femur.

CMNH: Humerus, radius, ulna, metacarpal VI, phalanx 1and 2, femur and tibia.

WILLISTON, 1902, 1903

Number:? Skull and skeleton.

 

BENNETT, 1991

Locality: Sec. 30, T15S, R26W, Gove County, Kansas.

Horizon: Lower Smoky Hill Member, Niobrara Formation.

Biostratigraphy:

Age: Santonian or Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

National Science Museum in Tokyo: Incomplete skeleton lacking skull.

 

BENNETT, 1993

Locality:

Horizon:

Biostratigraphy:

Age:

Material:

FHSM 2148: Notarium, synsacrum, scapulacoracoid, humerus, ulna, carpal, wing-metacarpal, wing-phalanage 1, wing-phalanx and pelvis of an immature individual.

KUVP 49401: Wing phalanx of an immature individual.

KUVP 85394: Wing metacarpal, wing phalange 1 of an immature individual.

KUVP 85463: Wing phalange 1, and wing phalanx of an immature individual.

USNM 93000: Skull, atlas-axis, notarium, synsacrum, humerus, radius, ulna, carpal, wing metacarpal, wing phalange 1, wing phalanx, pelvis, femur and tibia.

YPM 2303: Skull, radius, ulna, wing metacarpal, wing phalange 1 and wing phalanx of an immature individual.

YPM 2334: Skull, mandible, scapulacoracoid, humerus, radius, ulna, femur and tibia of an immature individual.

YPM 2341: Radius, ulna, wing phalanx of an immature individual.

YPM 2527: Mandible, notarium, humerus, radius, ulna, carpal, and wing phalanx of an immature individual.

 

LAMBE, 1914

Locality: Sampson’s pasture, Hackberry Creek, 13 miles south of Quinter, Gove County, Kansas.

Horizon: Niobrara Formation.

Biostratigraphy:

Age: Late Cretaceous.

Material:

CMN (old NMC) 8168: Humerus, radius, ulna, carpal, wing metacarpal, wing phalange 1, wing phalanx of an immature individual.

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Species: nanus (MARSH, 1881) EATON, 1910
Etymology: Latin, nanus, “dwarf.”
= Pteranodon nanus MARSH, 1881

Holotype: YPM 1182

Locality: Between Graham and Rooks counties, Kansas.

Horizon: Lower Smoky Hill Member, Niobrara Formation.

Biostratigraphy:

Age: Late Coniacian-Early Santonian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material: Left scapula, left coracoid, left humerus, dorsal vertebra with ankylosed rib, and other bone fragments.
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Species: lamegoi PRICE, 1953
Etymology: In honor of the geologist Alberto Ribeiro Lamego.
= Pteranodon (Nyctosaurus) lamegoi (PRICE, 1953)

Holotype: DGM. No. 238-R.

Locality: Fazeda Congo, Rio Gramame, Paraiba State, Brazil.

Horizon: Gramame Itamaraca Formation.

Biostratigraphy:

Age: Maastrichtian Stage, Upper Senonian Subepoch, Upper Gulf Epoch, Late Cretaceous.

Material: Humerus, missing the distal and proximal end.


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Species: bonneri (MILLER, 1972) WELLNHOFER, 1978
Etymology: In honor of O. W. Bonner, who first described the specimen in his thesis.
= Pteranodon (Nyctosaurus) bonneri MILLER, 1972
= Nyctosaurus sternbergi BONNER, 1964 (chironym) /HARKSEN, 1966 (as Pteranodon sternbergi)

Holotype: Sternberg Memorial Museum No. 11311

Locality: Elkader, Logan county, Kansas.

Horizon: Upper Smoky Hill Member, Niobrara Formation.

Biostratigraphy:

Age: Santonian or Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material: Skull, dentary missing anterior portion, atlas-axis, 8 cervical, 8 dorsal, a false sacral, and 3 sacral vertebrae, ischia, pubi, ilia, left wing is complete, right humerus, radius-ulna, 4th metacarpal, 4th phalanx, sternum, coracoids, scapulae, femora, tibia, ribs, notarium fragments, pteroids, and some metatarsal bones.

Referred material:

YPM 2395: Humerus, radius, ulna, metacarpal 1.