Genus: Pteranodon MARSH, 1876
Etymology: Greek, Ptero-, pteron, "wing" and Greek, -odon, "toothless."
= Ornithochirus COPE, 1872
Etymology:
= Ornithostoma WILLISTON, 1893 non SEELEY, 1870
= Subgenus: Longicepia MILLER, 1971 (published in 1972)
Etymology: The subgeneric name is derived from the specific name of P. longiceps and the species is designated as the type species.
= Subgenus: Pteranodon MILLER, 1972

Species: ingens (MARSH, 1872) MARSH, 1876
Etymology: Latin, ingens, “enormous.”
= Pterodactylus ingens MARSH, 1872
= Ornithostoma ingens (MARSH, 1872) WILLISTON, 1893
= Pteranodon (Longicepia) ingens (MARSH, 1872) MILLER, 1971 (published in 1972)
= Pteranodon (Pteranodon) ingens (MARSH, 1872) MILLER, 1972

Holotype: YPM 1170

Locality: North side of the Smoky (Hill) River, 18 miles (=30.0) km) east of Fort Wallace, 5 miles west of Russell Springs, Sec. 13 and 24, T12S, R36W, Logan County, Kansas.

Horizon: Smoky Hill Member, Niobrara Formation.

Biostratigraphy:

Age: Santonian or Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material: Distal end of right metacarpal IV, proximal end of ?metacarpal IV, proximal end of? ulna, distal end of 1st phalanx of digit IV.

Referred material:


YPM 1175: Mandublar ramus, scapula, coracoid, humerus, ulna, several wing bones, including an ulna, a radius, a humerus, a 4th metacarpal, 4d, 4v, femur, a metatarsal and a tooth?
Note: The specimens were found at 4 different locations and were very probably the remains of 4
Note: Mandublar ramus has associated hyperplasia on the external surface of the left side due to necroses (BENNETT, 2003).

Referred material to 1175: Femur, sacrum, notarial fragment, cervical ribs and several cervical vertebrae.

 

Locality: South side of the Smoky Hill River, 20 miles (=32.2 km) east of Fort Wallace, 3 miles west of Russell Springs, Sec. 19, 20, 29, 30, T13S, R35W, Logan County, Kansas.

Horizon: Upper Smoky Hill Member, Niobrara Formation.

Biostratigraphy:

Age: Santonian or Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

YPM 1169: Proximal end of ?1st phalanx of digit IV and numerous wing-bone fragments.

YPM 1171: Distal end of left metacarpal IV and bone fragments.

 


Locality: North side of the Smoky Hill River, 26 miles (=41.9 km), east of Fort Wallace, 3 miles east of Russell Springs, Sec. 32, 33, T13S, R34W, Logan County, Kansas.

Horizon: Upper Smoky Hill Member, Niobrara Formation.

Biostratigraphy:

Age: Santonian or Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

YPM 1172: Distal end of right metacarpal IV and 2 bone fragments of an immature individual.

 

LAMBE, 1914

Locality: Sampson’s pasture, Hackberry Creek, Gove County, 13 miles south of Quinter, Kansas.

Horizon: Niobrara Formation.

Biostratigraphy:

Age: Late Cretaceous.

Material:

CMN (old NMC) 8167: Humerus, ulna, metacarpal 4, 4d, wing phalange 1-4.

 

= Ornithochirus umbrosus COPE, 1872
= Ornithocheirus umbrosus (COPE, 1872) emend NEWTON, 1888
= Pteranodon umbrosus (COPE, 1872)
= Pterodactylus umbrosus (COPE, 1872)
Etymology: Latin, umbrosus, “shady.”

Holotype: AMNH 1571

Locality: [Twin] Butte Creek, Logan County, Kansas.

Horizon: Upper Smoky Hill Member, Niobrara Formation.

Biostratigraphy:

Age: Santonian or Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material: Fragmentary 4th metacarpal and the 1st phalanx, and 2 phalanges with claws.

_____________________________________________________________________________________

Species: longiceps MARSH, 1876
Etymology: Latin, longus, “long” and Latin, –ceps, “head.”
= Pteranodon (Longicepia) longiceps (MARSH, 1876) MILLER, 1971 (published in 1972)
= Pteranodon (Pteranodon) longiceps (MARSH, 1876) MILLER, 1972

Holotype: YPM 1177

Locality: Near Smoky Hill River, “Wallace” County, Kansas. Williston’s (1876) field notebook reads” “Loc. 3 mi. NE of Monument Rocks in fine yellow chalk. Wash gentle for 20 feet + then quite precipitous.” 3 miles northeast of Monument Rocks is in Gove County, not Wallace or Logan. Based on Examination of the exposures northeast of Monument Rocks, Bennett (1991) reported that the skull probably was collected from between 2 m above marker Unit 15 and Marker Unit 16, (Bennett, 1994)

Horizon: Smoky Hill Member, Niobrara Formation.

Biostratigraphy:

Age: Santonian or Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material: Skull, atlas and axis of a female.
Note: Additional remains not preserved in the slab include skull fragments, cervical rib, partial radius, proximal end of a wing phalanx 2 and other fragments.

Referred material:

YPM 5775: Skull.

SMM 11402: Skull missing the anterior portion and most of crest, atlas-axis, 3rd to 6th cervicals, dorsal vertebra, sternum, ribs, right and left humerus, right radius, ulna, right pteroid bones, 2 metacarpals, phalanx 1 of digit IV, phalanx 3 of digit IV, left femur, anterior right femur, 4th metatarsal, and several phalanges and a claw.


Locality: 4 miles Northeast of Monument Rocks, Kansas.

Horizon: Smoky Hill Member, Niobrara Formation.

Biostratigraphy:

Age: Santonian or Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

YPM 2473: Crest, radius, ulna, metacarpal 4, 4d, 4v, wing phalange 1.

 

ANONYMOUS, 1997

Locality: Kansas.

Horizon: Niobrara Formation.

Biostratigraphy:

Age: Late Cretaceous.

Material:

Private collection of Fred Nuss and Candace Williams: Skull lacking some of crest, part of lower jaw, 3 cervical vertebrae, fragment of scapula and wing.

 

BENNETT, 1991

Locality: 9 miles northeast of Elkader, Logan County, Kansas.

Horizon: Niobrara Formation.

Biostratigraphy:

Age: Coniacian Stage, Lower Senonian Subepoch, Lower Gulf Epoch, Late Cretaceous.

Material:

AMNH 149: Skull.

 


Locality: 11 miles South of Quintu (Quinter), Lane County (sic), northeastern Gove County, Kansas.

Horizon: Niobrara Formation.

Biostratigraphy:

Age: Coniacian Stage, Lower Senonian Subepoch, Lower Gulf Epoch, Late Cretaceous.

Material:

AMNH 5099: Wing phalange 3, femora, tibiae.

 


Locality: Twin Butte Creek, Logan County, Kansas.

Horizon: Niobrara Formation.

Biostratigraphy:

Age: Coniacian Stage, Lower Senonian Subepoch, Lower Gulf Epoch, Late Cretaceous.

Material:

CM 1539: Humerus, radius and metacarpal 4.

 


Locality: Twin Butte Creek, Kansas.

Horizon: Niobrara Formation.

Biostratigraphy:

Age: Coniacian Stage, Lower Senonian Subepoch, Lower Gulf Epoch, Late Cretaceous.

Material:

CM “UNC A”:

 


Locality: About 2 miles west of Russell Springs, just on the north side of the Smoky Hill River, Logan County, Kansas.

Horizon: Niobrara Formation.

Biostratigraphy:

Age: Coniacian Stage, Lower Senonian Subepoch, Lower Gulf Epoch, Late Cretaceous.

Material:

DMNH 1732: Fragmentary skull.


Locality: About 5 miles northwest of Elkader- 1/4 mile due east of a wing charger reversed to take static from a pipe line, Sec. 1, 12, 13, T14S, R33W, Kansas.

Horizon: Niobrara Formation.

Biostratigraphy:

Age: Coniacian Stage, Lower Senonian Subepoch, Lower Gulf Epoch, Late Cretaceous.

Material:

FHSM VP 184: Scapula, coracoid, humerus, ulna, metacarpal 4, 4d, 4v, wing phalange 1-4.

 

Locality: “A stone quarry West of Highway 25, Near Russell Spring, Localities 13, 26, and 27 and east and west of Russell Springs, Logan County, Kansas.

Horizon: Niobrara Formation.

Biostratigraphy:

Age: Coniacian Stage, Lower Senonian Subepoch, Lower Gulf Epoch, Late Cretaceous.

Material:

FHSM VP 2120: Radius, ulna, metacarpal 4, 4d, and wing phalange 1.

 


Locality: SE 1/4, Sec. 16, T15S, R34W, Logan County, Kansas.

Horizon: Niobrara Formation.

Biostratigraphy:

Age: Coniacian Stage, Lower Senonian Subepoch, Lower Gulf Epoch, Late Cretaceous.

Material:

FMNH PR 494: Metacarpal 4, 4v, wing phalange 1.

 

Locality: “Butte Creek”, Logan County, Kansas.

Horizon: Niobrara Formation.

Biostratigraphy:

Age: Coniacian Stage, Lower Senonian Subepoch, Lower Gulf Epoch, Late Cretaceous.

Material:

KUVP 976: Skull of an immature individual.

KUVP 977: Metacarpal 4, 4d, 4v, wing phalange 1.

 

Locality: Western part of Butte Creek, Logan County, Kansas.

Horizon: Niobrara Formation.

Biostratigraphy:

Age: Coniacian Stage, Lower Senonian Subepoch, Lower Gulf Epoch, Late Cretaceous.

Material:

LACM 50926: Fragmentary skull, right radius, ulna, left scapula, coracoid humerus, radius, metacarpal 4, both metacarpal 4d, both wing phalange 1-2, wing phalange 3, femora and right tibia.
Note: Mandibular ramus is badly deformed, necroses (BENNETT, 2003).
Note: An imbedded tooth from Cretoxyrhina mantelli (HONE, WITTON & HABIB, 2018).

LACM 50926 “A”: Ulna.

LACM “B”:

 


Locality: SE 1/4, Sec. 6, T15S, R35W, Logan County, Kansas.

Horizon: Niobrara Formation.

Biostratigraphy:

Age: Coniacian Stage, Lower Senonian Subepoch, Lower Gulf Epoch, Late Cretaceous.

Material:

LACM 51137: Fragmentary skull.

 


Locality: NE 1/4 of Sec. 24, T14S, R31W, Gove County, Kansas.

Horizon: Niobrara Formation.

Biostratigraphy:

Age: Coniacian Stage, Lower Senonian Subepoch, Lower Gulf Epoch, Late Cretaceous.

Material:

UNSM 50130: Scapula, humerus, radius, ulna, metacarpal 4, wing phalange 1.

 


Locality: 2-3 miles northeast of Monument Rocks, near the Smoky (Hill) River, Wallace County, Kansas.

Horizon: Upper Smoky Hill Member, Niobrara Formation.

Biostratigraphy:

Age: Santonian or Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

USNM 13656:

 


Locality: 12 miles south of Russell Springs, probably in the vicinity of localities 1, 6, 7, and 8, Logan County, Kansas.

Horizon: Niobrara Formation.

Biostratigraphy:

Age: Coniacian Stage, Lower Senonian Subepoch, Lower Gulf Epoch, Late Cretaceous.

Material:

UUPI R.197: Both scapulae, coracoid, left humerus, radius, metacarpal 4, 4d, 4v, wing phalange 1, both femora, tibia.

 


Locality: 1 miles south of Butte Creek, Wallace County, Kansas.

Horizon: Niobrara Formation.

Biostratigraphy:

Age: Coniacian Stage, Lower Senonian Subepoch, Lower Gulf Epoch, Late Cretaceous.

Material:

YPM 2431:

 


Locality: Southwest of Longe Butte, 1 1/2 mile-very small ravine from south, Sec. 13, T15S, R37W, Logan County, Kansas.

Horizon: Niobrara Formation.

Biostratigraphy:

Age: Coniacian Stage, Lower Senonian Subepoch, Lower Gulf Epoch, Late Cretaceous.

Material:

YPM 2440:

 


Locality: 4 miles northeast of Monument Rocks, 1 mile north of YPM 1177, Gove County, Kansas.

Horizon: Smoky Hill Member, Niobrara Formation.

Biostratigraphy:

Age: Santonian or Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

YPM 2470: 1st, 2nd, 3rd and 4th phalanx.

 

Locality: Kansas.

Horizon: Niobrara Formation.

Biostratigraphy:

Age: Coniacian Stage, Lower Senonian Subepoch, Lower Gulf Epoch, Late Cretaceous.

Material:

AMNH 5098: Fragmentary skull with possible regurgiated bolus of partially digested fish remains found between the mandibular rami (Bennett, 2001)

AMNH 138A, 139, 141, 144, 147-148, 150-153, 154, 155, 167, 168, 219, 1573, 1644, 1800, 1974, 2094, 2169, 2172-2173, 2175, 2177, 2180, 7515, 7608, 8595, 23701:

AMNH 140, 2095, 2122, 2123 “A”, 2170, 2176, 4906, KUVP 5071, 9786, 85411, 85472 “A”, LACM 1136 “A”, MCZ uncatalogued “B” , UCM F31228, YPM 1166, 2309, 2331, 2342, 2377, 2358, 2362, 2366, 2381, 2415, 2422, 2444 ,2510, 2521, 2561, 2588, 2595, 2600, 2609, 2611, 2615, 2624, 2629, 2703, 2710, 42818: Metacarpal 4ds.

AMNH 145, 2004, BMNH 4546, CM 11405, 11413, 11415, FHSM 184 “A”, FHSM 696, 2063, 2142, KUVP 944, 945, 40012, 40012 “A”, 57291, 65724, 65748, 69447, 84836, 84851, 85378, 85381, 85387, 85392, 85409, 85418, 85435, 85449, 921, 85467, 85472 “B”, LACM 51136 “B”, MCZ 279, MCZ uncatalogued “A”, MCZ uncatalogued “C”, USNM 407642, YPM 1165, 1184, 1193, 1198, 2277, 2300, 2308, 2333, 2344, 2375, 2380, 2396, 2457, 2458, 2434, 2477, 2509, 2522, 2533, 2534, 2550, 2551, 2563, 2564, 2573, 2575, 2576, 2580, 2593, 2635, 2636, 2638, 2643, 2661, 2664, 2670, 2671, 2673, 2677, 2681, 2688, 2696, 2705, 2716, 2726, 2734, 2736, 2739, 2750, 2752, 2753, 2777, 2780, 2812, 2830, 2832, 42801, 42804, 42808, 42813, 42823, 42824, 42837: Metacarpal 4d, metacarpal 4v.

AMNH 2085, BMNH 2934, FHSM 4522, 4526 “A”, FMNH P27476, FMNH UR 895, UCM F3116, USNM 407639, YPM 2266, 2278, 2417, 2459, 2511, 2566, 2665, 2607: Metacarpal 4v.

UM 12401: Metacarpal 4, 4d.

AMNH 153 “A”: Radius, and ulna.

AMNH 2174: Metacarpal 4d, metacarpal 4v, wing phalange 1.

AMNH 4907: Radius, ulna, wing phalange 2, and 3.

AMNH 4908: Humerus, radius, ulna, metacarpal 4d, metacarpal 4v, wing phalange 1, 2, femur and tibia.

AMNH 7519, UCM F747, F31159, YPM 2338, 2354, 2421, 2597, 2693: Femur.

BHI, 7 uncatalogued specimens as follows

“UNC A”: Wing phalange 3.

“UNC B”: Scapula coracoid, ulna and wing phalange 2.

“UNC C”: Skull, mandible and atlas/axis and wing phalanx 1 of an immature individual.

“UNC D”: Humerus.

“UNC E”: Metacarpal 4.

“UNC G”: Coracoid, metacarpal 4d, 4v.

“UNC I”: Coracoid and humerus.

BMNH R2928, 2930-2932, 2935-2939, 4007, 4078, 4536, 4545, and 1 uncatalogued specimen.

BMNH 2933, 2933 “A”, 4541, CM 11400-A, 11404, KUVP 980 “A”, 990, 27829, 66115, YPM 2409, 2499, 2648: Ulnare.

BMNH 2959, Right scapula, right coracoid, left humerus, radii, ulnae, both metacarpal 4, right metacarpal 4d, both metacarpal 4v, both wing phalange 1, wing phalange 2, left wing phalange 3, and both wing phalange 4.

BMNH 3299: Humerus, ulnae, metacarpal 4, metacarpal 4d, wing phalange 1, 2, femur and tibia.

BMNH 4006: Right humerus, radii, right ulnae, both metacarpal 4, right metacarpal 4d, both metacarpal 4v, both wing phalange 1, femora and tibiae.

BMNH 4008, CM 11420, KUVP 972, 56649: Humerii.

BMNH 4011, KUVP 987, 988, UM 12407, YPM 2460: Coracoid.

BMNH 4534: Scapula, coracoid, radius, ulnae, wing phalange 2, 3.

BMNH 4535, 4543: Femora and tibiae.

BMNH 4538: Humerus, radius, ulnae, metacarpal 4d, and wing phalange 4.

BMNH 4539: Humerus, ulnae.

BMNH 4542: Metacarpal 4, metacarpal 4d, metacarpal v, wing phalange 1.

BMNH 4544: Wing phalange 2, 3.

BMNH 4549: Both metacarpal 4d, 4v, and right wing phalange 1.

CM 11402-11403, 11406-11410, 11412, 11419, 11421, and 5 uncatalogued specimens.

CM 1540: Ulna and wing phalange 2.

CM 11400: Radius, metacarpal 4, 4v, wing phalange 2, 3, and femur.

CM 11411: Wing phalange 2.

CM 11416: Metacarpal 4, 4d, 4v.

CM 11418 “A”: Wing phalange 4.

CM uncatalogued “UNC A”: Ulna, metacarpal 4d, 4v, wing phalange 1, 2, 3.

CM uncatalogued “UNC B”: Metacarpal 4d.

CM uncatalogued “UNC C”: Metacarpal 4.

CM uncatalogued “UNC D”: Metacarpal 4d, 4v, wing phalange 1, 2.

FFM: 1 uncatalogued specimen, “9-1924”: Metacarpal 4d, wing phalange 2, 3, 4, femur.

FHSM 2067, 2145-2146, 2232, 2288, 4515, 5316:

FHSM 2059: Scapula, coracoid, radius, ulna.

FHSM 2062: Left scapula, both coracoids, humeri, left radius, both metacarpal 4, 4v, femora, and tibiae.

FHSM 2064: Wing phalange 1, and 2.

FHSM 8224: Wing metacarpal of an immature individual.

FHSM 2065: Notarium of a juvenile.

FHSM 2066: Metacarpal 4v.

FHSM 2066a: Wing metacarpal and first phalanx of an immature individual.

FHSM 2070: Left scapula, left humerus, radii, right ulna, right femur.

FHSM 2072, 2119: Radius, ulna, metacarpal 4d, 4v.

FHSM 2140, 2144, FMNH UC 1336, KUVP 948, 964, 981, 69446, 85472, MNHN “1908-17”, YPM 2433, 2466: Wing phalange 1s.

FHSM 2141: Wing phalange 2, femora.

FHSM 2143: Scapula, coracoid, humerus, wing phalange 2, 3.

FHSM 2147: Metacarpal 4d, 4v, wing phalange 1.

FHSM 2183: Humerus, radius, ulna, metacarpal 4d, wing phalange 1, 2, and femur.

FHSM 4526: Radius, ulna, metacarpal 4, 4d.

FHSM VP 221: Skull, notarium of a juvenile.

FMNH Pr 139, 1144, 1602:

FMNH UC 1337: Right wing phalange 3.

FMNH UC 1338: Wing phalange 4.

FMNH UC 1341 and 1 uncatalogued specimen.

KUVP 917-920, 922-926, 928-929, 931, 935, 937-940, 942, 949, 951-953, 955, 957-959, 963, 965, 969, 974, 989, 994-995, 997, 998, 16487, 27828, 28709, 40301-40302, 40304, 40335, 49164, 49417, 49502, 56648, 65725, 66137, 69448, 84844, 85027, 85372-85373, 85375, 85377, 85379-85380, 85382-85383, 85385, 85389, 85391, 85393, 85396-85398, 85400, 85402, 85404-85408, 85410, 85412-85414, 85416-85417, 85419-85426, 85431, 85437-85441, 85444, 85446, 85448, 85450-85451, 85453-85457, 85459, 85461, 85462, 85464-85466, 85468-85470, 85590:

KUVP 65742: An immature individual.

KUVP 69449, 85390 “A”, 85427: Metacarpal 4v.

KUVP 933: Wing phalange 2-4.

KUVP 930: Notarium of an immature individual.

KUVP 941: Both wing metacarpal 4d, and left carpal of an immature individual.

KUVP 954: Wing phalange 1 of an immature individual.

KUVP 961: Metacarpal 4, 4d, 4v.

KUVP 967: Tip of skull and mandible, left ulna, metacarpal 4, both metacarpal 4d, 4v, right femur.

KUVP 85436, YPM 2357, 2373, 2498, 2768: Scapula.

KUVP 968: Scapula, coracoid.

KUVP 970, UOCM F31157, USNM 407638: Wing phalange 3.

KUVP 971: Scapula, coracoid, and ulna.

KUVP 973: Radius and ulna.

KUVP 975: Ulna, metacarpal 4, 4d, 4v, wing phalange 1-4.

KUVP 978: Radius, and wing phalange 2.

KUVP 980: Right humerus, and both femora, wing phalanx of an immature individual.

KUVP 990 “A”: Humerus and ulna.

KUVP 993: Pelvic girdle of an immature individual.

KUVP 1952: Metacarpal 4, 4v, wing phalange 1-3.

KUVP 983, 991, 56650, YPM 2438: Tibiae.

KUVP 28710: Humerus and radius, metacarpal 4d, 4v of an immature individual.

KUVP 27827: Femora, tibiae.

KUVP 40303: Radius and ulna of an immature individual.

KUVP 40332: Wing phalange 1 of an immature individual.

KUVP 49398: Wing phalange 2-4.
Note: Pathological area due to necroses (BENNETT, 2003).

KUVP 49399: Humerus, tibia.

KUVP 65737: Mandible of an immature individual.

KUVP 66111: Scapulocoracoid, humerus, radius, carpal and wing metacarpal of an immature individual.

KUVP 66141: Metacarpal 4d, 4v, wing phalange 1-2.

KUVP 85371: Wing phalanx 3 of an immature individual.

KUVP 85374: Wing phalanx of an immature individual.

KUVP 85376: Humerus, wing carpal of an immature individual.

KUVP 85384: An immature individual.

KUVP 85386: Metacarpal 4d, 4v of an immature individual.

KUVP 85388: Wing phalange of an immature individual.

KUVP 85395: An immature individual.

KUVP 85399: Wing phalange 1-2.

KUVP 85401: Wing phalanx 1 of an immature individual.

KUVP 85403: Wing phalanx of an immature individual.

KUVP 85415: An immature individual.

KUVP 85432: Humerus of an immature individual.

KUVP 85433: Ulna of an immature individual.

KUVP 85434: Humerus, wing phalange 1, wing phalanx of an immature individual.

KUVP 85447: Wing phalanx of an immature individual.

KUVP 85452: Ulna, carpal, wingmetacarpal, wing phalange of an immature individual.

KUVP 85458: Humerus and carpal of an immature individual.

KUVP 85460: Scapula, coracoid, humerus, radius, ulna, metacarpal 4, 4d.

LACM 50924-50925, 5119, 51132-51134, 51138, 51760, 51762 and 2 uncatalogued specimens.

LACM 50927: Metacarpal 4d, femur.

LACM 1131, KUVP 49400, YPM 42797: Radius.

LACM 1131 “A”: Wing phalange 1, 3.

LACM 1136: Metacarpal 4, 4d, wing phalange 2.

LACM uncatalogued “A”: Radius, ulna, metacarpal 4, wing phalanges 1-4.

LACM uncatalogued “B”: Wing phalange 2.

MCZ 3815: Radius, ulna, metacarpal 4, 4d, wing phalanges 1-4, femur.

MCZ 3815 “A”: Humerus, ulna, radius, metacarpal 4v, wing phalanges 1-4.

MCZ uncatalogued “D”: Metacarpal 4d, wing phalange 2.

MCZ uncatalogued “E”: and 5 uncatalogued specimens.

MNHN uncatalogued:

ROM 26105: Scapulacoracoid, wing metacarpal, wing phalange, femur and tibia of an immature individual.

ROM 26106: Wing metacarpal and wing phalange 1 of an immature individual.

ROM 26107-26100:

SMNS 55056: Wing phalange 2-3.

SMNS uncatalogued “A”: Metacarpal 4, 4d, 4v.

UALVP 24239: Metacarpal 4, 4v, wing phalange 1-4.

UM 3222: Tibia of an immature individual.

UM 12396: Humerus of an immature individual.

UM 12400: Wing phalanx of an immature individual.

UM 12403: Mandible, wing metacarpal 4d, 4v of an immature individual.

UM 12404 “A”: Ulna, metacarpal 4, 4d, 4v, wing phalange 1-2, 4.

UM 12404 “B”: Radius, ulna, metacarpal 4, 4d, wing phalange 1-2.

UM 3611: Both metacarpal 4d, left metacarpal 4v.

UM 9074, 9075, 9083, 12397-12398, 12402, 12404:

UM V 56571:

UNSM 50128: Notarium, synsacrum, scapulacoracoid, humerus, pelvis and femur of an immature individual.

USNM 50129: Scapulacoracoid, humerus, carpal, wing carpal and wing phalanx of an immature individual.

USNM 50036: Humerii, right ulna, right metacarpal 4d, wing phalange 1-2, 4, femora, and tibia.

USNM 50037:

USNM uncatalogued “A”: Scapula, coracoid, wing phalange 4, femora, tibia.

UOCM F 746, 31158, 31162, 31203, 31224, 31231-31233:

USNM 9050, 11642, 13804, 16837, 20621, 20711, 407636-407637, 407640-407641, 407643-407644:

USNM 13868: Skull. 5 cm.

USNM 6075: Humerus, metacarpal 4, wing phalanges 1-4.

USNM 407634: Humerus, metacarpal 4d, 4v, wing phalange 3-4.

USNM 407635: Radius, ulna.

YPM 2489: Middle of snout, sacrum, 5 caudal vertebrae, right femur, tibia, nearly complete pes, metacarpal 4d, 4v of an immature individual.

YPM 1162, 1180, 1183, 1186, 1188, 1189, 1191, 1194-1197, 1199, 1455, 2264-2265, 2270, 2272, 2274-2275, 2279-2286, 2288, 2301, 2304-2305, 2307, 2311, 2320, 2332, 2336-2337, 2339-2340, 2343, 2346, 2353, 2355-2356, 2359-2361, 2372, 2374, 2376, 2378, 2390, 2399, 2401-2402, 2404-2408, 2410, 2412, 2416, 2418-2420, 2423-2424, 2426-2427, 2429-2430, 2435-2437,2439, 2442-2443, 2445-2447, 2449-2450, 2456, 2469. 2471, 2475, 2481, 2483-2486, 2488. 2500, 2503-2504, 2506, 2513, 2515, 2524, 2526, 2528-2531, 2535-2543, 2545, 2547-2549, 2552,2565, 2567-2568, 2571-2572, 2574, 2577-2579, 2581, 2583-2587, 2592, 2596, 2598-2599, 2599+, 2601-2602, 2604, 2608, 2610, 2612-2614, 2617-2620, 2623, 2627, 2630, 2637, 2639-2642, 2647, 2649-2652, 2654-2656, 2658-2659, 2666-2669, 2674, 2676, 2678, 2682, 2684, 2687, 2694, 2698-2700, 2704, 2706, 2713-2715, 2720-2725, 2728+, 2728 3/4, 2731-2732, 2735, 2737, 2740, 2743-2746, 2748-2749, 2751, 2758-2761, 2769, 2771-2773, 2775, 2779, 2781-2784, 2787-2790, 2810-2811, 2813, 42786-42794, 42796, 42798-42800, 42802-42803, 42805-42807, 42809, 42811-42812, 42814-42815, 42817, 42820-42822, 42825,42827, 42829, 42836, “UNC 2-8:

Seven uncatalogued specimens in private collections.

YPM 2269: Both metacarpal 4d, 4v.

YPM 2363, 2468, 2675, 2778: Wing phalange 3.

YPM 2271: Wing metacarpal of an immature individual.

YPM 2273: Metacarpal 4d, wing phalange 1.

YPM 2276: Wing metacarpal of an immature individual.

YPM 2289: Wing phalanx of an immature individual.

YPM 2306: Wing phalanx of an immature individual.

YPM 2310: Ulna, metacarpal 4d, 4v.
Note: Metacarpal with a small oval area of pathological bone due to necroses (BENNETT, 2003).

YPM 2345: Ulna, wing metacarpal, wing phalange 2, 4 of an immature individual.

YPM 2347: Ulna, carpal, wing metacarpal, wing phalange 1 of an immature individual.

YPM 2348: Metacarpal 4d, 4v, wing phalange 3-4.

YPM 2350: Metacarpal 4d, 4v, wing phalange 1.

YPM 2352: Scapulacoracoid, humerus, ulna, carpal, wing metacarpal, femur, tibia of an immature individual.

YPM 2364: Scapula, coracoid, humerus, ulna.

YPM 2371: Wing metacarpal, wing phalange 1 of an immature individual.

YPM 2379: Scapula of an immature individual.

YPM 2391: Wing phalange 1, wing phalanx of an immature individual.

YPM 2400: Humerus, metacarpal 4d, 4v.

YPM 2403: Wing phalange 2-4.

YPM 2413: Ulna carpal, wing metacarpal, and wing phalanx of an immature individual.

YPM 2414: Ulna, metacarpal 4d, 4v, wing phalange 1.

YPM 2432: Skull of an immature individual.

YPM 2441: Humerus, metacarpal 4d, 4v.

YPM 2448: Wing phalanx of an immature individual.

YPM 2453: Radius of an immature individual.

YPM 2454: Wing metacarpal of an immature individual.

YPM 2455: Humerus of an immature individual.

YPM 2467: Metacarpal 4d, 4v, wing phalange 1-3.

YPM 2474: Wing phalange 2-4.

YPM 2479: Ulna of an immature individual.

YPM 2480: Wing metacarpal and wing phalange 1 of an immature individual.

YPM 2490: Radius, ulna, carpal, wing metacarpal and wing phalange 1 of an immature individual.

YPM 2492: Humerus, carpal of an immature individual.

YPM 2497: Ulna and wing phalanx of an immature individual.

YPM 2501: Wing metacarpal, and wing phalange of an immature individual.

YPM 2505: Wing metacarpal of an immature individual.

YPM 2507: Radius, and wing phalange 1.

YPM 2508: Ulna of an immature individual.

YPM 2514: Scapulacoracoid of an immature individual.

YPM 2520: Humerus, ulna, metacarpal 4d, 4d, 4v, wing phalange 1.

YPM 2525: Skull, mandible, atlas-axis, scapulacoracoid and humerus of an immature individual.

YPM 2532: Humerus of an immature individual of an immature individual.

YPM 2544: Ulna, wing metacarpal and phalange 1 of an immature individual.

YPM 2553: Both metacarpal 4d, right 4v.

YPM 2562: Ulna, metacarpal 4, 4d.

YPM 2582: Wing metacarpal of an immature individual.

YPM 2589: Scapulacoracoid, humerus, ulna, wing metacarpal, wing phalange 1 of an immature individual..

YPM 2605: Wing phalange 1 of an immature individual.

YPM 2606: Humerus, metacarpal 4d.

YPM 2644: Metacarpal 4d, 4v of an immature individual.

YPM 2645: Femur and tibia.

YPM 2653, 2657, 2663: Wing phalange 1 of an immature individuals.

YPM 2662: Wing phalange 3-4.

YPM 2672: Wing metacarpal of an immature individual.

YPM 2679: Wing metacarpal of an immature individual.

YPM 2680: Radius, ulna, metacarpal 4d, 4v. wing phalange 3-4.

YPM 2685: Wing phalange 2 of an immature individual..

YPM 2689: Metacarpal 4d, 4v, wing phalange 1.

YPM 2690: Metacarpal 4d, 4v, wing phalange 3.

YPM 2695: Wing phalanx. of an immature individual.

YPM 2697: Metacarpal 4d, 4v, wing phalange 1.

YPM 2701: Scapulacoracoid and humerus of an immature individual.

YPM 2707: Skull of an immature individual.

YPM 2708: Wing phalange 1, 3.

YPM 2712, 2733: Metacarpal 4, 4d, 4v.

YPM 2728: Wing phalange 4.

YPM 2730: Right humerus, ulna, metacarpal 4v, wing phalange 2.

YPM 2741: Wing phalanx of an immature individual.

YPM 2742: Wing metacarpal, wing phalange 1 of an immature individual.

YPM 2747: Wing metacarpal of an immature individual.

YPM 2754: Wing phalange 1 and wing phalanx of an immature individual.

YPM 2755: Carpal of an immature individual.

YPM 2756: Wing phalanx of an immature individual.

YPM 2757: Wing phalanx of an immature individual.

YPM 2767: Right metacarpal 4, 4d, 4v, wing phalange 3, and both femora.

YPM 2776: Metacarpal 4.

YPM 2785: Wing metacarpal, wing phalange 1 of an immature individual.

YPM 2786: Wing metacarpal of an immature individual.

YPM 2799, 2834: Metacarpal 4, 4d, 4v.

YPM 2833: Metacarpal 4, 4d, 4v, wing phalange 1-3.

YPM 42795: Metacarpal 4d, 4v of an immature individual.

YPM 42810: Wing metacarpal, wing phalange 1, femur and tibia, metacarpal 4d, 4v of an immature individual.

YPM 42816: Femur, tibia.

YPM 42819: Radius, metacarpal 4, 4d, 4v.

YPM 42835: Metacarpal 4d, 4v, femur.

BENNETT, 2003

FMNH UC 609: Femur and pedal phalanges.
Note: First pedal phalanx has healed fracture.

FMNH uncatalogued: 4th wing phalange.
Note: Healed fracture.

KUVP 60617: Tibia.
Note: A large bony overgrowth on the anterolateral surface.

UM 12399: Mandible.
Note: Necrotic pit on the inferior margin of the left mandibular ramus.

YPM 1192: Tibia.
Note: A large bony overgrowth on the anterolateral surface.

YPM 2487: Mandibular ramus.
Note: Small overgrowth on the lingual side of the left mandibular ramus.

YPM 2727: Fragmentary specimen.
Note: Large area of pathological bone on the ventral side of a right first or second wing phalanx due to necroses.

YPM 2686: Badly deformed articular end.
Note: Osteorthritis.

YPM 2711: Pathology on the distal syncarpal.
Note: Osteorthritis.

 

Horizon: Specimens found in the Pierre Shale.

Biostratigraphy:

Age: Late Cretaceous.

Material:

AMNH 5819: Humerus.

AMNH 5840: Metacarpal 4d, wing phalange 1-3.

BHI Uncatalogued “UNC C”: Metacarpal 4d, wing phalange 1 of an immature individual.

CU 45062: Both coracoids, humerii, radii, ulnae, right metacarpal 4, both metacarpal 4d, wing phalanges 103, femora, tibiae.

CU 47280:

FMNH PR 464: Radius, ulna.

FMNH PR 468: Wing phalange 2.

FMNH PR 469, FMNH PR 1332, KUVP 50014: Metacarpal 4d.

FMNH PR 470, 676:

KUVP 27821, 50011, 50013, 50015-50019, 50089, 85445:

KUVP 36116, 33564: Metacarpal 4d, 4v.

LACM 126163, 127314:

USNM 18266: Metacarpal 4d, 4v.

 

EATON, 1910

Locality: Kansas.

Horizon: Smoky Hill Member, Niobrara Formation.

Biostratigraphy:

Age: Santonian or Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

YPM 1181: Right scapula, coracoid and bones of right wing.

YPM 2302: Left humerus.

YPM 2428: Digits II, III, IV of the right manus.

YPM 2451: Notarial vertebrae, 3 dorsal vertebrae and the anterior sacral vertebrae, humerus, metacarpal 4d, wing phalange 1, femur and tibia.

YPM 2452: Right ulna, right radius, left metacarpal V, 1st phalanx of the left digit V, 2nd phalanx, 3rd phalanx and 4th or terminal phalanx.

YPM 2472: Prepubis, metacarpal 4, 4d, 4v.

YPM 2478: Mid-section of skull and dentary.

YPM 2493: Distal ends of metacarpals, II, III, IV, right and left femora, right and left tibia, right coracoid, humerus, radius, ulna, metacarpal 4, 4d, wing phalange 1, 2, both femoria, tibiae.

YPM 2502: Distal end of right tibia and podial bones.

YPM 2512: Left capulacoracoid and humerus of an immature individual.

YPM 2546: Sternum and caudal vertebrae.

YPM 2570: Sacrum of an immature individual.

YPM 2591: 3rd and 4th phalanges of the 5th digit.
Note: 4th wing phalanx with a pathological lump due to necroses (BENNETT, 2003).

YPM 2616: 4 dorsal vertebrae, fragmentary sacrum, anterior portion of the sternum, left scapula and coracoid, humerus, radius, ulna, metacarpal 4d, 4v, wing phalange 1, 2, femur of an immature individual.

YPM 2628: Adjacent ends of metacarpal V and the 1st phalanx of the left wing.

YPM 2660: Conjoined fragments of the left metacarpal V and the 1st phalanx.

YPM 2683: Right carpus.

YPM 2692: 7th, 8th and 9th cervical vertebrae, right cervical rib, notarium, sternum, metacarpal 4d, 4v, and wing phalange 1.

YPM 2709: Left humerus.

YPM 2738: Skull, scapulacoracoid, right wing phalange 1, 2 and wing phalanx, both femora and tibiae of an immature individual.

YPM 2774: Distal end of left metacarpal and 1st, 2nd and 3rd phalanx of digit V.

YPM 2831: Left humerus.

 

= Pteranodon cf. longiceps EHRET & HARRELL, 2018

Locality: Harrell Station Paleontology site (HSPS), near Marion Junction, Dallas County, Alabama.

Horizon: Unnamed Member, Mooreville Chalk.

Biostratigraphy:

Age: Lower Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

ALMNH 8630: Wing phalanx dibit IV.

Note: Toothmarks from a Saurodon leanus?

 

HARGRAVE, 2007

Locality: SDSM Locality V614, Brown Ranch Locality, Pennington County, South Dakota.

Horizon: 0.74 m below the base of the 2nd bentonite stringer, Sharon Springs Formation, Lower Pierre Shale Group.

Biostratigraphy:

Age: Lower Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

SDSM 69042: 2 manual phalanges.

 

Locality: SDSM Locality V8121, Hesperornis Locality, Custer County, South Dakota.

Horizon: Sharon Springs Formation, Lower Pierre Shale Group.

Biostratigraphy:

Age: Lower Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

SDSM 51141: Proximal end of a right femur.

 

Locality: SDSM locality V491, Henderson Ranch, Fall River County, South Dakota.

Horizon: Sharon Springs Formation, Lower Pierre Shale Group.

Biostratigraphy:

Age: Lower Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

SM 497: Femur, tibia, wing metacarpal, wing phalange 1, and wing phalanage of an immature individual.

SDSM 56128: Radius, metacarpal, metacarpal IV, 2nd and 3rd wing phalanges, and wing fragments.

 

Locality: SDSM Locality V491, Henderson Ranch, Fall River County, South Dakota.

Horizon: 0.70 m above the Ardmore succession, Gammon Ferruginous Formation, Lowest Pierre Shale Group.

Biostratigraphy:

Age: Lowermost Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

SDSM 69041: 2 humeri.

 

 

Locality: SDSM locality V926, Slurp Flats, Fall River County, South Dakota.

Horizon: 0.61-0.91 m abve the Ardmore succession in the Sharon Springs Formation, Lower Pierre Shale Group.

Biostratigraphy:

Age: Lower Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

SDSM 45719: A vertebra, scapula-coracoid, sternal fragment, humerus, 2 ulnae, 1 complete carpal, 3 distal carpals, a ?lateral carpal, metacarpa IV, distal metacarpal adn carpal, ?metacarpal, 6 manual phalanges, femur, 5 metatarsi and fragments, pedla phalanx, ?ungual and unidentified fragments.

Note: 2 vertebrae of Enchodus sp were found in the stomach area?

 

Locality: SDSM Locality V938, Whitley's Wash, Fall River County, South Dakota.

Horizon: Near the 2nd stringer in the Sharon Springs Formation, Lower Pierre Shale Group.

Biostratigraphy:

Age: Lower Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

SDSM 56224: Partial dentary, basicranium, 11 skull fragments, and 4 wing fragments.

SDSM 56225: 5 wing fragments, 4 with articular surfaces.

 

Locality: SDSM Locality V8947, Conger Ranch, Fall River County, South Dakota.

Horizon: In the concretionary layer of the Sharon Springs Formation, Lower Pierre Shale Group.

Biostratigraphy:

Age: Lower Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

SDSM 69044: A complete metacarpal IV.

 

Locality: SDSM Locality V8948, Conger Ranch, Fall River County, South Dakota.

Horizon: 1.83 m below the bentonite couplete in Sharon Springs Formation, Lower Pierre Shale Group.

Biostratigraphy:

Age: Lower Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

SDSM 69046: 2 wing elements without articular surfaces.

 

Locality: SDSM Locality V8949, Conger Ranch, Fall River County, South Dakota.

Horizon: 0.61 below the Ardmore Sussionsinn the Gammon ferruginous Formation, Lowest Pierre Shale Group.

Biostratigraphy:

Age: Lowermost Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

SDSM 69045: Proximal humerus, ulna, radius, and proximal metacarpal.

 

Locality: SDSM Locality V9115, Wallace Ranch Locality 1, Fall River County, South Dakota.

Horizon: Just above the Ardmore Sucession in the basal Sharon Springs Formation, Lower Pierre Shale Group.

Biostratigraphy:

Age: Lower Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

SDSM 69038: A partial wing element.

 

Locality: SDSM Locality V9116, Wallace Ranch Locality 2, Fall River County, South Dakota.

Horizon: ~3.0-4.6m below the Ardmore succession, Gammon Ferruginous Formation, Lowest Pierre Shale Group.

Biostratigraphy:

Age: Lowermost Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

SDSM 56093: 5 wing fragments.

 

Locality: SDSM Locality V9116, Wallace Ranch Locality 2, Fall River County, South Dakota.

Horizon: 0.91 m above the 2nd bentonite stringer in the Upper Sharon Springs Formation, Lower Pierre Shale Group.

Biostratigraphy:

Age: Lower Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

SDSM 69039: 5 wing fragments.

 

Locality: SDSM Locality V9116, Wallace Ranch Locality 2, Fall River County, South Dakota.

Horizon: 0.76 m above 2nd bentonite stringer in the Upper Sharon Springs Formation, Lower Pierre Shale Group.

Biostratigraphy:

Age: Lower Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

SDSM 69040: A jaw fragment, ?carpal, distal phalanx.

Note: Fish vertebrae from Enchodus was found in the stomach area.

 

Locality: SDSM Locality V9162, Alum Creek Locality 1, Fall River County, South Dakota.

Horizon: Upper part of the Gammon Ferruginous Formation, Lowest Pierre Shale Group.

Biostratigraphy:

Age: Lowermost Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

SDSM 69043: 2 partial wing elements.

 

Locality: Mule Creek, Niobrara County, near the South Dakota border, Wyoming.

Horizon: Sharon Springs Formation, Lower Pierre Shale Group.

Biostratigraphy:

Age: Lower Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

SDSM 5817: Atlas-axis, anterior end of cervical vertebra, wing fragments, and left knee joint, consisting of the interlocking distal femur and proximal tibia.

SDSM 5818: Posterior end of right ramus, distal end of ulna, left proximal carpal, left distal carpal, left distal metacarpal IV, right distal metacarpal, right proximal phalanx 1 and incomplete elements.

SDSM 54358: Ulna, partial pubis, femora and tibiae.

 

 

HOWSE, 1986

Locality:?

Horizon:?

Biostratigraphy:

Age:

Material:

BMNH R2929, R4537: Postaxial cervicals.

 


Locality: Niobrara Chalk, Kansas.

Horizon: Smoky Hill Member, Niobrara Formation.

Biostratigraphy:

Age: Santonian or Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

Number: Not given: Skull missing the back of skull, mandible, ?cervical vertebra, 2 dorsal vertebrae, scapula and coracoid, humerus, ulna, phalange IV, femur, tibia and fibula.

 

Locality:

Horizon:

Biostratigraphy:

Age:

Material:

YPM 2626: Humerus, radius, ulna, phalanx 2, femur.

 

BENNETT, 1987

Locality: Western Kansas.

Horizon: Smoky Hill Member, Niobrara Formation.

Biostratigraphy:

Age: Santonian or Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

YPM 2462: 2nd, 5th, 7th-10th caudals, distal caudal rod.

AMNH 6158: Left coracoid, humerii, radii, ulnae, left wing phalange 1, 2, 3, 4, femur and tibia, caudal vertebrae.

 

WELLNHOFER, 1980

Locality: Kansas.

Horizon: Upper Smoky Hill Member, Niobrara Formation.

Biostratigraphy:

Age: Santonian or Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

YPM 2476: Left quadrate and articular.

 

Locality: South Dakota.

Horizon: Pierre Shale.

Biostratigraphy:

Age: Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

AMNH 5840A: Wing phalange 2.

AMNH 5841: Metacarpal 4d.

 


Locality: Butte Creek, 1 mile southeast of Long Butte, T15S, R37W, Logan County, Kansas.

Horizon: Smoky Hill Member, Niobrara Formation.

Biostratigraphy:

Age: Santonian or Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

YPM 2425: Left humerus, radius, ulna, the proximal distal carpals the metacarpals, pteroid bone, fragmentary proximal phalange, metacarpal 4, 4v, and wing phalange 1, 2.

 

= Pteranodon (Longicepia) marshi MILLER, 1971 (published in 1972)
= Pteranodon (Pteranodon) marshi (MILLER, 1971 (published in 1972)) MILLER, 1972
= Pteranodon marshi (MILLER, 1971 (published in 1972))
Etymology: In honor of Ornithel C. Marsh.

Holotype: YPM 2594

Locality: Near the Smoky (Hill) River, Wallace County, Kansas.

Horizon: Upper Smoky Hill Member, Niobrara Formation.

Biostratigraphy:

Age: Santonian or Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material: Skull, distal end of the quadrate bone of a male.
Note: Right marginal ridge of mandible shows extensive necrosis (BENNETT, 2003).



= Pteranodon (Sternbergia) walkeri MILLER, 1971 (published in 1972)
= Pteranodon walkeri (MILLER, 1971 (published in 1972))
= Pteranodon (Geosternbergia) walkeri (MILLER, 1971 (published in 1972)) MILLER, 1978
= Geosternbergia walkeri (MILLER, 1971 (published in 1972)) OLSHEVSKY, 1991
Etymology: In honor of Myrl V. Walker, of Fort Hays, Kansas.

Holotype: Sternberg Memorial Museum No. 2851

Locality: 2 miles northeast of Penokee, Sec. 13, T8S, R24W, Graham County, Kansas.

Horizon: Lower Smoky Hill Member, Niobrara Formation.

Biostratigraphy:

Age: Santonian or Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material: Skull.

 

Referred material:?

Locality: Kansas

Horizon: Upper Smoky Hill Member, Niobrara Formation.

Biostratigraphy:

Age: Santonian or Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

LACM: Skull lacking the crest. This skull is at LACM and is a very large skull; with a plaster crest it reaches 2M.