Species: brasilensis BUISONJE, 1980
Etymology: In reference to the country in which it was found, Brazil.

Holotype: GIUA M. 4894

Locality: Exact locality is not known, the provenance of the calcareous concretion which contained the holotype can be given, North-eastern exposure in the foothills of the Chapada do Araripe, Ceara State, north-eastern Brazil.

Horizon: Lower part of the Santana-Formation, Araripe Group.

Biostratigraphy:

Age: Aptian Stage, Middle Gallic Subepoch, Early Cretaceous Epoch, Early Cretaceous.

Material: Proximal part of right humerus together with the ankylosed glenoidal parts of the right scapula-coracoid.

Paratype: GIUA M 4895 (Cast MN 4737-V): 2 cervical vertebrae.

Referred material:

WELLNHOFER, BUFFETAUT & GIGASE 1983

Locality: Chapade do Araripe, Ceara State, Northeastern Brazil.

Horizon: Romualdo Member, Santana Formation, Araripe Group.

Biostratigraphy:

Age: Late Aptian or Early Albian Stage, Middle Gallic Subepoch, Upper Early Cretaceous Epoch, Early Cretaceous.

Material:

P. Gigase collection, nr. V-201: Notarium.

 

Locality: Barra do Jardim, Chapada do Araripe, Ceara State, North-east Brazil.

Horizon: Romualdo Member, Santana Formation, Araripe Group.

Biostratigraphy:

Age: Late Aptian or Early Albian Stage, Middle Gallic Subepoch, Upper Early Cretaceous Epoch, Early Cretaceous.

Material:

Bayerische Staatssammlung fur Palaontologie und historische Geologie, Munchen, BSP 1987 I 65: Right Scapula.

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Species: pricei WELLNHOFER, 1985
Etymology: In honor of Llewellyn Ivor Price (1905-1980), for his pioneer work on the pterosaurs from the Santana Formation of Chapada do Araripe.

Holotype: BSP 1980 I 122

Locality: Barra do Jardim, Chapada do Araripe, Ceara State, North-east Brazil.

Horizon: Romualdo Member, Santana Formation, Araripe Group.

Biostratigraphy:

Age: Late Aptian or Early Albian Stage, Middle Gallic Subepoch, Upper Early Cretaceous Epoch, Early Cretaceous.

Material: Left ulna, left radius, distal left carpal, left metacarpal IV and 1st left wing phalanx.

Paratypes:

BSP 1980 I 120: Ulna, radius, carpal, metacarpal IV.

BSP 1980 I 43: Humerus.

MZ A/III 522: 2 proximal phalanges of left wing finger.

Referred material:

AMNH 22552: Almost complete right wing including humerus, radius, 3 carpals, 1st and 2nd metacarpal, 3rd metacarpal, 4th metacarpal, the wing metacarpal, the pteroid bone, several phalanges of digits, I, II, and III, 3 phalanges of the wing finger, the proximal fragment of the 4th wing finger phalanx.

A cast of the type is at the Museu Nacional (MN 4738-V, Rio de Janeiro)

VELDMEIJER, 2002

SMNS 55883: Humerus.

SMNS 55410, 55411, 55413, 82001: Isolated ulnae and radii.

 

 

Locality: Chapade do Araripe, Ceara State, Northeastern Brazil.

Horizon: Romualdo Member, Santana Formation, Araripe Group.

Biostratigraphy:

Age: Late Aptian or Early Albian Stage, Middle Gallic Subepoch, Upper Early Cretaceous Epoch, Early Cretaceous.

Material:

Bayerische Staatssammlung fur Palaontologie und Hist. Geologie, Munchen, BSP 1987 I 1: Radius, ulna, distal carpus, carpal, pteroids, metacarpals and claws.

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Species: spixi WELLNHOFER, 1985
Etymology: In honor of the Bavarian Zoologist Johann Baptist von Spix (1781-1826), together with Carl Friedrich Philipp von Martius in the years 1817-1820) who discovered the fossil layers of Chapada do Araripe.

Holotype: BSP 1980 I 121

Locality: Chapade do Araripe, Ceara State, Northeastern Brazil.

Horizon: Romualdo Member, Santana Formation, Araripe Group.

Biostratigraphy:

Age: Late Aptian or Early Albian Stage, Middle Gallic Subepoch, Upper Early Cretaceous Epoch, Early Cretaceous.

Material: Distal portion of ulna and radius, left proximal carpal, left distal carpal, right distal carpal, right distal lateral carpal and left pteroid bone.