Genus: Tupandactylus KELLNER & CAMPOS, 2007
Etymology: Tupan or Tupa, god of the thunder in the Tupian culture and Greek, dactylus, "finger."
= Ingridia UNWIN & MARTIL, 2007
Etymology: In memory of Ingrid Wellnhofer, wife of Peter Wellnhofer, the leading authority on pterosaurs for much of the last four decades.

Species: imperator (CAMPOS & KELLNER, 1997) KELLNER & CAMPOS, 2007
Etymology: Latin, imperator, "emperor."
= Species Nova KELLNER & CAMPOS, 1992
= Ingridia imperator (CAMPOS & KELLNER, 1997) UNWIN & MARTIL, 2007

Note: Both Kellner, et al, 2007 and Unwin, et al., 2007 renamed Tapejara imperator. Kellner & Campos paper was publshed two months before Unwin & Martill and therefore has priority. Also, Unwin & Martill place Tapejara navigans into thier new genus Ingridia as a seperate species, though Kellner et al., 2007 referre Tapejara navigans to Tupandactylus imperator. I'm following both Unwin & Martill and place Tapejara navigans as a sperate species of Tupandactylus.

Holotype: DNPM MCT 1622-R

Locality: Near Nova Olinda, Chapada do Araripe, Ceara State, Northeastern Brazil.

Horizon: Crato Formation, Araripe Group.
Note: Was Crato Member, basal unit of the Santana Formation, Araripe Group.

Biostratigraphy:

Age: Aptian-Albian Stage, Middle Gallic Subepoch, Upper Early Cretaceous Epoch, Early Cretaceous.

Material: Nearly complete skull with a large crest.

Referred material:

PINHEIRO, FORTIER, SCHULTZ, ANDRADE, & BANTIM, 2011

CPCA 3590: Fragmentary skull and fragmentary dentary.

SAYAO & KELLNER, 1999

MN 4729-V: Articulated left wing formed by scapulacoracoid, humerus, radius, ulna, incomplete carpal, wing metacarpal McIV, metacarpal III, parts of digits I-III, and complete 4th digit.

FREY, TISCHLINGER, BUCHY & MARTILL, 2003

SMNK PAL 2839: Fragmentary skull.
Note: Has crest fibers.

UNWIN & MARTILL, 2007

Private collection: Fragmentary skull.

 

= Tupandactylus cf. imperator CINCOTTA, NICOLAI, CAMPOS, McNAMARA, D'ALBA, SHAWKEY, KISCHLAT, YANS, CARLEER, ESCUILLIE & GODEFROIT, 2022

Locality: Unknown, Chapada do Araripe, Ceara State, Northeastern Brazil.

Horizon: Crato Formation, Araripe Group.

Biostratigraphy:

Age: Aptian-Albian Stage, Middle Gallic Subepoch, Upper Early Cretaceous Epoch, Early Cretaceous.

Material:

MCT.R.1884: Partial skull and crest.

Note: With Soft tissue and feathers.

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Species: navigans (FREY, MARTILL & BUCHY, 2003b) PINHEIRO, FORTIER, SCHULTZ, ANDRADE, & BANTIM, 2011
= Genus: Nova? FREY, MARTILL & BUCHY, 2003b
= Tapejara navigans FREY, MARTILL & BUCHY, 2003b
Etymology: Latin, navigans, "sailing": pertaining to the shape of the cranial crest.
= Ingridia navigans (FREY, MARTILL & BUCHY, 2003b) UNWIN & MARTILL, 2007
= Genus: Nova MARTILL & FREY, 1998

Holotype: SMNK 2344 PAL

Locality: Araripe Basin, Ceara State, North East Brazil.

Horizon: Nova Olinda Member, Crato Formation.

Biostratigraphy:

Age: Aptian Stage, Middle Gallic Subepoch, Uppermost Early Cretaceous Epoch, Early Cretaceous.

Material: Fragmentary skull, similar to Tapejara imperator.
Note: Specimen shows a flap of skin on the front of the skull and has no crest extension behind the skull.

Referred material:

SMNK PAL 2343: Fragmentary skull.
Note: Specimen shows a flap of skin on the front of the skull and has no crest extension behind the skull.

CAMPO, PINHEIRO, NUNES, ANELLI & COSTA, 2019, BECCARI, PINHEIRO, NUNES, ANELLI, MATEUS & COSTA, 2021

GP/2E 9266: Almost complete, well-preserved and articulated material.