Genus: Megalocephalus BARKAS 1873
Etymology:
= Megacephalus HUENE, 1956 (sic)
Etymology:
= Mioganodon CASE, 1946
Etymology:
= Orthosauriscus KUHN, 1933
Etymology:
= Orthosaurus BARKAS, 1873 non St. HILAIRE, 1835 (Orthosaurus elaverensis, Crocodilia, Diplocynodontidae)
Etymology:

Species: pachycephalus (BARKAS, 1873) BARKAS 1873
Etymology:
= Orthosaurus pachycephalus BARKAS 1873
= Orthosauriscus pachycephalus (BARKAS, 1873) KUHN, 1933

Holotype: NEWHM: G015.22 (DMSW 2)

Locality: Low Main Seam, Newsham, Northumberland County, England, southern United Kingdom.

Horizon: Middle Coal Measures.

Biostratigraphy: Lower Similis-Pulchra zone.

Age: Westphalian B, Lower Upper Silesian series, Kashirskian Stage, Lower Moscovian Epoch, Early Middle Pennsylvanian, (Late Carboniferous).

Material: Nearly complete skull.

Referred material:

BEAUMONT, 1977

Locality: Jarrow Colliery, Castlecomer, Eire, Kilkenny County, Ireland.

Horizon:

Biostratigraphy: Lower Communis zone.

Age: Westphalian A, Middle Silesian Series, Upper Bashkirian – Lower Moscovian Epoch, Middle Carboniferous (Middle Pennsylvanian).

Material:

INM 187: Laterally compressed skull with lower jaw.

BMNH R8645: Juvenile skull.

 

Locality: Swanwick, Derbyshire County, England, southern United Kingdom.

Horizon: Blackshale or Silkstone seam.

Biostratigraphy:

Age: Westphalian A, Middle Silesian Series, Upper Bashkirian – Lower Moscovian Epoch, Middle Carboniferous (Middle Pennsylvanian).

Material:

KCL1: Right side of skull table and cheek region, overlain by part of lower jaw ramus.

KCL2: Orbital and interobital region, overlain by dentary.

 

Locality: Coalbrookdale, Shropshire County, England, southern United Kingdom.

Horizon: Blue Flats Ironstone, Middle Coral Measures.

Biostratigraphy: Lower Modiolaris zone.

Age: Westphalian B, Lower Upper Silesian series, Kashirskian Stage, Lower Moscovian Epoch, Early Middle Pennsylvanian, (Late Carboniferous).

Material:

BMNH R585: Skull and lower jaws.

 

Locality: Airdre or “mushet’s ‘ Blackband Ironstone, Larkhall, Strathclyde Province, Scotland, Northern United Kingdom.

Horizon: Middle Coal Measures.

Biostratigraphy: Upper Modiolaris zone.

Age: Westphalian B, Lower Upper Silesian series, Kashirskian Stage, Lower Moscovian Epoch, Early Middle Pennsylvanian, (Late Carboniferous).

Material:

HG V2016: Right jaw ramus.

 

(also from NEWMAN, McCLEAN & HUDSON, 1996)

Locality: Newsham, Northumberland County, England, southern United Kingdom.

Horizon: Low Main Seam, Middle Coal Measures.

Biostratigraphy: Lower Similis-Pulchra zone.

Age:, Westphalian B, Lower Upper Silesian series, Kashirskian Stage Lower Moscovian Epoch, Early Middle Pennsylvanian, (Late Carboniferous).

Material:

NEWHM: G015.21 (HMG 15.21, DMSW 1): Skull.

NEWHM: G015.23 (HMG 15.23, DMSW 3): Skull lacking the snout region and occiptial condyle with braincase.

NEWHM: G015.24 (DMSW 4): Skull lacking the snout anterior to the antorbital vacuities.

NEWHM: G015.25 (DMSW 5): An incomplete skull, consisting of the skull table, left quadrate ramus, left lateral half of snout, the palatal elements underlying these regions and the posterior part of the neurocranium.

NEWHM: G015.26 (DMSW 6): Left quadrate ramus with occiput and tabular region of the skull table.

R 3417: Posterior part of a large skull.
Note: Type of M. marcomma

NEWHM: G015.27 (HMG 15.27, DMSW 7): Right cheek region with epiptergyoid.

NEWHM: G015.28 (DMSW 8): Anterior part of the left maxilla, the anterior portion of the left jugal, the entirety of the left pre-frontal and the left palatine with tusk.

NEWHM: G015.29 (HMG 15.29, DMSW 9): Isolated basiocciptial, associated with palatine and other skull fragments.

NEWHM: G015.30 (HMG 15.30, DMSW 10): Middle region of snout.

NEWHM: G015.31 (DMSW 11): Incompletely preserved isolated left vomer bearing anterior tusk.

NEWHM: G015.32 (HMG 15.32, DMSW 12): Right palatine belonging to G15.23, showing posterior margin of choana and naso-lachrymal duct.

NEWHM: G015.33 (HMG 15.33, DMSW 13): Left vomer.

NEWHM: G015.34 (HMG 15.34, DMSW 14): Premaxilla, showing perforation for dentary tusk.

NEWHM: G015.35 (HMG 15.35, DMSW 15): Premaxilla, showing perforation for dentary tusk.

NEWHM: G015.38 (DMSW 18): A fragment of the central part of the length of the left mandibular ramus.

NEWHM: G015.39 (HMG 15.39, DMSW 19): Anterior region of left jaw ramus.

NEWHM: G015.40 (HMG 15.40, DMSW 20): Middle region of right jaw ramus.

NEWHM: G015.41 (DMSW 21): Length of left dentary and associated dermal bones.

NEWHM: G015.42 (DMSW 22): Articular extremity including the articular condyle of a left mandibular ramus.

NEWHM: G015.43 (HMG 15.43, DMSW 23): Middle region of right jaw ramus.

NEWHM: G015.44 (DMSW 44): Length of mid-section of a right mandibular ramus.

NEWHM: G015.46 (HMG 15.46, DMSW 26): Right jaw ramus.

NEWHM: G015.47: Right jaw articulation including the articular, quadrate and part of the pterygoid.

NEWHM: G015.48: Fragment of ornamented dermal bone bearing 3 incomplete teeth.

NEWHM: G015.49 (HMG 15.49): Snout and right orbital region.

NEWHM: G015.50: An isolated section of dermal skull roof of the snout comprising both frontals and right pre-frontal.

NEWHM: G015.51: Fragment of dentary.

NEWHM: G015.53: Fragment of dermal bone.

NEWHM: G015.54: Isolated left premaxilla.

NEWHM: G015.56: Isolated premaxilla bearing a length of lateral line sulcus.

NEWHM: G015.58: Isolated left jugal.

NEWHM: G015.59: Isolated left vomer with anterior tusk in situ.

NEWHM: G015.60: Vomerine tusk on fragment of dermal bone.

NEWHM: G015.62: Fragment of ornamented dermal bone bearing one (sectioned and polished) tooth base.

NEWHM: G015.63: The anterior 55mm of a left dentary bearing a parasymphysial tusk.

NEWHM: G015.64: 2 fragments of ornamented dermal bone.

NEWHM: G015.65, G015.66, G015.68, G086.46, G086.60, G090.28, G090.29, G090.30: Fragments of ornamented dermal bone.

NEWHM: G024.29: 2 isloated teeth.

NEWHM: G037.88: Isolated neural arch.

NEWHM: G053.30: Isolated tooth.

NEWHM: G055.56: Isolated left maxilla, lacking its anterior and posterior extremities.

NEWHM: G086.43: Slab with 2 fragments of ornamented dermal bone.

NEWHM: G086.44a-s: 19 isolated teeth.

NEWHM: G086.45a-t: 20 isolated teeth.

NEWHM: G086.62: A large left quadrate in assocation with fragments of the left quadratojugal and pterygoid.

NEWHM: G090.21: Fragment of premaxilla.

NEWHM: G140.63: Shale bearing impression of ornamented dermal bone.

NEWHM: G140.81: Isolated right jugal bone from a juvenile.

NEWHM: G141.30: Section of tooth row including the crowns of 3 successive teeth.

NEWHM: G142.95: Isolated left nasal.

NEWHM: G143.05: 12 isolated teeth.

 

Locality: Carnbroe, Strathclyde Province, Scotland, Northern United Kingdom.

Horizon: Palace Craig Ironstone, Middle Coal Measures.

Biostratigraphy: Lower Similus-Pulchra zone.

Age: Westphalian B, Lower Upper Silesian series, Kashirskian Stage, Lower Moscovian Epoch, Early Middle Pennsylvanian, (Late Carboniferous).

Material:

GS 28319, 28320: Snout and both jaw rami.

 

Locality: Longton, Staffordshire County, England, southern United Kingdom.

Horizon: Ash Ironstone, Middle Coal Measures.

Biostratigraphy: Upper Similis-Pulchra zone.

Age: Westephalian C, Middle Silesian series, Podolskian stage, Moscovian Epoch, Middle Carboniferous (Middle Pennsylvanian).

Material:

BMNH R2370: Middle region of skull.

 


Locality: Fenton, Staffordshire County, England, southern United Kingdom.

Horizon: Chalky Mine (-Golden Hill) Ironstone.

Biostratigraphy: Lower Phillipsii zone.

Age: Westephalian C, Middle Silesian series, Podolskian stage, Moscovian Epoch, Middle Carboniferous (Middle Pennsylvanian).

Material:

BMNH R2364: Skull table.

 

SMITHSON, 1985

Locality: Carbroe, near Airdrie, Strathclyde Province, Scotland, Northern United Kingdom.

Horizon: Palace Craig Ironstone.

Biostratigraphy:

Age: Westphalian B, Lower Upper Silesian series, Kashirskian Stage, Lower Moscovian Epoch, Early Middle Pennsylvanian, (Late Carboniferous).

Material:

Number: Not given:

 

Number: Not given.

Locality: Hamilton, Strathclyde Province, Scotland, Northern United Kingdom.

Horizon: Airdrie or “Mushet’s” Black Band Ironstone, Middle Coal Measures.

Biostratigraphy:

Age: Westphalian B, Lower Upper Silesian series, Kashirskian Stage, Lower Moscovian Epoch, Early Middle Pennsylvanian, (Late Carboniferous).

Material:

BMNH:

 

Locality: Sunnyside pit, Larkhall, Near Wishaw, Strathclyde Province, Scotland, Northern United Kingdom.

Horizon: Airdrie or “Muschet’s” Black Band Ironstone, Middle Coal Measures.

Biostratigraphy:

Age: Westphalian B, Lower Upper Silesian series, Kashirskian Stage, Lower Moscovian Epoch, Early Middle Pennsylvanian, (Late Carboniferous).

Material:

HMG:

 

Locality: Pirnie, near Leven, Fife Province, Scotland, Northern United Kingdom.

Horizon: Early Davids Parrot Coal, Middle Coal Measures.

Biostratigraphy:

Age: Westphalian A, Middle Silesian Series, Upper Bashkirian – Lower Moscovian Epoch, Middle Carboniferous (Middle Pennsylvanian).

Material:

Number: Not given.

 

Locality: Quarter, near Hamilton, Strathclyde Province, Scotland, Northern United Kingdom.

Horizon Airdrie or “Muschet’s” Black Bank Ironstone, Middle Coal Measures.

Biostratigraphy:

Age: Westphalian B, Lower Upper Silesian series, Kashirskian Stage, Lower Moscovian Epoch, Early Middle Pennsylvanian, (Late Carboniferous).

Material:

Number: Not given.

 


= Anthracosaurus edgei BAILY, 1884
Etymology:

Holotype:

Locality:

Horizon:

Biostratigraphy:

Age:

Material:

_____________________________________________________________________________________

Species: lineolatus (COPE, 1868) CASE, 1946
Etymology:
= Leptophractus lineolatus COPE, 1877

Holotype: AMNH 6826

Locality: Linton locality (Coal mines originally owned by the Ohio Diamond Coal Company), NE corner Sect. 13, T9N, R2W, Linton near Wellsvile, west bank near mouth of Yellow Creek, Saline Township, Jefferson County, Ohio.

Horizon: Canneloid shale underlying Upper Freeport Coal, Allegheny Group.

Biostratigraphy:

Age: Upper Westphalian D, Lower Upper Silesian series, Myachkovskian stage, Upper Moscovian Epoch, Middle Pennsylvanian (Late Carboniferous).

Material: Middle portion of skull and part of left mandibular ramus.

Referred material:

BEAUMONT, 1977

MCZ 2162: Posterior part of skull with 5 associated vertebrae, preserved in natural cast.

NHMUK (old BMNH) R2675: Part of the right side of a skull, from the frontal of the antorbital vacuity to the naris, preserved as natural cast.

AMNH 6946: Dentary.
Note: Leptophractus dentatus, MOODIE, 1916.

 

= Anisodenxis enchodus COPE, 1885
Etymology:
= Sauropleura enchodus (COPE, 1885) MOODIE, 1916
= Megalocephalus enchodus (COPE, 1885) HOOK & BAIRD, 1986

Holotype: AMNH FR 2558

Locality: Linton locality (Coal mines originally owned by the Ohio Diamond Coal Company), NE corner Sect. 13, T9N, R2W, Linton near Wellsvile, west bank near mouth of Yellow Creek, Saline Township, Jefferson County, Ohio.

Horizon: Canneloid shale underlying Upper Freeport Coal, Allegheny Group.

Biostratigraphy:

Age: Upper Westphalian D, Lower Upper Silesian series, Myachkovskian stage, Upper Moscovian Epoch, Middle Pennsylvanian (Late Carboniferous).

Material:

_____________________________________________________________________________________

Species: brevicornis (THOMSON & YOUNG, 1870) ROMER, 1947
= Pteroplax brevicornis THOMSON, & YOUNG, 1870
Etymology:
= Orthosaurus pachycephalus WATSON, 1926
Etymology:

Holotype: Two specimens destroyed in a fire at the Kilmarnock Museum in 1909 (SCHOCH & MILNER, 2014)

Locality: Quarter near Hamilton, Strathclyde, Scotland, Northern United Kingdom.

Horizon: Middle Coal Measures.

Biostratigraphy:

Age: Duckmantian Regional Stage, Bashkirian Epoch, Early Pennsylvanian Subperiod, Early Late Carbonifeorus Period.

Material: