Species: pusillus (FRAAS, 1881) SEELEY, 1882
Etymology: Latin, pusillus, "tiny."
= Simosaurus pusillus FRAAS, 1881
= Neusticosaurus (Pachypleura) pusillus (FRAAS, 1881) LYDEKKER, 1889

Holotype: BMNH R53

Locality: Kugelberg at Hoheneck-Dolomite at Hoheneck, near Ludwigsburg, Baden-Wurttemberg State, Germany.

Horizon: Lower Keuper, Hohenecker Kalk.

Biostratigraphy:

Age: Upper Ladinian Stage, Upper Mid Triassic Epoch, Middle Triassic.

Locality: Skull and fragmentary skeleton.

Paratype: BMNH Fore and hind limb.

Referred material:

Locality: Acqua del Ghiffo, Monte San Giorgio, Tessen Canton, Switzerland.

Horizon: Lower Meride Limestone, Cava Superior horizon.

Biostratigraphy:

Age: Middle Triassic.

PIMUZ: T3387, T3388, T3390, T3405, T3409, T3416, T3418, T3419, T3421, T3424, T3426, T3429, T3432, T3434, T3442, T3451, T3468, T3478, T3481, T3483, T3484, T3509, T3513-T3519, T3525-T3530, T3532-T3534, T3535a, T3535b, T3536, T3538, T3540, T3541, T3544, T3545, T3547a, T3547b, T3548a-T3548e, T3551, T3553a, T3553b, T3555, T3558-T3573a, T3775b-T3580, T3585, T3590, T3591, T3593-T3600a, T3600b-T3606, T3610-T3614, T3616-T3618, T3620-T3636a, T3636b-T3645, T3649a-T3649c, T3650a, T3650b, T3651-T3654, T3657-T3658a, T3658b-T3661, T3664, T3670-3673, T3703, T3714, T3717, T3720, T3721, T3725, T3737-T3740, T3741a-T3741e, T3743, T3745-T3748, T3750, T3751, T3756, T3757, T3761, T3763, T3765, T3767, T3777, T3779-T3782, T3782, T3784-T3786, T3790, T3792-T3796, T3798, T3802a-T3802d, T3803a-T3803d, T3805, T3852, T3905, T3907, T3909, T3912, T3934, T3936, T3937, T3938a-T3938h, T4289, T4296, T4300. Most of these are largely complete skeletons. The specimen numbers ending in a small letter indicate several skeletons on one slab.
Note: From SANDER, 1989, a single, presumably aberrant, gastroliths in two out of many specimens.

BMNH R5720, R5757-R5738. All of these are largely complete skeletons.

MCL, some uncatalogued specimens.

PMM a few uncatalogued specimens.

MCSNM: 3, 4 5, All of these are complete skeletons.

PMU R411: Skeletons without tail.

PMU R442: Skeleton without skull and anterior part of neck.

SMNS 7831: Slab with 12 partial skeletons.

SMNS 7382: 2 incomplete skeletons.

SMNS 8279: Large skeleton.

SMNS 9142: Partial skeleton.

SMNS 9748: Partial skeleton.

SMNS 9748a: Partial skeleton.

SMNS 53929: Slab with several incomplete skeletons.

SMNS 53930: Posterior trunk with proximal femora, poorly preserved.

SMNS 53931: Incomplete skeleton, poorly preserved.

SMNS 53932: Slab with three complete skeletons and nine partial skeletons.

GPIT 17968: Slab with four partial skeletons.

Five uncatalogued slabs with one to five partial skeletons.

T3392: Skull and skeleton.
Note: Paleopathology, humerus shows avascular necrosis (ROTHSCHILDS & STORRS, 2003)

 

= Neusticosaurus aff. pusillus DIEDRICH, 2013

Locality: Bad Sulza, Thuringia State, Germany.

Horizon: Lower Bad Sulza Member, Bad Sulza Formation, Upper Muschelkalk.

Biostratigraphy: atavus Biozone, Oolite beds.

Age: Middle Illyrian substage, Ansian Stage, Mid Triassic Epoch, Late Middle Triassic.

Material:

MB R.134.2, 78/328, 78/321a, b, c, 93/155, SHMBS unnumbered: Anterior teeth.

NME no. 39.78b, 39/78c, 78.500-1, 78.500-a: Dorsal centra.

 

= Neusticosaurus cf. pusillus DIEDRICH, 2015

Locality: Enodis/posseckeri bonebed, Lamerden, Hessen State, Germany.

Horizon: Upper Muscheklalk.

Biostratigraphy:

Age: Ladinian Stage, Late Mid Triassic Epoch, Late Middle Triassic.

Material:

LAM-5: Ilium.

LAM-564: Incomplete coracoid.

LAM-566: Femur.

LAM-1001: Dorsal vertebra centrum.

LAM-1653: Dorsal neural arch.

LAM-418-549, 550-561, 563, 565, 568-574, 576-589, 598, 998-1000, 1002-1246, 1475, 1661-1663,1707-1708, 1714-1725, 1747-1749:473 bones: 1 premaxillary fragment, 1 humerus, 1 coracoid fragments, 2 ilium bones, 2 femora, 403 vertebra centra, 14 nearly complete and about 26 half neural archs, 23 ribs adn fragments.

 

 

RENESTO, 2006

Locality: Unknown, Besano, Lombardia Province, North Italy.

Horizon: Lower Meride Limestone.

Biostratigraphy:

Age: Lower Ladinian Substage, Ladinian Stage, Upper Mid Triassic Epoch, Middle Triassic.

BES 273: Skull and skeleton of a juvenile.

 

= Neusticosaurus cf. N. pusillus RENESTO & PIERONI, 2013

Locality: Rasa Village, Varese Municipality, Lombardy, Northern Italy.

Horizon: Lower part of the Cunardo Formation.

Biostratigraphy:

Age: Ladinian Stage, Mid Triassic Epoch, Middle Triassic.

Material:

RIEPPEL, 1995

Locality: di Perledo near Varenna, Lombardia Province, North Italy.

Horizon: Calcre di Perledo.

Biostratigraphy:

Age: Upper Ladinian Stage, Upper Mid Triassic Epoch, Middle Triassic.

Material:

Museo Civico “Enrico Caffi”, Bergamo, Paleontological collection number B. 684: Fragmentary skeleton.

 

= Neuticosaurus cf. pusillus SCHOCH, 1999

Locality: N 49°12’52”, F9°40’52”, Autobahn near Kupferzell-Bauersbach, Hohenlohekreis, Baden-Wurttemberg State, Germany.

Horizon: Untere Grau Mergel, Upper Lettenkeuper.

Biostratigraphy:

Age: Longobardian, Upper Ladinian Stage, Upper Mid Triassic Epoch, Middle Triassic.

Material:

Number: Not given:

 

= Neusticosaurus pygmaeus FRAAS, 1896
Etymology:

Holotype:

Locality: Eglosheim, Germany.

Horizon:

Biostratigraphy:

Age:

Material: Skull and skeleton.

Referred material:

CORROY, 1924

Locality: Crainvillers, Vosges Department, France.

Horizon: Lettenkohle.

Biostratigraphy:

Age: Late Ladinian Substage, Ladinian Stage, Upper Mid Triassic Epoch, Middle Triassic.

Material:

Number: Not given: Fragmentary remains.

_____________________________________________________________________________________

Species: edwardsi (CORNALIA, 1854) SANDER, 1989
Etymology:
NOTE: The spelling of edwardsii has been spelled with one i (Cornalia, 1854; Lydekker, 1889; Peyer, 1928, 1932; Nopcsa, 1928; Zangerl, 1935) or two i’s (Broili, 1927; Nopcsa, 1928; Vialli, 1941).
= Pachypleura edwardsii CORNALIA, 1854
= Neusticosaurus (Pachypleura) edwardsii (CORNALIA, 1854) LYDEKKER, 1889
= Pachypleurosaurus (Pachypleura) edwardsii (CORNALIA, 1854) BROILI, 1927
= Pachypleurosaurus edwardsii (CORNALIA, 1854) BROILI, 1927
= Pachypleurosaurus edwardsii (CORNALIA, 1854) NOPCSA, 1928
= Pachypleurosaurus edwardsi (CORNALIA, 1854) NOPCSA, 1928

Holotype: Museo Civico di Storia Naturale uncatalogued.

Locality: Outcrops at Ca del Frate, Besano, Lombardia Province, North Italy.

Horizon: Lower Meride Limestone.

Biostratigraphy:

Age: Lower Ladinian Substage, Ladinian Stage, Upper Mid Triassic Epoch, Middle Triassic.

Material: Skull and fragmentary skeleton.

Referred material:

Locality: Alla Cascina at Monte San Giorgio, Tessen Canton, Switzerland.

Horizon:

Biostratigraphy:

Age: Middle Triassic.

Material:

PIMUZ T2466, T2810, T2811, T3407, T3425, T3427, T3430, T3435-T3440, T3447, T3450, T3454-T3455, T3457, T3459, T3460, T3702, T3708, T3711, T3719, T3732, T3733, T3735, T3736, T3749, T3758, T3766, T3769, T3769, T3775, T3776, T3778, T3797, T3806, T3910, T4015, T4055: Skulls and skeletons.

T3428: Skull and skeletons
Note: Paleopathology, 2 humeri, shows avascular necrosis (ROTHSCHILDS & STORRS, 2003)

T3935 : Skull and skeletons
Note: Paleopathology, humerus, shows avascular necrosis (ROTHSCHILDS & STORRS, 2003)

3806, 3458, 3456: Fragmentary skeletons.

3452: Skull and fragmentary skeleton.

3453: Fragmentary skull and fragmentary skeleton.

 

= Neusticosaurus cf. edwardsii DIEDRICH, 2015

Locality: Enodis/posseckeri bonebed, Lamerden, Hessen State, Germany.

Horizon: Upper Muscheklalk.

Biostratigraphy:

Age: Ladinian Stage, Late Mid Triassic Epoch, Late Middle Triassic.

Material:

LAM-562: Ilium.

LAM-563: Ischium.

LAM-630: Tooth.

LAM-639: Fibula.

LAM-1660 coll. no:

LAM-575: Thoraci rib.

LAM-1477: Dorsal enurla arch.

LAM-1494: Cervical neural arch.

LAM-1499: Dorsal vertebral centrum.

LAM-1660: Humerus shaft.

LAM-1683: Humerus shaft.

LAM-590, LAM-637, LAM-628, LAM-1761, LAM-1762, LAM-1763, LAM 1821-1828, LAM-1874-1875, LAM-1923-1918, LAM-1971: 43 bones including cranium fragments, 2 fang tooth, incomplete femur, 1 fibula, and compelte bone, vertebrae centra, neural arches, and some questionable ribs.

 

 

KUHN, 1941

Locality: Sasso Margone near Campinone, Italy.

Horizon:

Biostratigraphy:

Age: Late Triassic.

Material:

Number: Not given: Skull and skeleton of a juvenile.

 

STOCKAR & RENESTO, 2011

Locality: Bed 34, Cassina Beds, Monte San Giorgio, Tessen Canton, Switzerland.

Horizon: Lower Meride Limestone.

Biostratigraphy:

Age: Middle Ladinian Stage, Mid Triassic Epoch, Middle Triassic.

Material:

Number: Not given:

_____________________________________________________________________________________

Genus: toeplitschi (NOPSCA, 1928) RIEPPEL & LIN, 1995
Etymology:
= Psilotrachelosaurus toeplitschi NOPSCA, 1928

Holotype: LMKG, Nr. 201.

Locality: Not known but probably from Kellerberg Quarry, Stadlbachgraben, 2 km west of Toplitsch, Gailtaler Alps, Austria.

Horizon: Upper unit of the Partnach-Plattenkalk.

Biostratigraphy:

Age: Middle Ladinian Stage, Mid Triassic Epoch, Middle Triassic.

Material: Skeleton missing first few cervicals and distal end of caudal vertebrae.

Referred material:

RIEPPEL, 2000

Locality: Kopflachgraben, Gailtalter Alps, Austria.

Horizon: Partnach-Plattenkalk.

Biostratigraphy:

Age: Middle Ladinian Stage, Mid Triassic Epoch, Middle Triassic.

Material:

LMKG uncatalogued:

 

TICHY, 1998

Locality: 4.5 km WNW of the type locality, Kellerberg Quarry, Stadlbachgraben, 2 km west of Toplitsch, Gailtaler Alps, Austria.

Horizon: Upper unit of the Partnach-Plattenkalk.

Biostratigraphy:

Age: Middle Ladinian Stage, Mid Triassic Epoch, Middle Triassic.

Material:

Number: Not given: Skull and skeleton to the pelvic region.

 

WACHTLER, 2018

Locality: St. Lorenzen-Gitschtal, Eastern Alps, Switzerland.

Horizon:

Biostratigraphy:

Age: Middle Ladinian Stage, Mid Triassic Epoch, Middle Triassic.

Material:

George Kandutsch, Coll. Wachtler-Dolomhthos: Complete female specimen, fragmentary feamle specime, fragmetnary specimen, juvenile specimens.

Coll. Charly Weiss: Complete male specimen, fragmentary male specimens.

_____________________________________________________________________________________

Species: peyeri SANDER, 1989
Etymology: In honor of Prof. Bernhard Peyer.
= Pachypleurosaurus edwardsii CORNALIA (partim)

Holotype: PIMUZ T 3615

Locality: Acqua del Ghiffo, Monte San Giorgio, Tessen Canton, Switzerland.

Horizon: Livello 2 sopra (upper part of second bed), Cava Superiore horizon, Lower Meride Limestone.

Biostratigraphy:

Age: Middle Ladinian Stage, Mid Triassic Epoch, Middle Triassic.

Material: Skull and skeleton.

Referred material:

PIMUZ T3386, T3389, T3393-T3397, T3398a, T3398b, T3399, T3403,T3404, T3408, T3410-T3414, T3420, T3423, T3431, T3433, T3441, T3443-T3446, T3449, T3461-T3465, T3467, T3469, T3471, T3472, T3474-T3477, T3479, T3482, T3497, T3510, T3511, T3523, T3524, T3531, T3534, T3537, T3539, T3542, T3543, T3546, T3557, T3581-T3584, T3586-T3589, T3592, T3607, T3609, T3662, T3663, T3665, T3688, T3705, T3710, T3712, T3713, T3715, T3716, T3722, T3726, T3728, T3734, T3744, T3752-T3755, T3762, T3764, T3768, T3770, T3787-T3789, T3799, T3801, T3902, T3924, T3932, T4290-T4294, T4295, T4299: Fragmentary skull, and fragmentary skeletons.

PIMUZ T3705: Embryo skull and almost complete skeleton.

 

STOCKAR & RENESTO, 2011

Locality: Bed 34, Cassina Beds, Monte San Giorgio, Tessen Canton, Switzerland.

Horizon: Lower Meride Limestone.

Biostratigraphy:

Age: Middle Ladinian Stage, Mid Triassic Epoch, Middle Triassic.

Material:

MCSN 8076: Fragmentary skull and fragmentary skeleton.