Genus: Polycotylus COPE, 1869
Etymology: Greek, polys, “much, many” and Greek, kotyle, “hollow, cup”, + us: Very cupped vertebrae.

Species: latipinnis COPE, 1869
Etymology:

Holotype: USNM 27678

Locality: 8 km east of Fort Wallace, on the Smoky Hill River, Logan County, Kansas.

Horizon: Smoky Hill Member, Niobrara Formation.

Biostratigraphy: Hesperornis zone.

Age: Santonian or Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material: Fragmentary skull and fragmentary skeleton.
Note: Gastroliths were found (WILLISTON, 1893, WHITTLE & EVERHART, 2000)

Note: As per STORRS, 1999

Holotype: USNM 244534 and AMNH 1735

Locality: Approximately 8 km west of fort Wallace, on the plains near the Smoky Hill River, Wallace or Logan County, Kansas.

Horizon: Smoky Hill Member, Niobrara Formation.

Biostratigraphy:

Age: Santonian or Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material: 21 vertebrae, one or perhaps two at the American Museum of Natural History, a partial ilium and other pelvic fragments, and parts of a hind limb, distal end of femur, possible tibia, several tarsals, metatarsal, and phalanges. 13 of the smaller bones are at the American Museum, including the 9 figured by COPE in 1869.

Referred material:

AMNH 1735: Phalanges.

 

Paratype: YPM 1125

Locality: Along the Smoky Hill River, 22.5 km east of Fort Wallace, Logan County, Kansas.

Horizon: Smoky Hill Member, Niobrara Formation.

Biostratigraphy: Hesperornis Zone.

Age: Santonian or Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material: Nearly complete skeleton with skull fragments.

 

Locality: South Dakota.

Horizon: Sharon Springs Member, Pierre Shale.

Biostratigraphy:

Age: Upper Lower to Lower Middle Campanian Stage, Senonian Subepoch, Upper Gulf Epoch, Late Cretaceous

Material:

As per NICHOLLS, 1988

AMNH 5804, 16254, UCMP V76179/45205, UNSM 2812-66:

 

MARTIN, SCHUMACHER PARRIS & GRANDSTAFF, 1998

Locality: SDSM V9011, South Dakota.

Horizon: 1 meter below the Niobrara/Pierre contact.

Biostratigraphy:

Age: Late Cretaceous.

Material:

SDSM 23020: Nearly complete articulated skeleton.
Note: Paleopathology, humerus shows avascular necrosis (ROTHSCHILDS & STORRS, 2003)

 

O’KEEFE, 2003 (for 2002)

Locality: A ‘deep’ gully in an old field, 1.25 miles west and .75 miles north west of west Green, Greene County, Alabama.

Horizon: Unnamed lower member, Mooreville Chalk Formation.

Biostratigraphy:

Age: Early-Middle Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

FMNH PR 187, PR 1629: Nearly complete specimen, skull and skeleton.


O'KEEFE & CHIAPPE, 2011

Locality: Bonner Ranch, Logan County, Kansas.

Horizon: Sharon Springs Member, Pierre Shale.

Biostratigraphy:

Age: Upper Lower to Lower Middle Campanian Stage, Senonian Subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

LACM 129639: Skull and nearly complete skeleton.

Note: With a fragmentary skeleton of an embryo.

 

PERSSON

Locality: Maloje Serdoba, Oblast of Saratov, European Russia.

Horizon:

Biostratigraphy:

Age: Senonian Subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

Number: Not given: Vertebrae.

 

SCHUMACHER & MARTIN, 2016

Locality: SDSM locality V9011 (Conger Ranch Locality), Fall River County, Southwestern South Dakato.

Horizon: Uppermost Niobrara Formation.

Biostratigraphy:

Age: Lower Campanian Stage, Senonian Subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

SDSM 23020: Complete skull and nealry complete sksleton.

 

WILLISTON, 1903

Locality: Unknown, Smoky Hill River, east of Fort Wallace, Kansas.

Horizon: Niobrara Formation.

Biostratigraphy:

Age: Santonian or Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

KUVP 5916: Paddle.

 

Locality: Kansas.

Horizon: Niobrara Formation.

Biostratigraphy:

Age: Late Cretaceous.

Material:

Number: Not given: Caudal vertebrae and a portion of pelvis.

= Polycotylus dolichopus WILLLISTON, 1906
Etymology:

Holotype: YPM 1642

Locality: Wallace County, Kansas.
Note: Most probably from the present Logan County, STORRS, 1999.

Horizon: Member unknown, Niobrara Formation.

Biostratigraphy:

Age: Santonian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.
Note: But most probably Smoky Hill Chalk member, Santonian to Campanian, STORRS, 1999.

Material: Femur and most of the paddle.

Referred material:

YPM 1646: Humerus and some of the mesopodial and epipodial bones.

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Species: tenius HECTOR, 1874
Etymology:

Holotype: Welington Museum No. 5a

Locality: Amuri Bluff, South Island, New Zealand.

Horizon:

Biostratigraphy:

Age: Maastrichtian Stage, Upper Senonian Subepoch, Upper Gulf Epoch, Late Cretaceous.

Material: 9 large quadrate centra.
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Species: suprajurensis SAUVAGE, 1876
Etymology:

Holotype:

Locality: Boulogne-sur-mer, Pas-de-Calais Department, France.

Horizon: Kimmeridge Clay.

Biostratigraphy:

Age: Kimmeridgian Stage, Middle Malm Epoch, Late Jurassic.

Material: Propodial and fragmentary humerus.

Referred material:

SAUVAGE, 1894

Locality: Boulonnais, France.

Horizon: Late Portlandian.

Biostratigraphy:

Age: Middle Tithonian Stage, Upper Malm Epoch, Late Jurassic.

Material:

Number: Not given:

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Species: sopozkoi EFIMOV, MELESHIN, & NIKIFOROV, 2016
Etymology: In honor of G. A. Sopots'ko, a geologist, teacher, supervisor of the clug of young geologists of Orsk.

Holotype: UPM no. 247

Locality: Izhberda locality, 5 km west of the village of Izhberda, Gaiskii Distirct, 51°17'08.36"N, 57°24.57"E, Orenburge Region, Southern Urals, European Russia.

Horizon:

Biostratigraphy:

Age: Campanian Stage, Senonian Subepoch, Upper Gulf Epoch, Late Cretaceous.

Material: Occiptial part, palatal and dermal bones, posterior parts of both lower jaw rami, anterior and poterior limbs, atlas + epistropheus, 5 cervical, 10 pectoral, and 5 caudal vertebrae, fragments of pectoral girdle, neural arches, and ribs.