Genus: Clevosaurus SWINTON, 1939
Etymology:
= Dianosaurus YOUNG, 1982
Etymology:
= Glevosaurus ROBINSON, 1973
Etymology:
Species: hudsoni SWINTON, 1939
Etymology:
Syntypes: BMNH R5939
Locality: Slickstones (Cromhall) Quarry, (ST 704916), Gloucestershire (Avon?) County, England, Southern United Kingdom.
Horizon: Karstic fissures in Dinatian limestone.
Biostratigraphy:
Age: Carnian/Norian Stage, Upper Late Triassic Epoch, Late Triassic.
Material: Upper jaws portions of right and left mandibles.
Referred material:
AUP No. 11085: Left dentary.
AUP No. 11088: Left palatine from a juvenile.
AUP No. 11089: Left palatine.
AUP No. 11143: Left premaxilla.
AUP No. 11144, 11146, 11187: Maxillae.
AUP No. 11152: Right postorbital.
AUP No. 11153: Left jugal.
AUP No. 11157: Anterior fragment of a left vomer.
AUP No. 11158: Left palatine bearing a rudimentary second tooth row.
AUP No. 11169: Fragmentary articulated skull exposed in left lateral and dorsal view.
AUP No. 11171. Cervical vertebra.
AUP No. 11172: Dorsal vertebra.
AUP No. 11174: Mid-caudal vertebra.
AUP No. 11177: Right scapulocoracoid.
AUP No. 11178: Left ulna.
AUP No. 11179: Right humerus.
AUP No. 11180: Right ilium.
AUP No. 11184: Left tibia.
AUP No. 11185: Proximal end of left tibia.
AUP No. 11354: Tooth.
AUP No. 11371. 11372: 2 fragment of a juvenile right dentary.
AUP No. 11373: Juvenile right dentary.
UMZC T1264: Articulated right nasal and prefrontal, together with an anterior caudal vertebra.
UMZC T1265: Articulated frontals.
UMZC T1266: Left squamosal with articulated frontals.
UMZC T1267: Posterior fragment of a right quadrate and quadratojual.
UMZC T1268: Fragmentary paired vomers.
UMZC T1269: Articulated palate with paired palatines, pterygoids and left and right ectopterygoids.
UMZC T1270: Associated fragments of a juvenile skull including paired pterygoids, left palatine, left quadrate and the parabasisphenoid.
UMZC T1271: Articulated skeleton and skull missing the right fore and hind limbs and the tail posterior to caudal 8.
UMZC T1272: Partly preserved articulated skull exposed in ventral view.
UMZC T1273: Left femur with a pathological secondary outgrowth of the proximal head.
UMZC T1274: Left femur.
UMZC T1275: Associated right ilium and second sacral vertebra.
UMZC T1276: Left dentary.
UMZC T1277: A parabasisphenoid.
UMZC T1279: Left ilium.
BMNH R6100, R9249, R9251, R9252, R9255-R9258, R9260-R9267: Varied skeletal elements.
CHAMBI-TROWELL, WHITESIDE & BENTON, 2019
NHMUK PV R36832: A partial disarticulated skull, cervical vertbrae and fron limb.
O'BRIEN, WHITESIDE & MARSHALL, 2018
NHMUK PV R36832: Partially complete skull and skeleton.
NHMUK PV R36846: Partially complete hind limb.
= Clevosaurus aff. hudsoni FRASER, 1994
Locality: Holwell Quarry near Frome, Somerset County, England, Southern United Kingdom.
Horizon:
Biostratigraphy:
Age: Rhaetic Stage, Uppermost Late Triassic Epoch, Late Triassic or Early Jurassic.
Material:
Number: Not given: A single premaxilla.
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Species: minor FRASER & WALKDEN, 1983
Etymology:
Holotype: AUP No. 11377
Locality: Slickstones (Cromhall) Quarry, (ST 704916), Gloucestershire (Avon?) County, England, Southern United Kingdom.
Horizon: Karstic fissures in Dinatian limestone.
Biostratigraphy:
Age: Carnian/Norian Stage, Upper Late Triassic Epoch, Late Triassic.
Material: Left maxilla.
Paratype: Left palatine missing the anterior vomerine process and with a fragmented maxillary ramus.
AUP 11086, 11087, 11088: Left palatines.
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Species: petilus (YOUNG, 1982) WU, 1994
Etymology:
= Dianosaurus petilus YOUNG, 1982
Etymology:
Holotype: IVPP P 4007
Locality: Close to Dachong village, Lufeng County, Yunnan Province, Southwest China.
Horizon: Near the top of the lower part (Dull Purplish Beds), Lower Lufeng Formation.
Biostratigraphy:
Age: Lias Epoch, Early Jurassic.
Material: Skull with mandible, damaged anterior to orbits.
Referred material:
= Clevosaurus wangi WU, 1994
Etymology: In honor of Mr. Zheng-ju Wang, for his contributions to local vertebrate paleontology and Paleoanthropology during the past thirty years.Holotype: IVPP V.8271
Locality: Near Dawa village, Lufeng County, Yunnan Province, Southwest China.
Horizon: Near the middle of the Upper part (Dark Red Beds), Lower Lufeng Formation.
Biostratigraphy:
Age: Lias Epoch, Early Jurassic.
Material: An almost complete but cracked skull with mandible.
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Species: latidens FRASER, 1993
Etymology: From the Latin, latus, "broad" and Latin, dens, "tooth."
Holotype: VMNH 524
Locality: Slickstones (Cromhall) Quarry, (ST 704916), Gloucestershire (Avon?) County, England, Southern United Kingdom.
Horizon: Karstic fissures in Dinatian limestone.
Biostratigraphy:
Age: Carnian/Norian Stage, Upper Late Triassic Epoch, Late Triassic.
Material: Fragment of dentary.
Paratypes:
VMNH 525: Dentary fragments.
VMNH 526, 527, 528, 529: Maxillary fragments.
Referred specimens.
AUP 11192, 11223: Dentary fragments.
AUP 1120: Maxillary fragment.
AUP 11191: Premaxilla.
= aff. Clevosaurus latidens HECKERT, 2001
Locality: NMMNH 1430, at the head of the south fork of Home Creek north of Kalgary Locality, UTM 3701500N, 398850E, zone 14, NAD 27, Crosby County, Texas.
Horizon: Colorado City Member, Tecovas Formation, Chinle Group.
Biostratigraphy:Age: Adamanian, Latest Carnian Stage, Late Triassic Epoch, Late Triassic.
Material:
NMMNH P-26412: Dentary fragments.
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Species: mcgilli WU, 1994
Etymology: A testimonial to McGill University, Montreal, where Dr. Wu completed
his Ph. D. Training under the guidance of Professor Robert L. Carroll.
Holotype: IVPP V.8272
Locality: Near Dawa Village, Lufeng County, Yunnan Province, Southwest China.
Horizon: About the middle of the upper part (Dark Red Beds), Lower Lufeng Formation.
Biostratigraphy:
Age: Lias Epoch, Early Jurassic.
Material: A skull with mandible, but damaged in both the snout and occipital region.
Paratype: IVPP V.8273: A skull lacking the postorbital region, with the anterior
parts of the mandible.
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Species: bairdi SUES, SHUBIN & OLSEN, 1994
Etymology:
Holotype: NSM 988GF1.1
Locality: Sites K and K’, Wasson Bluff, near Parrsboro, Cumberland County, Nova Scotia Province, Canada.
Horizon: McCoy Brook Formation, Fundy Group, Newark Supergroup.
Biostratigraphy:
Age: Hettangian Stage, Lowermost Lias Epoch, Early Jurassic.
Material: An excellently preserved but slightly crushed partial skull with both dentaries in occlusion, several associated but disarticulated cranial and mandibular bones, and associated articulated phalanges of 3 digits of the right manus.
Hypodigm.
NSM 988GF4.1: Complete left dentary.
NSM 988GF3.1, 988GF32.1, 988GF33.1: Incomplete left maxillae.
NSM 990FG90.1: Nearly complete left maxilla.
MCZ 9105: Dorsoventrally crushed, incomplete skull with almost complete mandible in occlusion.
MCZ 9106: Well-preserved partial postcranial skeleton including left humerus and ulna, articulated left hind-limb and pes articulated distal portions of the right tibia and fibula and right pes, both ilia, and articulated partial vertebral column from the posterior dorsal to the distal caudal region.
MCZ 9110: Scattered palatal bones.
MCZ 9112: Incomplete left maxilla.
MCZ 9113: Incomplete left dentary and attached coronioid.
MCZ 9114: Complete right maxilla.
Locality: Sites F, Wasson Bluff, near Parrsboro, Cumberland County, Nova Scotia Province, Canada.
Horizon: McCoy Brook Formation, Fundy Group, Newark Supergroup.
Biostratigraphy:
Age: Hettangian Stage, Lowermost Lias Epoch, Early Jurassic.
Material:
NSM 988GF2.1: Incomplete left dentary.
NSM 988GF34.1: Nearly complete right maxilla.
SHUBIN, OLSEN & SUES, 1994
Locality: Between localities L and K’, Nova Scotia Province, Canada.
Horizon: McCoy Brook Formation, Fundy Group, Newark Supergroup.
Biostratigraphy:
Age: Hettangian Stage, Lowermost Lias Epoch, Early Jurassic.
Material:
Number: Not given:
Locality: Below the talus-slope on the western side of middle mini-graben, locality G, Sandstones between the two localites L and locality K, Cliff exposures of Wassons Bluff, Greenhill, Nova Scotia Province, Canada.Horizon: McCoy Brook Formation, Fundy Group, Newark Supergroup.
Biostratigraphy:
Age: Hettangian Stage, Lowermost Lias Epoch, Early Jurassic.
Material:
Number: Not given: A badly worn partial maxilla found in direct association with the gastraliths of Sauropodomorpha incertae sedis (orignally states as Ammosaurus sp.) (SUES, & OLSEN, 2016)
SUES, & OLSEN, 2016
Locality: Wasson Bluff, near Parrsboro, Cumberland County, Nova Scotia Province, Canada.
Horizon: McCoy Brook Formation, Fundy Group, Newark Supergroup.
Biostratigraphy:
Age: Hettangian Stage, Lowermost Lias Epoch, Early Jurassic.
Material:
FGM996GF050: Fragmenary skull and skeleotn.
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Species: convallis SAILA, 2005
Etymology: Latin, convallis, "enclosed valley"; chosen to describe the location
(Pant is Welsh for valley) and to indicate the material comes from a fissure-fill.
Holotype: BMNH R7530
Locality: Pant 4 fissure in Pant Quarry, Glamorgan, South Wales County, England, Southern United Kingdom.
Horizon:
Biostratigraphy:
Age: Lower Jurassic.
Material: Left premaxilla showing a short nasal process, a long maxillary process, and the fusion of the premaxillary teeth.
Referred material:
BMNH R7531, R7550: Left premaxillae.
BMNH R7532: Right premaxilla.
BMNH R7533: Left maxilla.
BMNH R7534-7537: Right maxillae.
BMNH R7538: Left palatine.
BMNH R7551: Right palatine.
BMNH R7539: Left dentary.
BMNH R7540-7541, R7552: Right dentaries.
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Species: brasiliensis BONAPARTE & SUES, 2006
Etymology: In reference to Brazil, and latin, -ensis, a suffix denoting a
place or location.
Holotype: UFRGS-PV0748T
Locality: Sesmaria do Pinhal, 200 km north-west of the city of Faxinal do Soturno, 8 km east of Candelaria, Rio Grande do Sul State, Brazil.
Horizon: Caturrita Formation.
Biostratigraphy: Icidosaur Assemblage Zone.
Age: Adamanian, Lower Coloradian age, Late Carnian-Early Norian Stage, Late Triassic Epoch, Late Triassic.
Material: A complete, slightly dorsoventrally crushed skull of an adult with mandible in articulation.
Referred material:
ARANTES, SOARES & SCHULTZ, 2009
UFRGS-PV0742T: 6 articulated dorsal vertebrae, second sacral vertebra with the first and second caudal vertebrae.
UFRGS-PV0751T: Phalange fragmens.
UFRGS-PVT0752T: Left femur.
UFRGS-PV0754T: Dorsal vertebra, 2 articulated dorsal vertebrae, right tibia.
UFRGS-PV0757T: Left fibula.
BONAPARTE, SCHULTZ, SOARES & MARTINELLI, 2010
Locality: Linha Sao Luiz Locality, an abandoned quarry located 1.5 km northwest of downtown do Soturno Faxinal, 53°30'23"E, 29°34'08"S, near Barrio Santo Antonino, Faxinal do Soturno, Rio Grande do Sul State, Brazil.
Horizon: Caturrita Formation.
Biostratigraphy: Icidosaur Assemblage Zone.
Age: Adamanian, Lower Coloradian age, Late Carnian-Early Norian Stage, Late Triassic Epoch, Late Triassic.
Material:
MCN-PV 2852: Fragmentary skull and dentary.
CHAMBI-TROWELL, MARTINELLI, WHITESIDE, de VIVAR, SOARES, SCHULTZ, GILL, BENTON, & RAYFIELD, 2021
UFSM-PV-0131: A nearly complete skull.
UFRGS-PV-0974-T: Nearly complete skull.
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Species: sectumsemper KLEIN, WHITESIDE, LUCAS, VIEGAS, & BENTON,
2015
Etymology: Latin, meaning "always cut" and allustion to the self-shaprening
teeth that remains sharp by cutting against each other and the lower jaw throught
the animals life.
Holotype: BRSUG 29381-56
Locality: Fissure 1 in Woodlease Quarry, South Gloucestershire County, England, Southern United Kingdom.
Horizon: Rhaetian Fissure fill in Lower Carboniferous Limestone.
Biostratigraphy:
Age: Rhaetian Stage, Uppermost Late Triasisc Epoch, Late Triassic.
Material: Left dentary.
Referred materal:
BRSUG 29381-1: Right pteryoid, vertebrae.
BRSUG 29381-2: Humeri, tibia.
BRSUG 29381-3: Right palatine.
BRSUG 29381-9: Righte premaxilla.
BRSUG 29381-11: Right maxilla.
BRSUG 29381-15: Right nasal.
BRSUG 29381-69: Ilium.
BRSUG 29381-85: Radial.
BRSUG 29381-87: Claw.
BRSUG 29382-16: Pubis.
BRSUG 29382-20: ulnare.
BRSUG 29382-23: Tarsometatarsal.
Note: Paleopathologies: An expanded metatarsal or metacarpal that looks tumorous, and many heald breaks. Many ribs and long bones are remoddled.
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Species: hadroprodon HSIOU, NYDAM, SIMOES, PRETTO, ONARY, MARTINELLI, LIPARINI, de VIVAR MARTINEX, SOARES, SCHULT & CALDWELL, 2019
Etymology: Greek, hadros, "large,", Greek, protos, "first," and Greek, odous, "tooth," in reference to the "big first tooth," i.e. the large tusk-like tooth of the premaxilla and the anteriormost/first dentary tooth position.
Holotype: MMACR PV-027-T
Locality: Linha Bernardion Locality, Candelaria Municipality, Parana Basin, Rio Grande do Sul State, Southern Brazil.
Horizon: Santa Maria Formation, Santa Maria Supersequence, Candelaria sequence, Rosario do Sul Group.
Biostratigraphy:
Age: Carnian Stage, Upper Late Triassic Epoch, Late Triassic.
Material: An incomplete right premaxilla, an incomplete right maxilla and right lower jaw in occlusion, and left lower jaw located below these elements.
Referred maerial:
MMACR PV-028-T: An incomplete left lower jaw.
MMACR PV-029-T: FRagment of dentary with 3 isoalted teeth.
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Species: nicholasi BHAT, DATTA, RAY & DATTA, 2023 (published 2024)
Etymology: In honor of the Britisch paleontologist Nicholas Campbell Fraser for his outstanding contributions ot the knowledge of early rhynchocephalians.
Holotype: IITKGPR570A
Locality: Near the village of Tihki, 23°56'N, 81°22'58"E, Shahdol District, Madhya Pradesh State, India.
Horizon; Tiki Formation.
Biostratigraphy:
Age: Late Carnian to early Middle Norian Stage, Late Traissic Epoch, Late Triassic.
Material: A partial symphyseal end of left dentary.
Referred material:
IITKGPR570B-IITKGPR570C: 2 partial left dentaries.
IITKGPR570D, IITKGPR571A-IITKGPR571C: 4 partial right dentaries.
IITKGPR575C: A partial right maxilla.
IITKGPR578A-IITKGPR578D: 4 partial premaxillae.