Genus: Adocus COPE, 1870
Etymology:
= Shineusemys SUKHANOV, 2000, (Unavailable name) SUKHANOV & NARMANDAKH, 2006
Etymology:

Species: beatus (LEIDY, 1868) COPE, 1870
Etymology:
= Emys beatus LEIDY, 1868

Syntypes: ANSP 9184

Locality: Green Sand of Mullica Hill, Gloucester County, New Jersey.

Horizon: Hornerstown Formation, Rancocas Group.

Biostratigraphy:

Age: Late Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material: Several plates and fragments of a carapace.

Referred material:

GALLAGHER, PARRIS & SPAMER, 1986

Locality: Crosswicks Creek, Ellisdale, Monmouth County, New Jersey.

Horizon: Marshalltown Formation, Matawan Group.

Biostratigraphy:

Age: Upper Campanian Stage, Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

NJSM 12711:

WHITE, 1972

Locality: Sewell, Gloucester County, New Jersey.

Horizon: Hornerstown Formation, Rancocas Group.

Biostratigraphy:

Age: Late Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

ANSP 15356: A nearly complete shell.

 

= Adocus pravus LEIDY
Etymology:

Holotype: AMNH (formerly RU 1. KV-9)

Locality: Tinton Falls, Monmouth County, New Jersey.

Horizon: Greensand (= Hornerstown Formation?).

Biostratigraphy:

Age: Late Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material: Most of the hyoplastron, a large part of the right hypoplastron, and the hinder ends of both xiphiplastra.

 

= Adocus punctatus MARSH, 1890
Etymology:

Holotype: YPM 780

Locality: Hornerstown, Monmouth County, New Jersey.

Horizon: Hornerstown Formation, Rancocas Group.

Biostratigraphy:

Age: Late Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material: Shell.

Referred material:

DARROUGH, PARRIS & GRANDSTAFF, 1996

Locality: Chronister farm, sec. 26, T31N, R9E, about 1 mile north of Glen Allen, Bollinger County, Missouri.

Horizon: McNairy sand Member, Ripley Formation.

Biostratigraphy: Middle-Uppermiddle Mosasaurus Acme-zone.

Age: Navesinkian age, Early Maastrichtian Stage? Upper Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

Number: Not given: Portions of carapace, plastron, and no-shell postcranialia.

 

GALLAGHER, PARRIS & SPAMER, 1986

Locality: Crosswicks Creek, Ellisdale, Monmouth County, New Jersey.

Horizon: Marshalltown Formation, Matawan Group.

Biostratigraphy:

Age: Upper Campanian Stage, Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

NJSM 12437:

 

SCHWIMMER, SANDERS, ERICKSON, & WEEMS, 2015

Locality: Site 1, Stokes Quarry, south side of County Road 407, ca. 0.25 mile (0.40 km) west of County Road 49, 2.85 miles (4.58 km) east of S.C. Route 320, USGS Florence West 7.5' quadrangle, 34°14'21.8"N, 79°30.0"W 50', Darlington County, South Carolina.

Horizon: Coachman Formation.

Biostratigraphy:

Age: Middle Campanian Stage, Senoniain Subepoch, Gulf Epoch, Late Cretaceous.

Material:

ChM PV8903: Hypoplastron fragment.

ChM PV8925: Plastron fragment.

ChM PV8930: Costal fragment.

ChM PV9129: First right peripheral.

ChM PV9135: Bridge Fragment.

ChM PV9144: Left hypoplastron fragment.

SMM P2006.1.1: 5th neural.

 

Locality: Site 8, Burches Ferry, west bank of Pee Dee River, ca. 0.75 mile (1.21 km) east of County Road 52, ca. 0.08 mile (0.19 km) northwest of junciton of Mill Branch and Pee Dee River, USGS Pamplico North 7.5' quadrangle, 34°03'50"N, 79°31'43"W, Florence County, South Carolina.

Horizon: Donoho Creek Formation/Peedee Formation.

Biostratigraphy:

Age: Early Late Campanian Stage/Early Maastrichtian Stage, Senoniain Subepoch, Gulf Epoch, Late Cretaceous.

Material:

SCSM 2005.25.1: Section of center carapace including all parts of neurals and proximal coastals, three, four, and five.

SCSM 2005.25.12: Costal fragment.

SCSM 2005.25.21: Neural.

 

= Adocus lacer HAY, 1908
Etymology:

Syntypes: ANSP 15357, 15356

Locality: New Jersey.

Horizon: Greensand.

Biostratigraphy:

Age: Late Cretaceous.

Material: Fragment of first left peripheral and 3 hinder peripherals.

Referred material:

Locality: New Jersey.

Horizon: Hornerstown Formation, Rancocas Group.

Biostratigraphy:

Age: Late Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

AMNH 1350:

 

Locality: Greensand, New Jersey.

Horizon: Hornerstown Formation, Rancocas Group.

Biostratigraphy:

Age: Late Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material:

ANSP 9698:

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Species: agilis COPE 1868
Etymology:

Holotype: AMNH 1135

Locality: Barnsboro, Gloucester County, New Jersey.

Horizon: Hornerstown Formation, Rancocas Group.

Biostratigraphy:

Age: Late Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material: Fragmentary plastron.
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Species: syntheticus COPE 1870
Etymology:

Holotype: AMNH 1466

Locality: Barnsboro, Gloucester County, New Jersey.

Horizon: Hornerstown Formation, Rancocas Group.

Biostratigraphy:

Age: Late Maastrichtian Stage, uppermost Senonian subepoch, Gulf Epoch, Late Cretaceous.

Material: Plastron lacking the entoplastron, the right epiplastron and the right hypoplastron, a peripheral bone from the bridge, 2 imperfect costals, and some smaller fragments.
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Species:? lineolatus COPE, 1874
Etymology:
= Compsemys lineolatus (COPE, 1874) COPE, 1877

Holotype: AMNH 1844

Locality: Bijou Creek, 40 miles east of Denver, Colorado.

Horizon: Arapahoe Formation, D1 sequence.

Biostratigraphy:

Age: Lancian Age, upper Maastrichtian Stage, uppermost Senonian Subepoch, Upper Upper Gulf Epoch, Late Cretaceous.

Material: 2 fragments of neural costals.

Referred material:

EBERTH, 1987

Locality: Dinosaur Provincial Park, Newell County, Alberta Province, Canada.

Horizon: Dinosaur Park Formation, upper Judith River Group.

Biostratigraphy:

Age: Judithian age, Middle Campanian Stage, middle Senonian subepoch, upper Gulf Epoch, Late Cretaceous.

Material:

Number: Not given:

 

GILMORE, 1916

Locality: 1 miles south of Fruitland, San Juan County, New Mexico.

Horizon: Fruitland Formation.

Biostratigraphy:

Age: Kirtlandian Age, Late Campanian Stage, Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

USNM 8348: 7th peripheral from the left side and other fragments.

 

KNOWLTON, 1911/LAMBE, 1902

Locality: Red Deer River, Alberta Province, Canada.

Horizon: Judith River Formation.

Biostratigraphy:

Age: Judithian age, Campanian Stage, Senonian subepoch, Upper Gulf epoch, Late Cretaceous.

Material:

AMNH 6105: Third or fifth costal.

 

/BROWN

Locality: Montana.

Horizon: Hell Creek Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

Number: Not given:

 

Locality: “Ceratops beds”, Converse County, Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

Number: Not given:

HAY, 1908

Locality: Montana.

Horizon: Hell Creek Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

AMNH 1014: Fragment of a costal, peripheral, the second of the left side.

 

Locality: Converse County, Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

AMNH 6107: Fragment of a costal plate and another of a plastral bone.

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Species: vigoratus HAY, 1910
Etymology:

Holotype: USNM 6554

Locality: Near Ojo Alamo, San Juan Basin, San Juan County, New Mexico.

Horizon: Naashoibito Member, Ojo Alamo Formation.

Biostratigraphy: Alamo Wash Local Fauna.

Age: Late-early or earliest Late Maastrichtian Stage, Uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material: First neural, first left peripheral, seventh left peripheral, section of carapace from the free border to the border that articulated with the free costal, section of the free borer of the base of the hind lobe.
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Species: bossi GILMORE, 1919
Etymology:

Holotype: USNM 8613

Locality: Head of Ah-shi-sle-pah Wash, 2 3/4 miles south of west of Kimbetoh, in sec. 3, T 22N, R 10W, San Juan County, New Mexico.

Horizon: Hunter Wash Local Fauna, Lower Kirtland Formation.

Biostratigraphy:

Age: Kirtlandian age, Upper Campanian Stage- Lower Maastrichtian Stage, Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material: Carapace and plastron, the former lacking the greater part of the costals, all the peripherals of the right side, and peripherals 4, 5, 9, and 10 of the left side.

Paratype:

Referred material:

DALMA & LUCAS, 2016

Locality: Unknown, San Juan Basin, New Mexico.

Horizon: Probably De-Na-zin Member, Kirkland Formation.

Biostratigraphy:

Age: Kirtlandian age, Upper Campanian Stage- Lower Maastrichtian Stage, Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

ACM 7979: Complete carapace.

 

GILMORE, 1935

Locality: 3 miles northeast of Hunter's Store (Bisti P. O.), SW 1/4, T24N, R13W, San Juan Basin, San Juan County, New Mexico.

Horizon: Lower Kirtland Formation.

Biostratigraphy:

Age: Kirtlandian age, Upper Campanian Stage- Lower Maastrichtian Stage, Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

USNM 12383, 12842, 12982, 12983: Nearly complete shells and except for being flattened are in a good state of preservation.

 

Locality: Meyers Creek, San Juan County, New Mexico.

Horizon: Hunter Wash Formation, Kirtland Formation.

Biostratigraphy:

Age: Kirtlandian Age, Late Campanian Stage, Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

USNM 11326: Fragmentary shell.

 

SULLIVAN, JASINSKI, & LUCAS, 2013

Locality: New Mexico.

Horizon: De-Na-Zin, Kirtland Formation.

Biostratigraphy:

Age: Kirtlandian Age, Late Campanian Stage, Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

USNM 11082: Carapace and plastron.

 

Locality: New Mexico.

Horizon: Hunter Wash, Kirtland Formation.

Biostratigraphy:

Age: Kirtlandian Age, Late Campanian Stage, Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

USNM 12838, 12842, 12982: Carapace and plastron.

USNM 12983: Carapace, plastron, and pelvis.

 

 

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Species: kirtlandius GILMORE, 1919
Etymology:

Holotype: USNM 8593

Locality: South side of Ah-shi-sle-pahWash, 3 1/2 miles south of west of Kimbetoh, near the south side of sec. 4, T. 22N, R 10W, San Juan County, New Mexico.

Horizon: Hunter Wash Local Fauna, Fruitland Formation.

Biostratigraphy:

Age: Kirtlandian Age, Late Campanian Stage, Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material: Fragmentary carapace and plastron.

Referred material:

Locality: San Juan Basin, New Mexico.

Horizon: Lower Kirtland Formation.

Biostratigraphy:

Age: Kirtlandian age, Upper Campanian Stage- Lower Maastrichtian Stage, Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

Number: Not given:

 

LICHTIG & LUCAS, 2023

NMMNH P-78110: Carapace, only missing the portion odorsal to the pelvis, an incomplete plastron, missing the right edge of the anterior lobe, the left humerus, a mostly complte pelvis, a cervical vertebra, and a coracoid.

NMMNH P-8291: An incompletecarapace and plasteron.

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Species: planicostatus RIABININ, 1930
Etymology:

Holotype:

Locality: Jiayin, Heilongjiang Province, China.

Horizon:

Biostratigraphy:

Age: Late Cretaceous.

Material: A single last costal plate and 2 broken neurals.
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Species: foveatus NESOV & KHOSATSKY in KHOSATZKY & NESOV, 1977
Etymology:

Holotype: ZIN OHT F64-11

Locality: Kansai Locality, Fergana Depression, Tajikistan.

Horizon: Yalovach Formation.

Biostratigraphy:

Age: Santonian Stage, Senoniain Subepoch, Gulf Epoch, Late Cretaceous.

Material: Left peripheral 4.

Referred material:

SYROMYATNIKOVA & DANILOV, 2009

ZIN PH 590/64: Costal 5.

ZIN PH 591/64: Costals 7 + 8.

ZIN PH 593/64: Undeterimend costal fragment.

ZIN PH 592/64: Entoplastron.

ZIN PH 582-588/84: Undetermined shell fragments.

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Species: kizylkumensis NESOV, 1981
Etymology:

Holotype: ZINPHT S75-15

Locality: Khodzhakul I (Khodzhakul locality), Chodzhakul Lake Locality, western Sultanuvais Ridge, Uzbekistan.

Horizon: Lower or middle Khodzhakul (Chodzhakul) Formation.

Biostratigraphy:

Age: Early Cenomanian Stage, Uppermiddle Gallic Subepoch, Lowermost Gulf Epoch, Middle Cretaceous.

Material: Lateral part of undetermined costal.

Referred material:

SYROMYATNIKOVA & DANILOV, 2009

ZIN PH 9/87: Bridge peripheral.

 

Locality: Khodzhakul I (Khodzhakul locality), Khodzhakulsay localites and Sheikhdzheli II (Sheikhdezheiil), Chodzhakul Lake Locality, western Sultanuvais Ridge, Uzbekistan.

Note: No breakdown stateing which specimen went to which locality.

Horizon: Lower or middle Khodzhakul (Chodzhakul) Formation.

Biostratigraphy:

Age: Early Cenomanian Stage, Uppermiddle Gallic Subepoch, Lowermost Gulf Epoch, Middle Cretaceous.

Material:

ZIN PH 13/87: Neural 1.

ZIN PH 8/87: Neural 4 (?).

ZIN PH S78-1: Costal 6.

ZIN PH 12/87: Peripheral 4.

ZIN PH 21/87: Undetermined costal fragment.

ZIN PH 3/87, 4/87: Peripherals 2.

ZIN PH 1/87: Peripheral 3.

ZI PH 2/87: Peripheral 7.

ZIN PH 5/87: Undetermined pheripheral fragment.

zI PH 6/87, 7/87: Bridge peripherals.

ZIN PH 20/87: Fragment of nuchal.

 

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Species: aksary NESOV & KRASNOVSKAYA, 1984
Etymology:
= Adocus aksari SUKHANOV, 2000 (sic)
= Lindholmemys elegans NESOV, 1986 (partim)
= Ferganemys sp NESOV, 1997

Holotype: CCMGE 10/12086

Locality: Dzharakuduk-II, central part of the Kyzylkum Desert, Uzbekistan.

Horizon: Bissekty Formation.

Biostratigraphy:

Age: Coniacian Stage, Lower Senonian Subepoch, Lower Gulf Epoch, Late Cretaceous.

Material: Left periphearl 3.

Note: In the original description the holotype was mistakently determined as a peripheral.

Referred material:

SYROMYATNIKOVA & DANILOV, 2009

ZIN PH 1-2/84, 4-6/84: Nuchals.

ZIN PH 24/84, 165/84: neurals 1.

ZIN PH 7/84: Neural 3.

ZIN PH 12/84,19/84: Neurals 4.

ZIN PH 13/84: neural 5.

ZIN PH 26/84, 27/84, 132/84: Suprapygals 2.

ZIN PH 28-32/84: Pygals.

ZIN PH 3/84, 163/84, 164/84: Costals 1.

ZIN PH154/84: Costal 3.

ZIN PH 144/84, 146/84: Costals 4.

ZIN PH 185/84: Costal 5.

ZIN PH 150/84: Costals 5 + 6 + neurals 5.

ZIN PH 143/84, 147/84: Costals 6.

ZIN PH 148/84: Costal 7.

ZIN PH 120-122/84: Costals 8.

ZIN PH 157/84: Undeterminate costal.

ZIN PH 34-40/84, 57/84: Peripherals 1.

CCMGE 30/12086, ZIN PH 21/84, 61-68/84, 74/84, 76-78/84: Pheripheral 3.

ZIN PH 170/84: Peripheral 6.

ZIN PH 60/84, 74,84, 76-78/84: Peripherals 7.

ZIN HP 79-91/94, 172/84, 173/84: Peripherals 8.

ZIN PH 171/84: Perpherals 7 +8.

CCMGE 29/12086, ZINPH 92-95/84, 98/84, 174/84: Peripherals 9.

ZIN PH 96/84, 97/84, 105/84, 175/84, 176/84, 177/84: Perpherals 10.

ZIN PH 111-118/84,195/84: Peripherals 11.

ZIN PH 69-71-71/84, 73/84, 168/84, 189-191/84: Bridge peripherals.

CCMGE 31/12086, ZIN PH 123-125/84: Epiplastra.

ZIN PH 126-129/84: Entoplastra.

ZIN PH 181/84: Hyoplastron + hypoplastron.

ZIN PH 119/84, 131/84: Hyoplastra.

ZIN PH 135-139/84: Xiphiplastra.

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Species: dzhurtasensis SYROMYATNIKOVA & DANILOV, 2009
Etymology: In reference to the Dzhurtas locality.

Holotype: ZIN PHT K27-1

Locality: Dzhurtas locality, Dzhungar Alatau Ridge, south-eastern Kazakhstan.

Horizon:

Biostratigraphy:

Age: Santonian Stage-early Campanian Stage, Lower Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material: Nuchal + left peripheral 1.

Referred material:

ZIN PH 3/92: Epiplastorn.

ZIN PH 1/92: Fragment of the anterior lobe, including epiplastra and entoplastron.

ZIPH 4/92: Hypoplastron.

ZIN PH 5/92: Xiphiplastron.

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Species: bostobensis SYROMYATNIKOVA & DANILOV, 2009
Etymology: In reference to the Bostobe Formation.

Holotype: ZIN PH 1/94

Locality: Baybishe I (Baybishe locality), Aral Sea area, Kazakhstan.

Horizon: Bostobe Formation.

Biostratigraphy:

Age: Santonian Stage-early Campanian Stage, Lower Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material: Nuchal fragment.

Referred material:

ZIN PH 14/94: Neural 1.

ZI PH 4/94: Undetermined costal fragment.

ZIN PH 2/94:Peripherals 2 + 3.

ZIN PH 3/94: Peripheral 8.

ZIN PH 11/94: Bridge peripheral.

ZIN PH 5/94: Hypoplastron.

ZIN PH 6/94: Hypoplastron + xiphipastron.

ZIN PH 7-13/94: Undetermined shell fragments.

 

Locality: Akkurgan locality, Aral Sea area, Kazakhstan.

Horizon: Bostobe Formation.

Biostratigraphy:

Age: Santonian Stage-early Campanian Stage, Lower Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

ZIN PH 1/96: Peripheral 1.

ZIN PH 2/96, 3/96: Peripherals 8.

ZIN PH 4/96: Undetermined shell fragment.

 

Locality: Buroynak Locality (Buroynak I and II), Aral Sea area, Kazakhstan.

Horizon: Bostobe Formation.

Biostratigraphy:

Age: Santonian Stage-early Campanian Stage, Lower Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

ZIN PH 6/95: Neural.

ZIN PH 1/95:Peripheral 2.

ZIN PH 7/95:Peripheral 9.

ZIN PH 2/95, 3/95: Undetermined fragments of peripherals.

ZIN PH 4/95, 5/95, 8/95, 9/95: Undetermined shell fragments.

 

Locality: Shakh-Shakah locality, Aral Sea area, Kazakhstan.

Horizon: Bostobe Formation.

Biostratigraphy:

Age: Santonian Stage-early Campanian Stage, Lower Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

ZIN PH 95/10: Peripheral 1.

ZIN PH 96/10: Peripheral 2.

ZIN PH 103/10, 104/10: Peripherals 8.

IZK 3970, 3971: Posterior peripherals.

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Species: planus (SUKANOV, 2000) DANILOV, SUKHANOV & SYROMYATNIKOVA, 2011
= Shineusemys plana SUKHANOV, 2000, (Unavailable name) SUKHANOV & NARMANDAKH, 2006
Etymology:

Holotype: PIN 4636-1

Locality: Shine Us Kkhuduk (= Shine Us Khudag) locality, Dornogov Aimag, Eastern Gobi, Mongolia.

Horizon: Lower part of the Bayanshiree Svita.

Biostratigraphy:

Age: Cenomanian-Early Turonian Stage, Middle-Upper Gallic Subepoch, Lower Gulf Epoch, Middle –Early Late Cretaceous.

Material: Partial shell with an almost complete plastron and a few carapace fragments on the steinkern.

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Species: kohaku HIRAYAMA, SONODA, UNO, HORIE, TSUTSUMI, SASAKI, MITSUZUKA & TAKISAWA, 2021
Etymology: Named after Japanese name of amber as this fossil was associated with numerous ambers.

Holotype: KAM 01

Locality: Kokuji, Kuji, Iwate Prefecture, Honshu Island, Japan.

Horizon: Tamagawa Formation, Kuji Group.

Biostratigraphy:

Age: Turonian Stage, Middle-Upper Gallic Subepoch, Lower Gulf Epoch, Middle –Early Late Cretaceous.

Material: A nearly coml;pete shell.

Referred material:

KAM 16: Anterior portion of carapace.

OSD 249: Right ninth peripheral.

OSD 520: Endoplastron.

OSD 1901: Partial carapace and plastron.

OSD 1929: Right third costal.

OSD 2278: Right costal.

OSD 2332: Nuchal.