Genus: Proterochersis E. FRAAS, 1913
Etymology:
= Keuperotesta SZCZYGIELSKI, & SULEJ, 2016
Etymology: In reference to Keuper, the geological unit where the animal
lived, and Latin, testa, "shell."
= Murrhardtia KARL & TICHY, 2000
Etymology: In reference to the type locality, Murrhardt.
= Proterochersys FRAAS, in
STROMER, 1912 (nomen nudum)
Species: robustata E. FRAAS, 1913
Etymology: Latin, robustus, "robust."
Holotype: SMNS 12777
Locality: Rudersberg, North Stuttgart, Baden-Wurttemberg State, Southern Germany.
Horizon: Löwenstein Formation, Lower Stubensandstein, Middle Keuper.
Biostratigraphy:
Age: Middle Norian Stage, Late Triassic Epoch, Late Triassic.
Material: Shell.
Referred material:
Localities: Murr Valley, and the vicinity of Stuttgart, Baden-Wurttemberg State, Germany.
Horizon: Stubensandstein, Middle Keuper.
Biostratigraphy:
Age: Middle Norian Stage, Late Triassic Epoch, Late Triassic.
Material: 12 shells and pelvis.
SZCZYGIELSKI, MARCHETTI & DRODZ, 2024
Locality: Quarry at Reichenbach at the river Fils, east of Stuttgart, Baden-Wurttemberg State, Germany.
Horizon: Löwenstein Fromation (formerly Stubensandstein).
Biostratigraphy:
Age: Norian Stage, Late Triassic Epoch, Late Triassic.
Material:
SMNS 15479: Largerly complete imprint of an unusually large plastron with no associated bone remains or steinkern.
SZCYGIELSKI, SLOWIAK & DROZDZ, 2018
Locality: In the vicinity of Lorch, Baden-Wurttemberg State, Germany.
Horizon: Stubensandstein, Middle Keuper.
Biostratigraphy:
Age: Middle Norian Stage, Late Triassic Epoch, Late Triassic.
Material:
SMNS 16603: Intern mold of a shell of a young individual with a caudal right part of carapace, a pelvis, a part of the right axillary buttress and an cranial fragment of the plastron cosisiting of the area of the right extragular and gular, a part of the left extragular, and medial parts of the humeral scutes.
Locality: Murrhardt-Mettelberg, 3 km from Kochersberg entfernt, Baden-Wurttemberg State, Germany.
Horizon: Upper Keuper, Lower Stubensandstein, Löwenstein Formation.
Biostratigraphy:
Age: ?Lower or Middle Norian Stage, Late Triassic Epoch, Late Triassic.
Material:
SMNS 17755: A left bridge and most of the left half of the plastron, consisting of the areas of the sixth to ninth and part of the 10th marginal, second third, and part of the first supramarginal, second and third inframarginal, second and third inframarginal, fragmetn of the humeral scute, most of the pectoral, both abdominals, femorla and fragment of anal scute, with visible sutures.
SMNS 17755a: A fragment of carapace consisting of the areas of the last vertebral scute, a broken off and dislocated fragment of the fourth vertebral scute area, a part of the third and complete fourth left pleural, a fragment of the 10th , most of the 11th, full 12th, and most of the 13th left marginal, with sacrum and dorsal parts of both ilia preserved inside and with visible sutures.
SMNS 18440: A left bridge fragment, comprising the area of small caudal fragments of the pectoral scute, nearly complete first and complete second abdominal scute, most of the femoral scute, a part of the first and complete second, third, and poorly preserved fourth inframarginal, a fragment of the fifth and 10th marginal scutes, and complete sixth, seventh, eighth, and nineth marginals, complte three supramarginals, and fragments of the second and third pleural scute, with sutures visible.
SMNS 50917: A fragmentary plastron consisting of an area of complete left femoral scute and fragmetns of the left pectoral, the first and second left abdominal, the right femoral and right and left anal scutes. Viscerally, a base of the pelvis and the epipubic process are preserved.
SMNS 56606: A mostly complete pelvis of a young individual with attached badly damaged caudal part of the carapace (natural surface preserved only in the partial area of the last and second-to-last left marginal scute and in some places along the rim of the caudal notch), at least one sacral vertebra and rib pair, caudal segment of the dorsal vertebral colum (approximately two complete and caudal half of the third centrum with associated rib heads), and poorly presrved caudal part of the plastron, cosisting of the areas of mostly intact intercaudal scute and damaged caudal sections of the anal scutes, with caudal processes broken off.
SMNS 81917: A partially preserved plastral bone (hypoplastron or hypoplastron) of a juvenile.
Locality: Oberbürden, Baden-Wurttemberg State, Germany.
Horizon: Upper Keuper, Lower Stubensandstein, Löwenstein Formation.
Biostratigraphy:
Age: ?Lower or Middle Norian Stage, Late Triassic Epoch, Late Triassic.
Material:
SMNS 17930: An internal mold of the carapace with embedded right part of the pelvis and small parts of the right bridge, and a large part of the carapace including the areas of the cervical scute, complete first vertebral and increasingly more fragmentary second, third, fourth, and small fragment of the fifth vertebral scute, complete first pair of pleurals, fragmentary second, third, and fourth right pleural, damaged right supramarginals, and marginal scutes: the first and second, a fragment of the third, complete fourth, fragmentary fifth and 1oth, complete 11th, and a fragment of the 12th.
= Proterochersis intermedia E.
FRAAS, 1913
Etymology:
Holotype: SMNS 11396
Locality: Stuttgart-Rohracker, Baden-Wurttemberg State, Germany.
Horizon: Löwenstein Formation, Stubensandstein, Middle Keuper.
Biostratigraphy:
Age: Middle Norian Stage, Late Triassic Epoch, Late Triassic.
Material: Fragmentary specimen, consisting of a natural interior mould of the posterior part of the carapace.
= Murrhardtia staeschei KARL & TICHY, 2000
Etymology:
Holotype: CSMM o. Nr.
Locality: Murrhardt-Mettelberg, 3 km from Kochersberg entfernt, Baden-Wurttemberg State, Germany.
Horizon: Upper Keuper, Lower Stubensandstein, Löwenstein Formation.
Biostratigraphy:
Age: ?Lower or Middle Norian Stage, Late Triassic Epoch, Late Triassic.
Material: Fragmentary shell.
Referred material:
SMNS 16442: Damaged anterior part of carpace, with nuchal bone, first few neurals, parts of first few costals, adn possibly a few peripherals, a damaged part of the carapace margin, with fragment of the anterior part of bridge, anterior lobe of the plastron, humeral scute area and entoplasteron, with epiplastera seeming missing, a rock impression of the latter, a posterior lobe of the plastron, area of femorals and anals with a base of the right caudal porcess, and some difficult to interpret fragments and impressions of the plaster (one o f them porbably of a marginal sulci).
SMNS 17561: Complete carapace.
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Species: porebensis SZCZYGIELSKI, & SULEJ, 2016
Etymology: In reference to Poreba, Poland, the place of discovery.
Holotype: ZPAL V.39/48
Locality: Poreba, Silesina Voivodeship, Poland.
Horizon: Zbasynek Beds.
Biostratigraphy: Subzone IVb of the Corollina Meyeriana Zone.
Age: Middle-Upper Norian Stage, Late Triassic Epoch, Late Triassic.
Material: Complete shell, with only a small fragment of the seond vertebral scute ara, most of the right dorsal porcess of epiplastron, and some small elements from the bridge area, wiht the second pair of the sacral ribs missing.
Paratypes:
ZPAL 39/34: Nearly complete and mostly well-preserved shell.
ZPAL 39/49: Nearly complete shell.
ZPAL V.39/72: A mostly complete carapace, missing only parts of the areas of the third, fourth, and fifth vertebral scute, fragments of the second left and third and fourth right pleural scute, part of the second and most of the third right supramarginal scute, as well as the eighth and cranial part of the ninth right marginal scutes. Associated with a proximal caudal vertebra.
ZPAL V.39/370: Part of the posterior left section of a carapace with most of the dorsal part of the left ilum.
Referred material:
SZCYGIELSKI, SLOWIAK & DROZDZ, 2018, SZCYGIELSKI & SLOWIAK, 2022
ZPAL V.39/1: A fragment of a middle right section of a carapace consisting of partial areas of teh first and teh secont vertebral sctue, including proximal parts of three ribs.
ZPAL V.39/2: A fragment of a middle left section of a carapace, consisting of partial areas of teh second adn the ghird verteral scute, including proximal parts of five ribs.
ZPAL V.39/3: A large part of an isoalted costal with sutural edges.
ZPAL V.39/4: An isolated neural bone with transverse, w-shaped sulcus.
ZPAL V.39/6: Caudal right section of a carapace, including partial area of the fifth vertebral scute, two las marginal scute areas, and partial area of the third-to-last marginal scute, with part of teh cudal process of the ilium.
ZPAL V.39/8: A fragmetn of a left bridge region, consisting of the areas of complete first and cranial pat of the second supramarginal scute, a part of the fifth, complete sixth, and part of the seventh marginal scute, a fragment of the first pleural scute, and minute fragments of the axillary and the first two inframarginal scutes.
ZPAL V.39/9: ?Costal bone.
ZPAL V.39/18: Caudal right part of the carapace, consisting of the last marginal scute area, partial area of the second-to-last and third -to-last marginal scute, partial fourth and fifh vertebral scute area, and partial area of the fourth right pleural, wiht part of the caudal process of the ilium.
ZPAL V.39/20: Posterior costal bone without teh overlying osteodermal mosaic.
ZPAL V.39/22: A fragment of the cranial left nuchal part of the carapace, including left half of the cervical cute, pat of the first vertebral scute, and parts of the cranial two marginals.
ZPAL V.39/23: A fragment of the caudal left carapacial rim, consisting of the area of the last marginal, a caudal part of the preceeding marginal, and a small fragment of the last vertebral sctue, with sutural edge.
ZPAL V.39/28: Right tenth marginal and parts of the third and fourth pleural and elevenenth marginal area.
ZPAL V.39/56: Left caudal process of plastron.
ZPAL V.39/57: Cranial left part of teh carapae, consisting of partial areas of the cervical scute, first vertebral, first left pleural, and marginal scute areas: complete first, fragmentary second, complete third and fourth and partial fifth.
ZPAL V.39/59: Caudal left part of the carapace, consisting of two last marginal scute areasand partial last vertebra sctue area, with part of the caudal process of the ilium.
ZPAL V.39/60: A fragment of a carapace, consisting of a complete ninth sctue area and fragmentary areas of the third supramarginal scute and the third pleural scute.
ZPAL V.39/61: ?Ninth marginal and third supramarginal area (peripheral and costal, possibly supernumerary carpacial ossification).
ZPAL V.39/63: Caudal right part of the carapace, consisting of partial areas of the fourth and fifth vertebral scutes, and a posterodorsal part of the fourth pleural scute area, with attached doral section of right ilium.
ZPAL V.39/66: Caudal plastral lobe consisting of the area of the caudal part of the anal scutes, nearly complete intercaudal scute, proxmial part of the right and complete left caudal sute, with the basal part of the ischia attached to the visceral strucutre.
ZPAL V.39/68: Caudal plastral lobe consisting of the area of the caudal part of the anal scutes, the complete intercaudal scute, and proximal parts of caudal scutes, with the basal part of the ischiia attache to the visceral surfae.
ZPAL V.39/69: Caudal plastral lobe consisting of the area of the anal, intercaudal, and caudal scutes, with bases of the lateral pubic processes and basal part of the ischia and pubis attached to the visceral surface.
ZPAL V.39/70: Caudal plastral lobe consisting of the area of the caudal part of the anal scutes, nearly complete intercaudal scute, proximal part of the left and complete right caudal scute, with the basal part of the ischia attached to the visceral surface.
ZPAL V.39/71: Caudal plastral lobe consisting of the area of the caudal pat of the anal scutes, nearly complete intercaudal scute, proximal part of the left and complete right caudal scute, with the basal part of the ishcia attached to the visceral surface.
ZPAL V.39/157: A fragmentary plastron, consisting of partial sections of the right femoral and right anal scute areas with attached right lateral pubic process.
ZPAL V.39/165: A partially preserved plastral bone (likely right hypoplastron) of a juvenile.
ZPAL V.39/169: A fragment of the dorsal vertebral column consisting of one complete and small fragments of two surrounding vertebrae and attached part of a carapace with intervertbral culcus.
ZPAL V.39/176: A fragment of a costal with sutural edges.
ZPAL V.39/186: Left gular tubercle of a large specimen.
ZPAL V.39/187: A fragment of cranial plastral lobe consisgint of the area of the right gular and extragular.
ZPAL V.39/189: A fragment of the cranial plastral lobe consisting of teh area of the right gular and medial partf of the right extragular.
ZPAL V.39/195: Plastral bone from the bridge region (?right first hyoplastron).
ZPAL V.39/197: A partial preserved plastral bone (likely right hyoplastron) of a juvenile.
ZPAL V.39/199: Left caudal process of plastron.
ZPAL V.39/200: (?) left caudal process of plastron.
ZPAL V.39/213: A fragment of the caudal left carapacial rim, consisting of the area of the last marginal, a caudal edge of the preceeding marginal, and a small fragment of the last vertebral scute, with suture running at the base of the peripheral.
ZPAL V.39/277: A partially preserved plastral bone of a juvenile.
ZPAL V.39/333: A fragment of the right part of the cranial plastral lobe, consisting of the complete right gular, extragular, and a cranial part of the humeral scute area, and small fragmetn of the left humeral scute area.
ZPAL V.39/377: An unankylosed fragment of the dorsal vertebral column of a young specimen, consisting of one complete veertebra and an cranal half of a succeeding vertebra.
ZPAL V.39/378: An ankylosed fragment of the dorsal vertebral column, consisting of one complete vertebra and fragmetns of a preceding and succeeding vertebral wiht associated rib head.
ZPAL V.39/379: A fragment of right part of the cranial plastral lobe, consisting of the complete right gular, extragular, and a cranial part of the right humeral scute area.
ZPAL V.39/380: Caudla right part of the carapace, consisting of two last marginal scute areas and partial last vertbral scute area, with part of the caudal process of ilium.
ZPAL V.39/381: A fragmentary cosal of a hatchling.
ZPAL V.39/382: A fragmetn of a large costal.
ZPAL V.39/383: A partially preserved plastral bone of a juvenile.
ZPAL V.39/384: A partially preserved plastral bone of a juvenile.
ZPAL V.39/385: A nearly complete cranial plastral lobe with bases of the dorsal processes of the epiplastra and complete caudal process of the entoplatron preserved on the visceral side, and areas of both gular, both extragular, two supernumerary, right humeral and right pectoral cutes as well as partially preserved left humeral and pectoral, and the first pair of abdominals.
ZPAL V.39/386: Caudal left part of the carapace, consisting of partial area of the last and third-to-last marginal scute, complete area of the second to last marginal scute, and partial fourth and fifth vertebral scute areas, with part of the caudal process of ilium.
ZPAL V.39/387: A fragment of the cranial plastral lobe consisting of the area of the left gular and extragular, and an cranial part of the left humeral scute area.
ZPAL V.39/388: A fragment of the cranial plastral lobe consisting of partial areas of the right gular and extragular.
ZPAL V.39/390: Cranial left part of carapace consisting of partial areas of the cervical scute, the first vertebral scutes, and damaged two cranialmost left marginal scutes.
ZPAL V.39/392: ?Posterior pheripheral.
ZPAL V.39/401: Hyoplastron and posterior process of teh entoplastron.
ZPAL V.39/402: Caudal part of the carapce consisting of the middle area of the fifth vertebral with coosified last dorsal vertebra and both scral vertebrae.
ZPAL V.39/416: Neural bone.
ZPAL V.39/417: Supernumerary carapacial ossification (?thrid or fourth pleural boss).
ZPAL V.39/420: A fragment of the cranial plastral lobe consisting of the area of the left gular and extragular.
ZPAL V.39/476: Plastral bone from the bridge region (?right hyoplastron).
ZPAL V.39/477, 479, 480: Costal bone.
ZPAL V.39/479: neural bone.
ZPAL V.39/482: neuchal region of the carapace.
= Proterochersis cf. porebensis CZEPINSKI, SULEJ, SZCZYGIELSKI, PAWLAK, DROZDZ & LEWCZUK, 2019, CZEPINSKI, DRODZDZ, SZCZ7GIELSKI, TALANDA, PAWLAK, LEWCZUK, RYTEL & SULEJ, 2021
Locality: Kocury (Kotzuren, Koczury, Koczurry, Koturen) near Dobrodzien (Kotzuren forest), near Guttenberg, Gorny Slask, Upper Silesia, southern Poland.
Horizon: Patoka Member, Grabowa Formation ('Lisow Breccia' [Lissauer Breccie], Middle Keuper).
Biostratigraphy:
Age: Late Triassic.
Material:
ZPAL V.66/9, /11: Carapace fragment.
ZPAL V.66/10: Bridge fragment.
ZPAL V.66.12: Part of a right femur.
ZPAL V.66/14, /25, /35: Shell fragment.
ZPAL V.66/15, /34: Costal fragment.
ZPAL V.66/16: Plastron fragment.
ZPAL V.66/18: Posterior rigth part of teh carapace.
ZPAL V.66/20: Left pubis.
ZPAL V.66/21: Shell (?carapace) fragment.
ZPAL V.66/37: Plastron (?hypoplastron) fragment.
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Species: limendorsa (SZCZYGIELSKI, & SULEJ, 2016) JOYCE, 2017
Etymology:
Latin, limen, "threshold" and Latin, dorsum, "back"; referring to the surface
of the shell.
= Proterochersis robusta JOYCE, SCHOCH, & LYSON, 2013
= Keuperotesta limendorsa SZCZYGIELSKI, & SULEJ, 2016
Holotype: SMNS 17757
Locality: Rudersberg, Baden-Wurttemberg State, Germany.
Horizon: Löwenstein Formation.
Biostratigraphy:
Age: ?Lower or Middle Norian Stage, Late Triassic Epoch, Late Triassic.
Material: A natural mould of the visceral surface of costals, with bone tissue remaining in some sections (fragments of the areas of the cervical scute and vertebrals I-V, left marginals I-IV, and first and fragment of the second left pleural, whole fourth right pleural, fragment of the first left supramarginal), and most of the plastron (excluding the anteriormost and posteriormost parts, and the bridge region).