Genus: Toxochelys COPE, 1873
Etymology:

Species: latiremis COPE, 1873
Etymology:

Holotype: AMNH 2362

Locality: Near the forks of the Smoky Hill River, Kansas.

Horizon: Probably Pierre shale.

Biostratigraphy:

Age: Late Cretaceous.

Material: Fragmentary lower jaw and a coracoid.

Referred material:

CASE, 1898

Locality: Unknown, Gove County, Kansas.

Horizon: Smoky Hill Chalk Member, Niobrara Formation.

Biostratigraphy:

Age: Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

KUVP 1215: Greater portion of skull.

 

HAY, 1896

Locality:

Horizon:

Biostratigraphy:

Age:

Material:

Number: Not given:

Fragmentary skull.

 

NICHOLLS, 1988

Locality: Manitoba Province, Canada.

Horizon: Pembina Member, Pierre Shale.

Biostratigraphy:

Age: Middle Campanian Stage, Senonian Subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

MMMN unnumbered: Partially articulated specimen.


Locality: South Dakota.

Horizon: Sharon Springs Member, Pierre Shale.

Biostratigraphy:

Age: Upper Lower to Lower Middle Campanian Stage, Senonian Subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

As per NICHOLLS, 1988

AMNH 1773, 1774, 1776, 1777, 1814, 2362, 5818, 5820, 5843, 6080, 8600, 14220, FMNH PR 213, 463, 417, 472, 473, 474, 513, 586, 587, 592, 593, 594, 595, 614, 648, 655, 656, 657, 658, 659, 660, 672, 675, 678, FMNH uncatalouged, KUVP 19500, 19503, 27841, 36117, 36118, 49637, 49638, 49639, 49640, 49641, 49642, 49643, 49644, 49645, 49646, 49647, 46949, 49650, 50002, 50003, 50004, 50005, 50006, 50007, 50008, 50009, 50020, LACM 52573, SDSM 425, 482, 4510, 4613, 4614, 5839, 6710, 52143 (2 specimens), 54348, 56130, SDSM EDW SP-2, SDSM 2 uncatalogued specimens, UCM 20561 (2 specimens), UCM 36388, 45438, 45442, UCM 3 uncatalogued specimens, UCMP V5407/44700, V5407/45199, V5407/45200, V5407/45201, UNSM 6-70, 6-74, USNM 2 uncatalouged, USNM 11064, 18252, 18362, 18279, 18279, 357134, 357135, 364380, YPM 9525:

 

NICHOLLS, TOKARYK & HILLS, 1990

Locality: MDM quarry 1, Reichert Pit, SW ¼, sec. 21, tp. 2, rge. 6, Manitoba Province, Canada.

Horizon: Pembina Member, Pierre Shale.

Biostratigraphy:

Age: Middle Campanian Stage, Senonian Subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

MDM T77.15.01: Scapula.

MDM T73.01.01: Partial skull.

 

Locality : MDM Quarry 4, MT. Nebo pit, SW ¼, sec. 18, tp. 4, rge. 6, WM1, Manitoba Province, Canada.

Horizon: Pembina Member, Pierre Shale.

Biostratigraphy:

Age: Middle Campanian Stage, Senonian Subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

MDM T74.01.04: Peripherals, scapular fragments.

 


Locality: MDM quarry 14, Robertoson south pit, NE ¼, sec. 13, tp. 4, rge. 7, WM1, Manitoba Province, Canada.

Horizon: Pembina Member, Pierre Shale.

Biostratigraphy:

Age: Middle Campanian Stage, Senonian Subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

MDM T80.03.12: Skull, mandible, front limbs, and girlds, 3 cervical vertebrae and carapace fragments.

 

Locality: MDM quarry 17, Baker pit, SW ¼, sec. 25, tp. 4, rge. 7, WM1, Manitoba Province, Canada.

Horizon: Pembina Member, Pierre Shale.

Biostratigraphy:

Age: Middle Campanian Stage, Senonian Subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

MDM T78.01.17: Incompelte mandible.

MDM T82.01.17: Limbs, girdles, partial carapace, and plastron.

MDM T82.02.17: Humerus.

 

RUSSELL, 1988

Locality: Sec. 23, T14, R6, 3 miles southwest of Castle Rock, Gove County, Kansas.

Horizon: Niobrara Formation.

Biostratigraphy:

Age: Upper Santonian,-Lower Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

ROM 28563: Almost complete specimen, lacking hind limbs, forelimbs distal to humerus and left ramus of mandible.


Locality:

Horizon:

Biostratigraphy:

Age:

Material:

KUVP 1244, FMNH PR 123, YPM 3602: Fragmentary carapaces.

 

ZANGERL, 1953

Locality: Butte Creek, Logan County, Kansas.

Horizon: Niobrara Formation.

Biostratigraphy:

Age: Late Cretaceous.

Material:

AMNH 5118: Skull.

 

Locality: Near fork of Smoky Hill, Kansas.

Horizon: Niobrara Formation.

Biostratigraphy:

Age: Late Cretaceous.

Material:

YPM 3602: Skull and partial shell.

YPM 3609, 3604, AMNH 1497: Skulls and mandibles.

YPM 361: Skull, vertebrae, girdle bones, epiplastron and limb-bones.

YPM 1386, AMNH 1042, USNM 11639, 11558: Partial skulls.

YPM 3610, 1387, AMNH 5118, 1498, 181, 5999, USNM 12011, 12010, KU (VP) 8352, KU (VP) 1210, KU (VP) 8256, FMNH UR3, FMNH PR124: Mandibles.

AMNH 1835: Skull fragments and mandible of 2 individuals.

USNM 11559: Skull.

USNM 1496, 1496: Partial skulls and mandibles.

FMNH PR125, FMNH UR9: Fragmentary juvenile skulls.

FMNH UR4: Mandible, vertebrae of a juvenile.

 

Locality: Unknown, Kansas.

Horizon: Smoky Hill Chalk Member, Niobrara Formation.

Biostratigraphy:

Age: Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

KUVP 8258: Scapula and right coracoid.

KUVP 1227: Femur and lower jaw.

KUVP 1225: Femur, ischium, ilium, and other fragments.

KUVP?

KUVP 1225, 1227:

 

Locality: 3.5 miles southwest of Castle Rock, Sec. 22-23, T14S, R26W, Gove County, Kansas.

Horizon: Smoky Hill Chalk Member, Niobrara Formation.

Biostratigraphy:

Age: Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

KUVP 65547: 4 elements comprising the first and fourth for fifth finger and other finger elements.

KUVP 1203: Skull.

 

Locality: Eagle Tail Creek, near Sharon Springs, T13S, R40W, Wallace County, Kansas.

Horizon: Smoky Hill Chalk Member, Niobrara Formation.

Biostratigraphy:

Age: Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

KUVP 1293: Posterior portion of skull.

 


Locality: Near Lisbon, Sec. 8, T12S, R36W, Logan County, Kansas.

Horizon: Smoky Hill Chalk Member, Niobrara Formation.

Biostratigraphy:

Age: Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

KUVP 1294: Fragment lower jaw.

 

MATZKE, 2007

Locality: 2 miles (3.3 km) north of pyramids, Gove County, Kansas.

Horizon: Niobrara Formation.

Biostratigraphy:

Age: Late Cretaceous.

Material:

USNM 11560: Almost complete skull.

 

Locality: 3 or 4 miles (5-6.6km) north of monument rocks, Gove County, Kansas.

Horizon: Niobrara Formation.

Biostratigraphy:

Age: Late Cretaceous.

Material:

Broken skull with preserved rostrum basisphenoidale.


WIELAND,


Locality: Kansas.

Horizon: Niobrara Formation.

Biostratigraphy:

Age: Late Cetaceans.

Material:

KUVP 2491: Skull, lower jaw, shell and skeleton.

 

= Toxochelys serrifer COPE, 1875
Etymology:

Holotype: AMNH 1835

Locality: Kansas.

Horizon: Niobrara Formation.

Biostratigraphy:

Age: Late Cretaceous.

Material: Fragmentary skull.

 

= Toxochelys brachyrhinus CASE, 1898
Etymology:

Holotype: KU (VP) 1212

Locality: Near fork of Smoky Hill River, Kansas.

Horizon: Niobrara Formation.

Biostratigraphy:

Age: Late Cretaceous.

Material: Fragmentary skull.

 

 

= Porthochelys browni HAY, 1905
Etymology:
= Toxochelys browni ZANGERL, 1953

Holotype: AMNH 6080

Locality: 20 miles south of Edgemont, Fall County, South Dakota.

Horizon: Pierre Formation.

Biostratigraphy:

Age: Late Cretaceous.

Material: Skull and dentary.

Referred material:

Skulls, mandibles and appendicular elements.

 

BARDACK, 1968

Locality: No Data, probably from a quarry near Thornhill, Manitoba Province, Canada.

Horizon: Vermilion River Formation.

Biostratigraphy:

Age: Late Cretaceous.

Material:

MMMN uncatalogued: Anterior limb including scapula, coracoid, humerus, radius, ulna, some carpals, metacarpals, and phalanges.

 

BREITHAUPT, 1985

Locality: Wyoming.

Horizon: Pierre Shale.

Biostratigraphy:

Age: Late Cretaceous.

Material:

 

= Toxochelys sp . cf. T. browni CARRINO, 2007

Locality: South Dakota.

Horizon: Basal Gammon Ferruginous Formation.

Biostratigraphy:

Age: Middle Campanian Stage,Senoniain Subepoch, Gulf Epoch, Late Cretaceous.

Material:

1 specimen.

 

Locality: South Dakota.

Horizon: Sharon Springs Formation.

Biostratigraphy:

Age: Campanian Stage, Senoniain Subepoch, Gulf Epoch, Late Cretaceous.

Material:

21 specimens.

 

Locality: South Dakota.

Horizon: Mitten Black Shale Formation.

Biostratigraphy:

Age: Campanian Stage, Senoniain Subepoch, Gulf Epoch, Late Cretaceous.

Material:

4 specimens.


ZANGERL, 1953

Locality: 20 miles southeast of Edgemont, Fall County, South Dakota.

Horizon: Pierre Formation.

Biostratigraphy:

Age: Late Cretaceous.

Material:

AMNH 5843: Large skull and attached mandible.

AMNH 1773: Snout of medium-sized individual.

AMNH 1774: Small skull and mandible.

AMNH unnumbered: Mandible.

FMNH PR214: Snout of medium-sized individual.

FMNH OR213: Skull and mandible.

SDSM 425: Large skull and mandible.

SDSM 4610: Large skull.

KU(VP) 1221: Partial skull.

KU(VP) 1222: Partial mandible.

 

= Phylemys barberi SCHMIDT, 1944
Etymology:
= Toxochelys barberi ZANGERL, 1953

Holotype: FM P27047

Locality: Wido Cox Farm, about 1 mile northeast of the junction of the Hollywood-Okolona Road to Arkadelphia, NNW 1/4, sec. 28, T 7S, R 20W, Clark County, Arkansas.

Horizon: Marlbrook Marl, Gulf Series.

Biostratigraphy:

Age: Late Cretaceous.

Material: Fragmentary shell.

 

= Toxochelys weeksi COLLINS, 1951
Etymology:

Holotype: UT K20

Locality: Tennessee.

Horizon: Ripley Formation.

Biostratigraphy: Middle-Uppermiddle Mosasaurus Acme-zone.

Age: Navesinkian age, Early Maastrichtian Stage? Upper Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material: Incomplete plastron and a few peripherals.

_____________________________________________________________________________________

Species: gigantea OERTEL, 1914 (nomen dubium)
Etymology:

Holotype:

Locality: Germany.

Horizon:

Biostratigraphy:

Age: Aptian Stage, Lower Gallic Subepoch, Upper Early Cretaceous Epoch, Middle Cretaceous.

Material: Incomplete skull.
_____________________________________________________________________________________

Species: moorevillensis ZANGERL, 1953
Etymology:

Holotype: FMNH P27330

Locality: Moore Brothers Farm, Harrell Station area, Dallas County, Alabama.

Horizon: Mooreville Chalk, Selma Formation.

Biostratigraphy: Middle Tylosaurus Acme-zone – Clidastes Acame zone.

Age: Niobraran age, Upper Santonian-Lower Campanian Stage, Senoniain Subepoch, Gulf Epoch, Late Cretaceous.

Material: A nearly complete shell, vertebral fragments and girdle bones.

Referred material:

Locality: Eutaw area, Greene County, Alabama.

Horizon: Mooreville Chalk, Selma Formation.

Biostratigraphy: Middle Tylosaurus Acme-zone – Clidastes Acame zone.

Age: Niobraran age, Upper Santonian-Lower Campanian Stage, Senoniain Subepoch, Gulf Epoch, Late Cretaceous.

Material:

FMNH PR168: Costal plates.

FMNH P27434, PR32: Fragmentary specimens.

FMNH P27348: Partial carapace and plastron.

CHHM P27438, P27444: Peripherals.

 

Locality: SW 1/4, SW 1/4, NW 1/4, sec. 33, T 16N, R 16E, Burkville area, Lowndes County, Alabama.

Horizon: Mooreville Chalk, Selma Formation.

Biostratigraphy: Middle Tylosaurus Acme-zone – Clidastes Acme zone.

Age: Niobraran age, Upper Santonian-Lower Campanian Stage, Senoniain Subepoch, Gulf Epoch, Late Cretaceous.

Material:

FMNH PR136: Nearly complete shell, girdle bones, some limb-bones and some vertebrae.

 


Locality: NE 1/4, NE 1/4, NE 1/4, sec. 5, T 15N, R 16E, Burkville area, Lowndes County, Alabama.

Horizon: Mooreville Chalk, Selma Formation.

Biostratigraphy: Middle Tylosaurus Acme-zone – Clidastes Acme zone.

Age: Niobraran age, Upper Santonian-Lower Campanian Stage, Senoniain Subepoch, Gulf Epoch, Late Cretaceous.

Material:

FMNH PR 218 : andible.

 


Locality: Clinton area, Greene County, Alabama.

Horizon: Mooreville Chalk, Selma Formation.

Biostratigraphy: Middle Tylosaurus Acme-zone – Clidastes Acme zone.

Age: Niobraran age, Upper Santonian-Lower Campanian Stage, Senoniain Subepoch, Gulf Epoch, Late Cretaceous.

Material:

FMNH PR113: Fragment.

 

Locality: Harrell Station area, Dallas County, Alabama.

Horizon: Mooreville Chalk, Selma Formation.

Biostratigraphy: Middle Tylosaurus Acme-zone – Clidastes Acame zone.

Age: Niobraran age, Upper Santonian-Lower Campanian Stage, Senoniain Subepoch, Gulf Epoch, Late Cretaceous.

Material:

FMNH P27341: A nearly complete carapace, fragments of plastron, vertebrae and a good pelvis.

FMNH PR224: Shell fragments.

FMNH PR188, PR109, P27544, P27549, P27556, P27555, P27349: Fragmentary specimens.

FMNH P27367, PR191, PR189: Fragmentary plastrons.

FMNH PR199: Anterior part of carapace.

FMNH PR264, P27358, PR266, PR140: Peripharals.

FMNH PR265, P27652, P27562, P27345: Costal plate.

FMNH P27347: Fragmentary plastron.

FMNH P27544: Costal fragments.

FMNH P27436, P27539: Peripheral fragments.

FMNH PR194: Neural and costal plates.

FMNH P27550: Partial skeleton.

FMNH PR144: Partial carapace.

FMNH PR219: Skull.

FMNH P27338: Posterior portion of skull.

 


Locality: T. 11, West of Highway 13, Cedarville area, Hale County, Alabama.

Horizon: Mooreville Chalk, Selma Formation.

Biostratigraphy: Middle Tylosaurus Acme-zone – Clidastes Acme zone.

Age: Niobraran age, Upper Santonian-Lower Campanian Stage, Senoniain Subepoch, Gulf Epoch, Late Cretaceous.

Material:

FMNH P27346: Fragmentary specimen.

 

Locality: West Greene Area, Greene County, Alabama.

Horizon: Mooreville Chalk, Selma Formation.

Biostratigraphy: Middle Tylosaurus Acme-zone – Clidastes Acme zone.

Age: Niobraran age, Upper Santonian-Lower Campanian Stage, Senoniain Subepoch, Gulf Epoch, Late Cretaceous.

Material:

FMNH PR167: Nearly complete shell and pelvis.

FMNH PR155: Carapace fragments.

FMNH PR154: Fragmentary specimen.

FMNH PR253: Peripherals.

FMNH PR166: Partial carapace.

 

Locality: Mt. Hebron area, Greene County, Alabama.

Horizon: Mooreville Chalk, Selma Formation.

Biostratigraphy: Middle Tylosaurus Acme-zone – Clidastes Acme zone.

Age: Niobraran age, Upper Santonian-Lower Campanian Stage, Senoniain Subepoch, Gulf Epoch, Late Cretaceous.

Material:

FMNH PR28: Nearly complete carapace and several vertebrae.

FMNH PR26: Peripherals.

 

Locality: Boligee area, Greene County, Alabama.

Horizon: Mooreville Chalk, Selma Formation.

Biostratigraphy: Middle Tylosaurus Acme-zone – Clidastes Acme zone.

Age: Niobraran age, Upper Santonian-Lower Campanian Stage, Senoniain Subepoch, Gulf Epoch, Late Cretaceous.

Material:

FMNH PR111, PR112, PR33: Fragmentary specimens.

FMNH PR60: Costal plates.

FMNH PR59: Anterior peripherals.