ZAMBIA
SOUTHERN PROVINCE
MID-ZAMBEZI BASIN
APPROXIMATELY 2 KM WEST OF MAZILA
LOCALITY L140
BASE OF THE UPPER MEMBER
MADUMABSIA MUDSTONE FORMATION
LOWER KAROO GROUP
TATARIAN AGE
LOPINGIAN SUBEPOCH
ZECHSTEIN EPOCH
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPID BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: ANOMODONTIA WATSON & ROMER, 1956
INFRAORDER: DICYNODONTIA OWEN, 1860
Superfamily: ENDOTHIODONTIDEA Owen, 1876 (sensu Cluver & King, 1983)
Family: ENDOTHIODONTIDAE Owen, 1876
Endothiodon sp
_____________________________________________________________________________________
ZAMBIA
SOUTHERN PROVINCE
MID-ZAMBEZI BASIN
APPROXIMATELY 2 KM WEST OF MAZILA
LOCALITY L143
BASE OF THE UPPER MEMBER
MADUMABSIA MUDSTONE FORMATION
LOWER KAROO GROUP
TATARIAN AGE
LOPINGIAN SUBEPOCH
ZECHSTEIN EPOCH
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPIDA BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: THERIODONTIA OWEN, 1881
SUBORDER: GORGONOPIA SEELEY, 1895
Superfamily: GORGONOPIOIDEA Lydekker, 1890
Family: GORGONOPIDAE Lydekker, 1890
Gorgonops sp
_____________________________________________________________________________________
ZAMBIA
SOUTHERN PROVINCE
MID-ZAMBEZI BASIN
APPROXIMATELY 2 KM WEST OF MAZILA
LOCALITY L144
BASE OF THE UPPER MEMBER
MADUMABSIA MUDSTONE FORMATION
LOWER KAROO GROUP
TATARIAN AGE
LOPINGIAN SUBEPOCH
ZECHSTEIN EPOCH
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPID BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: ANOMODONTIA WATSON & ROMER, 1956
INFRAORDER: DICYNODONTIA OWEN, 1860
Superfamily: ENDOTHIODONTIDEA Owen, 1876 (sensu Cluver & King, 1983)
Family: ENDOTHIODONTIDAE Owen, 1876
Endothiodon sp
_____________________________________________________________________________________
ZAMBIA
SOUTHERN PROVINCE
MID-ZAMBEZI BASIN
APPROXIMATELY 2 KM WEST OF MAZILA
LOCALITY L202
BASE OF THE UPPER MEMBER
MADUMABSIA MUDSTONE FORMATION
LOWER KAROO GROUP
TATARIAN AGE
LOPINGIAN SUBEPOCH
ZECHSTEIN EPOCH
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPID BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: ANOMODONTIA WATSON & ROMER, 1956
INFRAORDER: DICYNODONTIA OWEN, 1860
Superfamily: ENDOTHIODONTIDEA Owen, 1876 (sensu Cluver & King, 1983)
Family: ENDOTHIODONTIDAE Owen, 1876
Endothiodon sp
_____________________________________________________________________________________
ZAMBIA
SOUTHERN PROVINCE
MID-ZAMBEZI BASIN
APPROXIMATELY 2 KM WEST OF MAZILA
LOCALITY L213
BASE OF THE UPPER MEMBER
MADUMABSIA MUDSTONE FORMATION
LOWER KAROO GROUP
TATARIAN AGE
LOPINGIAN SUBEPOCH
ZECHSTEIN EPOCH
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPID BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: ANOMODONTIA WATSON & ROMER, 1956
INFRAORDER: DICYNODONTIA OWEN, 1860
Superfamily: ENDOTHIODONTIDEA Owen, 1876 (sensu Cluver & King, 1983)
Family: ENDOTHIODONTIDAE Owen, 1876
Endothiodon sp
_____________________________________________________________________________________
ZAMBIA
SOUTHERN PROVINCE
MID-ZAMBEZI BASIN
APPROXIMATELY 2 KM WEST OF MAZILA
LOCALITY L220
BASE OF THE UPPER MEMBER
MADUMABSIA MUDSTONE FORMATION
LOWER KAROO GROUP
TATARIAN AGE
LOPINGIAN SUBEPOCH
ZECHSTEIN EPOCH
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPID BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: ANOMODONTIA WATSON & ROMER, 1956
INFRAORDER: DICYNODONTIA OWEN, 1860
Superfamily: ENDOTHIODONTIDEA Owen, 1876 (sensu Cluver & King, 1983)
Family: ENDOTHIODONTIDAE Owen, 1876
Endothiodon sp
_____________________________________________________________________________________
ZAMBIA
SOUTHERN PROVINCE
MID-ZAMBEZI BASIN
APPROXIMATELY 2 KM WEST OF MAZILA
LOCALITY L225
BASE OF THE UPPER MEMBER
MADUMABSIA MUDSTONE FORMATION
LOWER KAROO GROUP
TATARIAN AGE
LOPINGIAN SUBEPOCH
ZECHSTEIN EPOCH
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPID BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: ANOMODONTIA WATSON & ROMER, 1956
INFRAORDER: DICYNODONTIA OWEN, 1860
Superfamily: ENDOTHIODONTIDEA Owen, 1876 (sensu Cluver & King, 1983)
Family: ENDOTHIODONTIDAE Owen, 1876
Endothiodon sp
_____________________________________________________________________________________
ZAMBIA
NORTHERN PROVINCE
LUANGWA BASIN
NEAR THE SOUTHERN BORDER OF NORTH LUANGWA NATIONAL PARK
LOCALITY L 53
UPPER MEMBER
MADUMABISA MUDSTONE FORMATION
EQUIVALENT TO THE CISTECEPHALUS ZONE
WUCHIAPINGIAN STAGE
TATARIAN AGE
LOPINGIAN SUBEPOCH
ZECHSTEIN EPOCH
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPID BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: ANOMODONTIA WATSON & ROMER, 1956
ORDER: ANOMODONTIA WATSON & ROMER, 1956
INFRAORDER: DICYNODONTIA OWEN, 1860
Superfamily: DIICTODONTIA Cluver & King, 1983
Family: EMYDOPOIDEA Cluver & King, 1983
Subfamily: CISTEOCEPHALINAE Haughton & Brink, 1954 (sensu Cluver & King, 1983)
*Kembawacela kitchingi
_____________________________________________________________________________________
ZAMBIA
NORTHERN PROVINCE
LUANGWA BASIN
NEAR THE SOUTHERN BORDER OF NORTH LUANGWA NATIONAL PARK
0.2 KM SOUTHEAST OF THE TYPE LOCALITY L53
LOCALITY L 55
UPPER MEMBER
MADUMABISA MUDSTONE FORMATION
EQUIVALENT TO THE CISTECEPHALUS ZONE
WUCHIAPINGIAN STAGE
TATARIAN AGE
LOPINGIAN SUBEPOCH
ZECHSTEIN EPOCH
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPID BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: ANOMODONTIA WATSON & ROMER, 1956
ORDER: ANOMODONTIA WATSON & ROMER, 1956
INFRAORDER: DICYNODONTIA OWEN, 1860
Superfamily: DIICTODONTIA Cluver & King, 1983
Family: EMYDOPOIDEA Cluver & King, 1983
Subfamily: CISTEOCEPHALINAE Haughton & Brink, 1954 (sensu Cluver & King, 1983)
Kembawacela kitchingi
_____________________________________________________________________________________
ZAMBIA
EASTERN PROVINCE
LUANGWA BASIN
1.5 KM SOUTH OF THE VILLAGE OF OLD MPUNDA
LOCALITY L 4
UPPER MEMBER
MADUMABISA MUDSTONE FORMATION
EQUIVALENT TO THE CISTECEPHALUS ZONE
WUCHIAPINGIAN STAGE
TATARIAN AGE
LOPINGIAN SUBEPOCH
ZECHSTEIN EPOCH
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPID BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: ANOMODONTIA WATSON & ROMER, 1956
ORDER: ANOMODONTIA WATSON & ROMER, 1956
INFRAORDER: DICYNODONTIA OWEN, 1860
Superfamily: DIICTODONTIA Cluver & King, 1983
Family: EMYDOPOIDEA Cluver & King, 1983
Subfamily: CISTEOCEPHALINAE Haughton & Brink, 1954 (sensu Cluver & King, 1983)
Kembawacela kitchingi
_____________________________________________________________________________________
ZAMBIA
EASTERN PROVINCE
LUANGWA BASIN
1 KM NORTHEAST OF DRYSDALL & KITCHING'S LOCALITY 3
LOCALITY L 7
UPPER MEMBER
MADUMABISA MUDSTONE FORMATION
EQUIVALENT TO THE CISTECEPHALUS ZONE
WUCHIAPINGIAN STAGE
TATARIAN AGE
LOPINGIAN SUBEPOCH
ZECHSTEIN EPOCH
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPID BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: ANOMODONTIA WATSON & ROMER, 1956
ORDER: ANOMODONTIA WATSON & ROMER, 1956
INFRAORDER: DICYNODONTIA OWEN, 1860
Superfamily: DIICTODONTIA Cluver & King, 1983
Family: EMYDOPOIDEA Cluver & King, 1983
Subfamily: CISTEOCEPHALINAE Haughton & Brink, 1954 (sensu Cluver & King, 1983)
Kembawacela kitchingi
_____________________________________________________________________________________
ZAMBIA
LUANGWA BASIN
MUCHINGA PROVINCE
NEAR THE VILLAGE OF SITWE
LOCLAITY L310
UPPER MADUMABISA MUDSTONE FORMATION
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPID BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: ANOMODONTIA WATSON & ROMER, 1956
INFRAORDER: DICYNODONTIA OWEN, 1860
Superfamily: PRISTERODONTIA Cluver & King, 1983
Family: DICYNODONTOIDAE Owen, 1859
Subfamily: CRYPTODONTINAE Owen, 1859
Tribe: DICYNODONTINI Owen, 1859
Dicynodon angielczyki
_____________________________________________________________________________________
ZAMBIA
NORTHERN PROVINCE
SOUTHERN BORDER OF NORTH LUANGWA NATIONAL PARK
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
_____________________________________________________________________________________SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPIDA BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: THERIODONTIA OWEN, 1881
SUBORDER: GORGONOPIA SEELEY, 1895
Superfamily: GORGONOPIOIDEA Lydekker, 1890
Family: GORGONOPIDAE Lydekker, 1890
Gen. sp indet.
ZAMBIA
SOUTHERN PROVINCE
MID-ZAMBEZI BASIN
APPROXIMATELY 2 KM WEST OF MAZILA
LOCALITY L145
MADUMABSIA MUDSTONE FORMATION
MIDDLE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPID BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: ANOMODONTIA WATSON & ROMER, 1956
INFRAORDER: DICYNODONTIA OWEN, 1860
Superfamily: ENDOTHIODONTIDEA Owen, 1876 (sensu Cluver & King, 1983)
Family: ENDOTHIODONTIDAE Owen, 1876
Abajuodon kaayai
_____________________________________________________________________________________
ZAMBIA
SOUTHERN PROVINCE
MID-ZAMBEZI BASIN
APPROXIMATELY 2 KM WEST OF MAZILA
LOCALITY L148
MADUMABSIA MUDSTONE FORMATION
MIDDLE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPID BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: ANOMODONTIA WATSON & ROMER, 1956
INFRAORDER: DICYNODONTIA OWEN, 1860
Superfamily: ENDOTHIODONTIDEA Owen, 1876 (sensu Cluver & King, 1983)
Family: ENDOTHIODONTIDAE Owen, 1876
Abajuodon kaayai
_____________________________________________________________________________________
ZAMBIA
SOUTHERN PROVINCE
MID-ZAMBEZI BASIN
APPROXIMATELY 2 KM WEST OF MAZILA
LOCALITY L160
MADUMABSIA MUDSTONE FORMATION
MIDDLE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPID BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: ANOMODONTIA WATSON & ROMER, 1956
INFRAORDER: DICYNODONTIA OWEN, 1860
Superfamily: ENDOTHIODONTIDEA Owen, 1876 (sensu Cluver & King, 1983)
Family: ENDOTHIODONTIDAE Owen, 1876
Abajuodon kaayai
_____________________________________________________________________________________
ZAMBIA
SOUTHERN PROVINCE
MID-ZAMBEZI BASIN
APPROXIMATELY 2 KM WEST OF MAZILA
LOCALITY L163
MADUMABSIA MUDSTONE FORMATION
MIDDLE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPID BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: ANOMODONTIA WATSON & ROMER, 1956
INFRAORDER: DICYNODONTIA OWEN, 1860
Superfamily: ENDOTHIODONTIDEA Owen, 1876 (sensu Cluver & King, 1983)
Family: ENDOTHIODONTIDAE Owen, 1876
Abajuodon kaayai
_____________________________________________________________________________________
ZAMBIA
SOUTHERN PROVINCE
MID-ZAMBEZI BASIN
APPROXIMATELY 2 KM WEST OF MAZILA
LOCALITY L165
MADUMABSIA MUDSTONE FORMATION
MIDDLE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPID BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: ANOMODONTIA WATSON & ROMER, 1956
INFRAORDER: DICYNODONTIA OWEN, 1860
Superfamily: ENDOTHIODONTIDEA Owen, 1876 (sensu Cluver & King, 1983)
Family: ENDOTHIODONTIDAE Owen, 1876
Abajuodon kaayai
_____________________________________________________________________________________
ZAMBIA
SOUTHERN PROVINCE
MID-ZAMBEZI BASIN
APPROXIMATELY 2 KM WEST OF MAZILA
LOCALITY L181
MADUMABSIA MUDSTONE FORMATION
MIDDLE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPID BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: ANOMODONTIA WATSON & ROMER, 1956
INFRAORDER: DICYNODONTIA OWEN, 1860
Superfamily: ENDOTHIODONTIDEA Owen, 1876 (sensu Cluver & King, 1983)
Family: ENDOTHIODONTIDAE Owen, 1876
Abajuodon kaayai
_____________________________________________________________________________________
ZAMBIA
LUANGWA BASIN
LOCALITY 49
MADUMABISA MUDSTONE
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPID BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: ANOMODONTIA WATSON & ROMER, 1956
SUBORDER: DICYNODONTIA OWEN, 1860
Superfamily: ENDOTHIODONTIDEA Owen, 1876 (sensu Cluver & King, 1983)
Family: ENDOTHIODONTIDAE Owen, 1876
Endothiodon sp
_____________________________________________________________________________________
ZAMBIA
SOUTHERN PROVINCE
GWEMBE DISTRICT
APPROXIMATELY 18 KM SW OF HAMWE
LOCALITY L70
MIDDLE MEMBER
MADUMABISA MUDSTONE
WUCHIAPINGIAN STAGE
EARLY LOPANGIAN
ZECHSTEIN EPOCH
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
_____________________________________________________________________________________SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
INFRACLASS DINOMORPHA IVAKNENKO, 2008
SUPERORDER: GORGONTIA IVAKNENKO, 2008
ORDER: DEINOCEPHALIA OLSEN, 1962
SUBORDER: EUDEINOCEPHALIA APPLEBY, CHARIG, COX, KERMACK & TARLO, 1967
INFRAORDER: EOTITANOSUCHIA BOONSTRA, 1963
Incertae sedis
*Wantulignathus gwembensis
ZAMBIA
NORTHERN PROVINCE
MUNYAMADZI GAME MANAGEMENT PARK
MADUMABISA MUDSTONE
WUCHIAPINGIAN STAGE
EARLY LOPANGIAN
ZECHSTEIN EPOCH
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPIDA BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: THERIODONTIA OWEN, 1881
EUTHERIODONTIA
SUBORDER: SCALOPOSAURIA BRINK, 1963
SUBORDER: SCALOPOSAURIA BRINK, 1963
SUBORDER: EUTHEROCEPHALIA HOPSON & BARGHUSEN, 1986
Incertae sedis
*Ichibengops munyamadziensis
INFRAORDER: BAURINI WATSON, 1917
Superfamily: BAUROIDEA Watson, 1911 (sensu Brink, 1986)
Family: KARENITIDAE Tatarinov, 1997
*Mupashi migrator
_____________________________________________________________________________________
ZAMBIA
NORTHERN PROVINCE
SOUTHERN BORDER OF NORTH LUANGWA NATINOAL PARK
LOCALITY L322
MADUMABISA MUDSTONE
WUCHIAPINGIAN STAGE
EARLY LOPANGIAN
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPIDA BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: THERIODONTIA OWEN, 1881
SUBORDER: GORGONOPIA SEELEY, 1895
INFRAORDER: ESTEMMENOSUCHOIDEA IVENKNENKOV, 2003
BURNETIAMORHA BROOM, 1923
Incerate sedis
*Isengops luangwensis
ORDER: ANOMODONTIA WATSON & ROMER, 1956
INFRAORDER: DICYNODONTIA OWEN, 1860
Superfamily: DIICTODONTIA Cluver & King, 1983
Family: EMYDOPOIDEA Cluver & King, 1983
Digalodon cf Digalodon rubidgei
_____________________________________________________________________________________
ZAMBIA
LUANGWA BASIN
LOCALITY 3
MADUMABISA MUDSTONE
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPID BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: ANOMODONTIA WATSON & ROMER, 1956
SUBORDER: DICYNODONTIA OWEN, 1860
Superfamily: ENDOTHIODONTIDEA Owen, 1876 (sensu Cluver & King, 1983)
Family: ENDOTHIODONTIDAE Owen, 1876
Endothiodon sp
_____________________________________________________________________________________
ZAMBIA
LUANGWA BASIN
LOCALITY 5
MADUMABISA MUDSTONE
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPID BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: ANOMODONTIA WATSON & ROMER, 1956
INFRAORDER: DICYNODONTIA OWEN, 1860
KISTECEPHALIA SEELEY, 1894
Family: CISTEOCEPHALINAE Haughton & Brink, 1954 (sensu Cluver & King, 1983)
Genus: Nova
_____________________________________________________________________________________
ZAMBIA
LUANGWA BASIN
LOCALITY 6
MADUMABISA MUDSTONE
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPID BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: ANOMODONTIA WATSON & ROMER, 1956
SUBORDER: DICYNODONTIA OWEN, 1860
INFRAORDER: DICYNODONTIA OWEN, 1860
Superfamily: EUMANTELLIA Broom, 1935
Family: EUMANTELLIDAE BROOM, 1935
Subfamily: EUMANTELLINAE BROOM, 1935
Pristerodon mackayi
_____________________________________________________________________________________
ZAMBIA
LUANGWA BASIN
LOCALITY 7
MADUMABISA MUDSTONE
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPID BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: ANOMODONTIA WATSON & ROMER, 1956
SUBORDER: DICYNODONTIA OWEN, 1860
INFRAORDER: DICYNODONTIA OWEN, 1860
Superfamily: EUMANTELLIA Broom, 1935
Family: EUMANTELLIDAE BROOM, 1935
Subfamily: EUMANTELLINAE BROOM, 1935
Pristerodon mackayi
_____________________________________________________________________________________
ZAMBIA
LUANGWA BASIN
LOCALITY 26
MADUMABISA MUDSTONE
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPID BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: ANOMODONTIA WATSON & ROMER, 1956
SUBORDER: DICYNODONTIA OWEN, 1860
INFRAORDER: DICYNODONTIA OWEN, 1860
THEROCEHELONIA SEELEY, 1894
Superfamily: EMYDOPOIDEA van Hoepen, 1934
Family: COMPSODONTIDAE van Hoepen, 1934
Compsodon helmoedi
_____________________________________________________________________________________
ZAMBIA
LUANGWA BASIN
LOCALITY 29
MADUMABISA MUDSTONE
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPID BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: ANOMODONTIA WATSON & ROMER, 1956
INFRAORDER: DICYNODONTIA OWEN, 1860
KISTECEPHALIA SEELEY, 1894
Superfamily: KINGORAMOIDEA Cluver & King, 1983
Family: KINGORIDAE Cluver & King, 1983
Dicynodontoides cf. D. nowacki
_____________________________________________________________________________________
ZAMBIA
LUANGWA BASIN
LOCALITY 30
MADUMABISA MUDSTONE
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPID BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: ANOMODONTIA WATSON & ROMER, 1956
INFRAORDER: DICYNODONTIA OWEN, 1860
BIDENTALIA vide OWEN, 1876
Superfamily: DICYNODONTOIDEA Owen, 1859
Family: DICYNODONTOIDAE Owen, 1859
Subfamily: CRYPTODONTINAE Owen, 1859
Tribe: OUDENODONTINI Broom, 1935 (sensu Cluver & King, 1983)
Oudenodon baini
_____________________________________________________________________________________
ZAMBIA
LUANGWA BASIN
LOCALITY 31
MADUMABISA MUDSTONE
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPID BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: ANOMODONTIA WATSON & ROMER, 1956
SUBORDER: DICYNODONTIA OWEN, 1860
INFRAORDER: DICYNODONTIA OWEN, 1860
THEROCEHELONIA SEELEY, 1894
Superfamily: PYLAECEPHALIA van Hoepen, 1934
Family: PYLAECEPHALIDAE van Hoepen, 1934
Subfamily: PYLAECEPHALINAE van Hoepen, 1934
Diictodon feliceps
_____________________________________________________________________________________
ZAMBIA
LUANGWA BASIN
LOCALITY 32
MADUMABISA MUDSTONE
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPID BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: ANOMODONTIA WATSON & ROMER, 1956
SUBORDER: DICYNODONTIA OWEN, 1860
Superfamily: ENDOTHIODONTIDEA Owen, 1876 (sensu Cluver & King, 1983)
Family: ENDOTHIODONTIDAE Owen, 1876
Endothiodon sp
_____________________________________________________________________________________
ZAMBIA
LUANGWA BASIN
LOCALITY 37
MADUMABISA MUDSTONE
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPID BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: ANOMODONTIA WATSON & ROMER, 1956
INFRAORDER: DICYNODONTIA OWEN, 1860
BIDENTALIA vide OWEN, 1876
Superfamily: DICYNODONTOIDEA Owen, 1859
Family: DICYNODONTOIDAE Owen, 1859
Subfamily: CRYPTODONTINAE Owen, 1859
Tribe: OUDENODONTINI Broom, 1935 (sensu Cluver & King, 1983)
Oudenodon baini
_____________________________________________________________________________________
ZAMBIA
LUANGWA BASIN
LOCALITY 38
MADUMABISA MUDSTONE
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPID BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: ANOMODONTIA WATSON & ROMER, 1956
SUBORDER: DICYNODONTIA OWEN, 1860
INFRAORDER: DICYNODONTIA OWEN, 1860
THEROCEHELONIA SEELEY, 1894
Superfamily: PYLAECEPHALIA van Hoepen, 1934
Family: PYLAECEPHALIDAE van Hoepen, 1934
Subfamily: PYLAECEPHALINAE van Hoepen, 1934
Diictodon feliceps
_____________________________________________________________________________________
ZAMBIA
LUANGWA BASIN
LOCALITY 45
MADUMABISA MUDSTONE
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPID BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: ANOMODONTIA WATSON & ROMER, 1956
SUBORDER: DICYNODONTIA OWEN, 1860
INFRAORDER: DICYNODONTIA OWEN, 1860
THEROCEHELONIA SEELEY, 1894
Superfamily: EMYDOPOIDEA van Hoepen, 1934
Family: COMPSODONTIDAE van Hoepen, 1934
Compsodon helmoedi
_____________________________________________________________________________________
ZAMBIA
LUANGWA BASIN
LOCALITY L48
MADUMABISA MUDSTONE
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPID BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: ANOMODONTIA WATSON & ROMER, 1956
SUBORDER: DICYNODONTIA OWEN, 1860
INFRAORDER: DICYNODONTIA OWEN, 1860
THEROCEHELONIA SEELEY, 1894
Superfamily: PYLAECEPHALIA van Hoepen, 1934
Family: PYLAECEPHALIDAE van Hoepen, 1934
Subfamily: PYLAECEPHALINAE van Hoepen, 1934
Diictodon feliceps
_____________________________________________________________________________________
ZAMBIA
LUANGWA BASIN
LOCALITY 49
MADUMABISA MUDSTONE
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPID BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: ANOMODONTIA WATSON & ROMER, 1956
SUBORDER: DICYNODONTIA OWEN, 1860
Superfamily: ENDOTHIODONTIDEA Owen, 1876 (sensu Cluver & King, 1983)
Family: ENDOTHIODONTIDAE Owen, 1876
Endothiodon sp
_____________________________________________________________________________________
ZAMBIA
LUANGWA BASIN
LOCALITY 50
MADUMABISA MUDSTONE
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPID BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: ANOMODONTIA WATSON & ROMER, 1956
SUBORDER: DICYNODONTIA OWEN, 1860
INFRAORDER: DICYNODONTIA OWEN, 1860
Superfamily: EUMANTELLIA Broom, 1935
Family: EUMANTELLIDAE BROOM, 1935
Subfamily: EUMANTELLINAE BROOM, 1935
Pristerodon mackayi
_____________________________________________________________________________________
ZAMBIA
LUANGWA BASIN
LOCALITY 52
MADUMABISA MUDSTONE
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPID BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: ANOMODONTIA WATSON & ROMER, 1956
SUBORDER: DICYNODONTIA OWEN, 1860
INFRAORDER: DICYNODONTIA OWEN, 1860
Superfamily: EUMANTELLIA Broom, 1935
Family: EUMANTELLIDAE BROOM, 1935
Subfamily: EUMANTELLINAE BROOM, 1935
Pristerodon mackayi
THEROCEHELONIA SEELEY, 189
Superfamily: EMYDOPOIDEA van Hoepen, 1934
Family: EMYDOPIDAE van Hoepen, 1934
Emydops sp
_____________________________________________________________________________________
ZAMBIA
LUANGWA BASIN
LOCALITY 53
MADUMABISA MUDSTONE
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPID BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: ANOMODONTIA WATSON & ROMER, 1956
INFRAORDER: DICYNODONTIA OWEN, 1860
KISTECEPHALIA SEELEY, 1894
Family: CISTEOCEPHALINAE Haughton & Brink, 1954 (sensu Cluver & King, 1983)
Genus: Nova
_____________________________________________________________________________________
ZAMBIA
LUANGWA BASIN
LOCALITY 55
MADUMABISA MUDSTONE
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPID BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: ANOMODONTIA WATSON & ROMER, 1956
INFRAORDER: DICYNODONTIA OWEN, 1860
KISTECEPHALIA SEELEY, 1894
Family: CISTEOCEPHALINAE Haughton & Brink, 1954 (sensu Cluver & King, 1983)
Genus: Nova
_____________________________________________________________________________________
ZAMBIA
LUANGWA BASIN
LOCALITY 59
MADUMABISA MUDSTONE
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPID BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: ANOMODONTIA WATSON & ROMER, 1956
INFRAORDER: DICYNODONTIA OWEN, 1860
KISTECEPHALIA SEELEY, 1894
BIDENTALIA
cf. Katumbia parringtoni
Superfamily: DICYNODONTOIDEA Owen, 1859
Family: DICYNODONTOIDAE Owen, 1859
Subfamily: CRYPTODONTINAE Owen, 1859
Tribe: OUDENODONTINI Broom, 1935 (sensu Cluver & King, 1983)
Oudenodon baini
_____________________________________________________________________________________
ZAMBIA
LUANGWA BASIN
LOCALITY 61
MADUMABISA MUDSTONE
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPID BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: ANOMODONTIA WATSON & ROMER, 1956
INFRAORDER: DICYNODONTIA OWEN, 1860
Superfamily: PRISTERODONTIA Cluver & King, 1983
Family: DICYNODONTOIDAE Owen, 1859
Subfamily: CRYPTODONTINAE Owen, 1859
Tribe: DICYNODONTINI Owen, 1859
Syops vanhoepeni
_____________________________________________________________________________________
ZAMBIA
LUANGWA BASIN
LOCALITY 64
MADUMABISA MUDSTONE
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPID BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: ANOMODONTIA WATSON & ROMER, 1956
INFRAORDER: DICYNODONTIA OWEN, 1860
KISTECEPHALIA SEELEY, 1894
Superfamily: KINGORAMOIDEA Cluver & King, 1983
Family: KINGORIDAE Cluver & King, 1983
Dicynodontoides cf. D. nowacki
_____________________________________________________________________________________
ZAMBIA
LUANGWA BASIN
LOCALITY L424
UPPER MEMBER
MADUMABISA MUDSTONE
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPID BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: ANOMODONTIA WATSON & ROMER, 1956
INFRAORDER: DICYNODONTIA OWEN, 1860
Superfamily: DICYNODONTOIDEA Owen, 18593
Family: DICYNODONTOIDAE Owen, 1859
Incertae sedis
Gen. sp Indet.
_____________________________________________________________________________________
ZAMBIA
LUANGWA BASIN
KERR'S 1974 LOCALITY 1
MADUMABISA MUDSTONE
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
_____________________________________________________________________________________SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPID BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: ANOMODONTIA WATSON & ROMER, 1956
INFRAORDER: DICYNODONTIA OWEN, 1860
Superfamily: DICYNODONTIA Owen, 1859
Family: DICYNODONTOIDAE Owen, 1859
Subfamily: CRYPTODONTINAE Owen, 1859
Tribe: DICYNODONTINI Owen, 1859
Dicynodon huenei
Superfamily: KANNEMEYEROIDEA Huene, 1948
Family: KANNEMEYERIDAE Huene, 1948
Subfamily: LYSTROSAURINAE Cope, 1870 (sensu Broom, 1903)
Genus: Nova = Lystrosaurus cf. L. curvatus
ZAMBIA
LUANGWA BASIN
KERR'S 1974 LOCALITY 2
MADUMABISA MUDSTONE
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPID BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: ANOMODONTIA WATSON & ROMER, 1956
INFRAORDER: DICYNODONTIA OWEN, 1860
Superfamily: PRISTERODONTIA Cluver & King, 1983
Family: DICYNODONTOIDAE Owen, 1859
Subfamily: CRYPTODONTINAE Owen, 1859
Tribe: RHACHIOCEPHALINI Maisch, 2000
Kitchinganomodon crassus
_____________________________________________________________________________________
ZAMBIA
LUANGWA BASIN
KERR'S 1974 LOCALITY 5
MADUMABISA MUDSTONE
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPID BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: ANOMODONTIA WATSON & ROMER, 1956
INFRAORDER: DICYNODONTIA OWEN, 1860
Superfamily: PRISTERODONTIA Cluver & King, 1983
Family: DICYNODONTOIDAE Owen, 1859
Subfamily: CRYPTODONTINAE Owen, 1859
Tribe: DICYNODONTINI Owen, 1859
Syops vanhoepeni
_____________________________________________________________________________________
ZAMBIA
LUANGWA BASIN
KERR'S 1974 LOCALITY 6
MADUMABISA MUDSTONE
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPID BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: ANOMODONTIA WATSON & ROMER, 1956
INFRAORDER: DICYNODONTIA OWEN, 1860
Superfamily: PRISTERODONTIA Cluver & King, 1983
Family: DICYNODONTOIDAE Owen, 1859
Subfamily: CRYPTODONTINAE Owen, 1859
Tribe: DICYNODONTINI Owen, 1859
Dicynodon huenei
_____________________________________________________________________________________
ZAMBIA
LUANGWA BASIN
KERR'S 1974 LOCALITY 7
MADUMABISA MUDSTONE
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPID BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: ANOMODONTIA WATSON & ROMER, 1956
INFRAORDER: DICYNODONTIA OWEN, 1860
Superfamily: PRISTERODONTIA Cluver & King, 1983
Family: DICYNODONTOIDAE Owen, 1859
Subfamily: CRYPTODONTINAE Owen, 1859
Tribe: DICYNODONTINI Owen, 1859
Dicynodon huenei
_____________________________________________________________________________________
ZAMBIA
LUANGWA BASIN
KERR'S 1974 LOCALITY 11
MADUMABISA MUDSTONE
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPID BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: ANOMODONTIA WATSON & ROMER, 1956
SUBORDER: DICYNODONTIA OWEN, 1860
INFRAORDER: DICYNODONTIA OWEN, 1860
THEROCEHELONIA SEELEY, 1894
Superfamily: PYLAECEPHALIA van Hoepen, 1934
Family: PYLAECEPHALIDAE van Hoepen, 1934
Subfamily: PYLAECEPHALINAE van Hoepen, 1934
Diictodon feliceps
BIDENTALIA
Superfamily: DICYNODONTOIDEA Owen, 1859
Family: DICYNODONTOIDAE Owen, 1859
Subfamily: CRYPTODONTINAE Owen, 1859
Tribe: OUDENODONTINI Broom, 1935 (sensu Cluver & King, 1983)
Oudenodon baini
_____________________________________________________________________________________
ZAMBIA
LUANGWA BASIN
KERR'S 1974 LOCALITY 13
MADUMABISA MUDSTONE
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPID BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: ANOMODONTIA WATSON & ROMER, 1956
SUBORDER: DICYNODONTIA OWEN, 1860
INFRAORDER: DICYNODONTIA OWEN, 1860
THEROCEHELONIA SEELEY, 1894
Superfamily: PYLAECEPHALIA van Hoepen, 1934
Family: PYLAECEPHALIDAE van Hoepen, 1934
Subfamily: PYLAECEPHALINAE van Hoepen, 1934
Diictodon feliceps
BIDENTALIA
Superfamily: DICYNODONTOIDEA Owen, 1859
Family: DICYNODONTOIDAE Owen, 1859
Subfamily: CRYPTODONTINAE Owen, 1859
Tribe: OUDENODONTINI Broom, 1935 (sensu Cluver & King, 1983)
Oudenodon baini
_____________________________________________________________________________________
ZAMBIA
LUANGWA BASIN
KERR'S 1974 LOCALITY 16
UPPER NTAWERE FORMATION
ANISIAN STAGE
LOWER MID TRIASSIC EPOCH
LOWER MIDDLE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPID BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: ANOMODONTIA WATSON & ROMER, 1956
INFRAORDER: DICYNODONTIA OWEN, 1860
Superfamily: KANNEMEYEROIDEA Huene, 1948
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
ZAMBIA
LUANGWA BASIN
APPROXIMATELY 30 KILOMETERS NORTHWEST OF THE VILLAGE OF CHITUNGULU
NEAR THE SOUTHERN BORDER OF NORTH LUANGWA NATIONAL PARK
WITHIN 500 METERS OF LOCALITY L39
UPPER MEMBER OF THE MADUMABISA MUDSTONE FORMATION
LOWER KAROO GROUP
CORRELATES WITH THE CISTECEPHALUS ZONE
TATARIAN AGE
LOPINGIAN SUBEPOCH
ZECHSTEIN EPOCH
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
_____________________________________________________________________________________SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
INFRACLASS DINOMORPHA IVAKNENKO, 2008
SUPERORDER: GORGONTIA IVAKNENKO, 2008
ORDER: DEINOCEPHALIA OLSEN, 1962
SUBORDER: EUDEINOCEPHALIA APPLEBY, CHARIG, COX, KERMACK & TARLO, 1967
INFRAORDER: EOTITANOSUCHIA BOONSTRA, 1963
Family: EOTITANOSUCHIDAE CHUDINOV, 1960
Biarmosuchus sp
ZAMBIA
CHIKONTA
CISTECEPHALUS ZONE
TATARIAN AGE
LOPINGIAN SUBEPOCH
ZECHSTEIN EPOCH
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPID BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: ANOMODONTIA WATSON & ROMER, 1956
INFRAORDER: DICYNODONTIA OWEN, 1860
Superfamily: PRISTERODONTIA Cluver & King, 1983
Family: DICYNODONTOIDAE Owen, 1859
Subfamily: CRYPTODONTINAE Owen, 1859
Tribe: OUDENODONTINI Broom, 1935 (sensu Cluver & King, 1983)
Oudenodon baini
_____________________________________________________________________________________
EAST AFRICA
ZAMBIA
3 TO 4 M N OF N’AWERE
LUANGWA VALLEY
HORIZON 5
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
_____________________________________________________________________________________SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPID BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: ANOMODONTIA WATSON & ROMER, 1956
INFRAORDER: DICYNODONTIA OWEN, 1860
Superfamily: PRISTERODONTIA Cluver & King, 1983
Family: DICYNODONTOIDAE Owen, 1859
Subfamily: CRYPTODONTINAE Owen, 1859
Tribe: DICYNODONTINI Owen, 1859
*Dicynodon roberti
EAST AFRICA
ZAMBIA
3 TO 4 M N OF N’AWENE
HORIZON 5
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
_____________________________________________________________________________________SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPID BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: ANOMODONTIA WATSON & ROMER, 1956
INFRAORDER: DICYNODONTIA OWEN, 1860
Superfamily: PRISTERODONTIA Cluver & King, 1983
Family: DICYNODONTOIDAE Owen, 1859
Subfamily: CRYPTODONTINAE Owen, 1859
Tribe: OUDENODONTINI Broom, 1935 (sensu Cluver & King, 1983)
Oudenodon baini
= Dicynodon roberti
= Oudenodon roberti
EAST AFRICA
ZAMBIA
“LUANGWA VALLEY”
UPPER MADUMABSIA MUDSTONE
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
_____________________________________________________________________________________SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPID BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: ANOMODONTIA WATSON & ROMER, 1956
INFRAORDER: DICYNODONTIA OWEN, 1860
Superfamily: PRISTERODONTIA Cluver & King, 1983
Family: DICYNODONTOIDAE Owen, 1859
Subfamily: CRYPTODONTINAE Owen, 1859
Tribe: RHACHIOCEPHALINI Maisch, 2000
Odontocyclops dubius
= Rhachinocephalus dubius
= Platypodosaurus dubius
SOUTH AFRICA
ZAMBIA
LUANGWA VALLEY
3 M FROM CHIKONTA ON MPUNDA ROAD
HORIZON 4
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPID BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: ANOMODONTIA WATSON & ROMER, 1956
INFRAORDER: DICYNODONTIA OWEN, 1860
Superfamily: PRISTERODONTIA Cluver & King, 1983
Family: DICYNODONTOIDAE Owen, 1859
Subfamily: CRYPTODONTINAE Owen, 1859
Tribe: OUDENODONTINI Broom, 1935 (sensu Cluver & King, 1983)
*Oudenodon luangwaensis
= Dicynodon luangwaensis
_____________________________________________________________________________________
SOUTH AFRICA
ZAMBIA
LUANGWA VALLEY
LOCALITY 13
N’TAWRE FORMATION
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
_____________________________________________________________________________________SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPIDA BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: THERIODONTIA OWEN, 1881
EUTHERIODONTIA
SUBORDER: SCALOPOSAURIA BRINK, 1963
INFRAORDER: SCALOPOSAURINI BRINK, 1963
Superfamily: ICTIDOSUCHOIDEA Broom, 1903 (sensu Brink, 1986)
Family: NANICTIDOPSIDAE Watson & Romer, 1956
Ictidosuchops intermedius
= Ictidosuchoides intermedius
EAST AFRICA
N ZAMBIA
MID-LUANGWA VALLEY
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPID BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: ANOMODONTIA WATSON & ROMER, 1956
INFRAORDER: DICYNODONTIA OWEN, 1860
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
ZAMBIA
BUSI VALLEY
GUNYANKAS KRAAL
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPID BROOM, 1905
GRANDORDER: EUTHERAPSIDA
ORDER: DEINOCEPHALIA OLSEN, 1962
SUBORDER: TAPINOCEPHALIA BOONSTRA, 1955
INFRAORDER: TITANOSUCHIA
Superfamily: TAPINOCEPHALOIDEA Lydekker, 1890 (sensu Huene, 1956)
Family: TAPINOCEPHALIDAE Lydekker, 1890
Subfamily: TAPINOCEPHALINAE Lydekker, 1890 (sensu Kuhn, 1961)
*Criocephalus gunyankaensis
_____________________________________________________________________________________
ZAMBIA
LUANGWA VALLEY
NORTH SIDE OF MUNYAMDZI RIVER
EAST SIDE OF HUNTER’S TRACK FROM LUANGWA RIVER
MADUMABISA MUDSTONE
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPID BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: ANOMODONTIA WATSON & ROMER, 1956
INFRAORDER: DICYNODONTIA OWEN, 1860
Superfamily: KANNEMEYEROIDEA Huene, 1948
Family: KANNEMEYERIDAE Huene, 1948
Subfamily: LYSTROSAURINAE Cope, 1870 (sensu Broom, 1903)
Lystrosaurus sp
_____________________________________________________________________________________
EAST AFRICA
ZAMBIA
LOCALITY 4
UPPER LUANGWA VALLEY
HOEDEMAKER MEMBER
MIDDLE TEEKLOOF FORMATION
TROPIDOSTOMA ZONE (LOWER CISTECEPHALUS ZONE KITCHING, 1970, 1977)
TATARIAN AGE
LOPINGIAN SUBEPOCH
ZECHSTEIN EPOCH
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
_____________________________________________________________________________________SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPIDA BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: THERIODONTIA OWEN, 1881
SUBORDER: GORGONOPIA SEELEY, 1895
Superfamily: GORGONOPIOIDEA Lydekker, 1890
Family: GORGONOPIDAE Lydekker, 1890
Subfamily: GORGONOPSINAE Sigogneau, 1970
Aelurognathus quadrata
= Dixeya quadrata
EAST AFRICA
ZAMBIA
LOCALITY 4
UPPER LUANGWA
CISTECEPHALUS ZONE
TATARIAN AGE
LOPINGIAN SUBEPOCH
ZECHSTEIN EPOCH
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
_____________________________________________________________________________________CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPIDA BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: THERIODONTIA OWEN, 1881
SUBORDER: GORGONOPIA SEELEY, 1895
Superfamily: GORGONOPIOIDEA Lydekker, 1890
Family: GORGONOPIDAE Lydekker, 1890
Subfamily: RUBIDGEINAE Broom, 1938
Aelurognathus nasuta
= Dixeya nausta
= Dixeya quadrataAelurognathus sp
EAST AFRICA
ZAMBIA
LOCALITY 5
UPPER LUANGWA VALLEY
HOEDEMAKER MEMBER
MIDDLE TEEKLOOF FORMATION
TROPIDOSTOMA ZONE (LOWER CISTECEPHALUS ZONE KITCHING, 1970, 1977)
TATARIAN AGE
LOPINGIAN SUBEPOCH
ZECHSTEIN EPOCH
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
_____________________________________________________________________________________SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPIDA BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: THERIODONTIA OWEN, 1881
SUBORDER: GORGONOPIA SEELEY, 1895
Superfamily: GORGONOPIOIDEA Lydekker, 1890
Family: GORGONOPIDAE Lydekker, 1890
Subfamily: RUBIDGEINAE Broom, 1938
Aelurognathus quadrata
= Dixeya quadrataProrubidgea sp
Sycosaurus laticeps
SOUTH AFRICA
ZAMBIA
UPPER LUWANGWA VALLEY
LOCALITY 5
DICYNODON ZONE (UPPER CISTECEPHALUS ZONE WATSON, 1914)
TATARIAN AGE
LOPINGIAN SUBEPOCH
ZECHSTEIN EPOCH
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
_____________________________________________________________________________________SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPIDA BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: THERIODONTIA OWEN, 1881
SUBORDER: GORGONOPIA SEELEY, 1895
Superfamily: RUBIDGEOIDEA Broom, 1938
Family: RUBIDGEIDAE Broom, 1938
Subfamily: RUBIDGEINAE Broom, 1938 (sensu Sigogneau, 1970)
Prorubidgea sp
ZAMBIA
CISTECEPHALUS ZONE
TATARIAN AGE
LOPINGIAN SUBEPOCH
ZECHSTEIN EPOCH
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: PARAREPTILIA OLSON 1947
INFRACLASS: SAUROPSIDA HUXLEY, 1864
ORDER: PROCOLOPHONOMORPHA ROMER, 1964
SUBORDER: PAREIASAUROIDEA SELEY, 1888
Family: PARAEIASAURIDAE Cope, 1869
Subfamily: PAREIASAURINAE Nopcsa, 1923
Pareiasuchus nasicornis
= Pareiasaurus nasicornis
_____________________________________________________________________________________
ZAMBIA
UPPER LUANGWA VALLEY
CISTECEPHALUS ZONE
TATARIAN AGE
LOPINGIAN SUBEPOCH
ZECHSTEIN EPOCH
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: PARAREPTILIA OLSON 1947
INFRACLASS: SAUROPSIDA HUXLEY, 1864
SUPERORDER: EUREPTILIA OLSON, 1947
ORDER: CAPTORHINOMORPHA WATSON, 1917
SUBORDER: CAPTORHINOMORPHA WATSON, 1917
Family: CAPTORHINIDAE Case, 1911
Subfamily: CAPTORHININAE Case, 1911 (sensu De Ricqles, et AL 1982)
Captorhinus sp
_____________________________________________________________________________________
EAST AFRICA
ZAMBIA
MIDDLE LUANGWA VALLEY
MADUMABISA MUDSTONE
EQUIVELENT TO DICYNODON ASSEMBLAGE ZONE (FORMERLY DAPTOCEPHALUS ZONE OF KITCHING, 1870, 1977)
TATARIAN AGE
LOPINGIAN SUBEPOCH
ZECHSTEIN EPOCH
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPIDA BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: THERIODONTIA OWEN, 1881
EUTHERIODONTIA
SUBORDER: CYNODONTIA OWEN, 1860
INFRAORDER: PROCYNOSUCHIA
Family: PROCYOSUCHIDAE Broom, 1937
Procynosuchus delaharpeae
_____________________________________________________________________________________
SOUTH AFRICA
ZAMBIA
LUANGWA VALLEY
UPPER ENDOTHIODON OR HOEDEMAKER MEMBER
MIDDLE TEEKLOOF FORMATION
TROPIDOSTOMA ZONE (LOWER CISTECEPHALUS ZONE KITCHING, 1970, 1977)
TATARIAN AGE
LOPINGIAN SUBEPOCH
ZECHSTEIN EPOCH
LATE PERMIAN
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPIDA BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: THERIODONTIA OWEN, 1881
SUBORDER: GORGONOPIA SEELEY, 1895
Superfamily: GORGONOPIOIDEA Lydekker, 1890
Family: GORGONOPIDAE Lydekker, 1890
Subfamily: CYNARIOPINAE Watson & Romer, 1956
Scylacops sp
_____________________________________________________________________________________
NE ZAMBIA
N’TAWERE FORMATION
EARLY TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
MAGNORDER: PROTEROSUCHIFORMES nova
ORDER: PROTEROSUCHIA BROOM, 1906
Family: PROTEROSUCHIDAE Huene, 1914
Proterosuchus sp
= Chasmatosaurus ?
_____________________________________________________________________________________
EAST AFRICA
NORTHERN ZAMBIA
NORTHERN PART OF THE LUANGWA VALLEY
UPPER NTAWERE FORMATION
LATE ANISIAN STAGE/EARLY LADINIAN STAGE
MID TRIASSIC EPOCH
MIDDLE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPIDA BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: THERIODONTIA OWEN, 1881
EUTHERIODONTIA
SUBORDER: CYNODONTIA OWEN, 1860
EPICYNODONTIA HOPSON & KITCHING, 2001
INFRAORDER: EUCYNODONTIA
MAMMALIAMORPHA ROWE, 1988
Superfamily: TRITYLODONTIA Cope, 1884 (sensu Kuhne, 1943)
Family: TRAVERSODONTIDAE Huene, 1936
Sufamily: TRAVERSODONTINAE Huene, 1936
*Luangwa drysdalli
= Scalenodon drysdalli
_____________________________________________________________________________________
EAST AFRICA
NORTHERN ZAMBIA
LOKOKWA RIVER LOCALITY
UPPER NTAWERE FORMATION
LATE ANISIAN STAGE/EARLY LADINIAN STAGE
MID TRIASSIC EPOCH
MIDDLE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPIDA BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: THERIODONTIA OWEN, 1881
EUTHERIODONTIA
SUBORDER: CYNODONTIA OWEN, 1860
EPICYNODONTIA HOPSON & KITCHING, 2001
INFRAORDER: EUCYNODONTIA
MAMMALIAMORPHA ROWE, 1988
Superfamily: TRITYLODONTIA Cope, 1884 (sensu Kuhne, 1943)
Family: TRAVERSODONTIDAE Huene, 1936
Sufamily: TRAVERSODONTINAE Huene, 1936
Luangwa drysdalli
= Scalenodon drysdalli
_____________________________________________________________________________________
ZAMBIA
UPPER LUANGWA VALLEY
LOCALITY 14
N'TAWERE FORMATION
?UPPER ANISIAN STAGE
LOWER MID TRIASSIC EPOCH
LATE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
CLADE NAME: ARCHOSAURIFORMES GAUTHIER, KLUGE & ROWE, 1988
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
PARACROCODYLOMORPHA
LORICATA
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
ZAMBIA
UPPER LUANGWA VALLEY
LOCALITY 15
N'TAWERE FORMATION
?UPPER ANISIAN STAGE
LOWER MID TRIASSIC EPOCH
LATE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LABYRINTHODONTIA OWEN, 1860
SUPERORDER: TEMNOSPONDYLIA ZITTEL, 1887
MICROORDER: LIMNARCHIA YATES & WARREN, 2000
ORDER: BRACHYOPOMORPHA WARREN & MARSICANO, 2000
SUBORDER: BRACHYOPOIDEA ROMER, 1947
Superfamily: BRACHYOPOIDEA Lydekker, 1885 (sensu Save-Soderbergh, 1935)
Family: BRACHYOPIDAE Lydekker, 1885
ORDER: CAPITOSAURIA ROMER, 1947
Superfamily: CAPITOSAUROIDEA Watson, 1919 (sensu Romer, 1947)
Family: MASTODONSAURIDAE Lydekker, 1885
*Cherninia megarhina
= Parotosuchus megarhinus
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
Incertae sedis
Gen. sp indet.
CLADE NAME: ARCHOSAURIFORMES GAUTHIER, KLUGE & ROWE, 1988
Incertae sedis
Gen. sp indet.
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
PARACROCODYLOMORPHA
LORICATA
Incertae sedis
Gen. sp indet.
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
Family: LEWISUCHIDAE PAUL, 1988 emend Olshevsky, 1991
*Lutungutali sitwensis
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPID BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: ANOMODONTIA WATSON & ROMER, 1956
INFRAORDER: DICYNODONTIA OWEN, 1860
Superfamily: PRISTERODONTIA Cluver & King, 1983
Family: KANNEMEYERIDAE Huene, 1948
Subfamily: DINODONTOSAURINAE Kalandadze, 1973
Tribe: RECHNISAURINI Maisch, 2001
*Sanguasaurus edentatus
Subfamily: STAHLECKERIINAE Huene, 1935
Tribe: STAHLECKERINI Lehman, 1961 non sensu Keyeser & Cruckshank, 1979
*Zambiasaurus submersus
ORDER: THERIODONTIA OWEN, 1881
SUBORDER: CYNODONTIA OWEN, 1860
EPICYNODONTIA HOPSON & KITCHING, 2001
INFRAORDER: EUCYNODONTIA
MAMMALIAMORPHA ROWE, 1988
Superfamily: TRITYLODONTIA Cope, 1884 (sensu Kuhne, 1943)
Family: TRIRACHODONTIDAE Crompton, 1955
Cricodon metabolus
Family: TRAVERSODONTIDAE Huene, 1936
Gen. sp indet.
_____________________________________________________________________________________
EAST AFRICA
ZAMBIA
APPROXIMATELY ONE KILOMETERE NORTH OF SITWE IN THE UPPER LUANGWA VALLEY
LOCALITY NO. 16
NTAWERE FORMATION
?UPPER ANISIAN STAGE
LOWER MID TRIASSIC EPOCH
MIDDLE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: SYNAPSIDA OSBORN, 1903
SUPERORDER: THERASPID BROOM, 1905
GRANDORDER: EUTHERAPSIDA
MICRORDER: NEOTHERAPSIDA
ORDER: ANOMODONTIA WATSON & ROMER, 1956
INFRAORDER: DICYNODONTIA OWEN, 1860
Superfamily: PRISTERODONTIA Cluver & King, 1983
Family: KANNEMEYERIDAE Huene, 1948
Subfamily: KANNEMEYERINAE Huene, 1948
*Dolichuranus latirostris
= Kannemeyeria latirostris
= Shansiodon latirostris*Kannemeyeria lophorhinus
= Kannemeyeria cristarhynchus
= Rechnisaurus cristarhynchusORDER: THERIODONTIA OWEN, 1881
SUBORDER: CYNODONTIA OWEN, 1860
EPICYNODONTIA HOPSON & KITCHING, 2001
INFRAORDER: EUCYNODONTIA
Superfamily: CYNOGNATHOIDEA Seeley, 1895
Family: CYNOGNATHIDAE Seeley, 1895
Cynognathus crateronotus
MAMMALIAMORPHA ROWE, 1988
Superfamily: TRITYLODONTIA Cope, 1884 (sensu Kuhne, 1943)
Family: DIADEMODONTIDAE Haughton, 1925
Diademodon tetragonus
Family: TRIRACHODONTIDAE Crompton, 1955
Cricodon metabolus
_____________________________________________________________________________________
NORTHEASTERN ZAMBIA
EASTERN PROVINCE
UPPER LUANGWA VALLEY
UPPER NTAWERE L74
NTAWERE FORMATION
ANSIAN STAGE
LOWER MID TRIASSIC EPOCH
MIDDLE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
Family: SILESAURIDAE Nesbitt, sidor, irmis, angielczyk, Smith & Tsuji, 2019
Gen. sp indet.
_____________________________________________________________________________________
NORTHEASTERN ZAMBIA
EASTERN PROVINCE
UPPER LUANGWA VALLEY
UPPER NTAWERE L75
NTAWERE FORMATION
ANSIAN STAGE
LOWER MID TRIASSIC EPOCH
MIDDLE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LABYRINTHODONTIA OWEN, 1860
SUPERORDER: TEMNOSPONDYLIA ZITTEL, 1887
MICROORDER: LIMNARCHIA YATES & WARREN, 2000
ORDER: STEREOSPONDYLOMORPHA YATES & WARREN, 2000
SUBORDER: STEREOSPONDYLI ZITTEL, 1887-90
Incertae sedis
Gen. sp indet.
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
Family: SHUVOSAURIDAE Chatterjee, 1993
Incertae sedis
Gen. sp indet.
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992Family: UncertainGen. sp indet.
Family: SILESAURIDAE Nesbitt, sidor, irmis, angielczyk, Smith & Tsuji, 2019
Gen. sp indet.
_____________________________________________________________________________________
NORTHEASTERN ZAMBIA
EASTERN PROVINCE
UPPER LUANGWA VALLEY
UPPER NTAWERE L76
NTAWERE FORMATION
ANSIAN STAGE
LOWER MID TRIASSIC EPOCH
MIDDLE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
Family: SILESAURIDAE Nesbitt, sidor, irmis, angielczyk, Smith & Tsuji, 2019
Gen. sp indet.
_____________________________________________________________________________________
NORTHEASTERN ZAMBIA
EASTERN PROVINCE
UPPER LUANGWA VALLEY
UPPER NTAWERE L77
NTAWERE FORMATION
ANSIAN STAGE
LOWER MID TRIASSIC EPOCH
MIDDLE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
Family: SILESAURIDAE Nesbitt, sidor, irmis, angielczyk, Smith & Tsuji, 2019
Gen. sp indet.
_____________________________________________________________________________________
NORTHEASTERN ZAMBIA
EASTERN PROVINCE
UPPER LUANGWA VALLEY
UPPER NTAWERE L78
NTAWERE FORMATION
ANSIAN STAGE
LOWER MID TRIASSIC EPOCH
MIDDLE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
Family: SILESAURIDAE Nesbitt, sidor, irmis, angielczyk, Smith & Tsuji, 2019
Gen. sp indet.
_____________________________________________________________________________________
NORTHEASTERN ZAMBIA
EASTERN PORVINCE
UPPER LUANGWA VALLY
IN THE HILLS OF SITWE
DRYSDALL & KITCHINGS LOCALITY 15
UPPER NTAWERE L 82
NTAWERE FORMATION
ANISIAN STAGE
LOWER MID TRIASSIC
MIDDLE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LABYRINTHODONTIA OWEN, 1860
SUPERORDER: TEMNOSPONDYLIA ZITTEL, 1887
MICROORDER: LIMNARCHIA YATES & WARREN, 2000
ORDER: STEREOSPONDYLOMORPHA YATES & WARREN, 2000
SUBORDER: STEREOSPONDYLI ZITTEL, 1887-90
Incertae sedis
Gen. sp indet.
ORDER: CAPITOSAURIA ROMER, 1947
EUCYCLOTOSAUIRA SCHOCH, 2008
'Stanocephalosaurus pronus'
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
Incertae sedis
Gen. sp indet.
CLADE NAME: ARCHOSAURIFORMES GAUTHIER, KLUGE & ROWE, 1988
Incertae sedis
Gen. sp indet.
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
Family: SILESAURIDAE Nesbitt, sidor, irmis, angielczyk, Smith & Tsuji, 2019
Gen. sp indet.
_____________________________________________________________________________________
NORTHEASTERN ZAMBIA
EASTERN PROVINCE
UPPER LUANGWA VALLEY
UPPER NTAWERE L78
NTAWERE FORMATION
ANSIAN STAGE
LOWER MID TRIASSIC EPOCH
MIDDLE TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
Family: SILESAURIDAE Nesbitt, sidor, irmis, angielczyk, Smith & Tsuji, 2019
Gen. sp indet.
_____________________________________________________________________________________
ZAMBIA
LUANGWA VALLEY
IN THE VICINITY OF LUSITU VILLAGE
BANK OF THE LUSITU RIVER
'RED SANDSTONE'
EARLY JURASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
SUPERORDER: SAUROPODOMORPHA HUENE, 1932
Incertae sedis
Gen. sp indet.
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