MUTATION

Hyphalosaurus lingyuanensis (two headed embryo)


PALAEOPATHOLOY

STEM: TETRAPODA GOODRICH, 1930

Incertae sedis

Genus: Ossinodus WARREN & TURNER, 2004
O. puerorum emend WARREN & TURNER, 2004 (Type)

SUPERCLASS: AMNIOTA HAECKEL, 1866

Incertae sedis

Gen. sp indet.

Bibliography

CLASS: AMPHIBIA LINNAEUS, 1758

SUBCLASS: LABYRINTHODONTIA OWEN, 1860

SUPERORDER: TEMNOSPONDYLIA ZITTEL, 1887

MICROORDER: EUSKELIA YATES & WARREN, 2000

ORDER: EYROPODIA

Superfamily: DISSOROPHOIDEA Boulenger, 1902 (sensu Bolt, 1969)

Family: MICROMELERPETONTIDAE Boy, 1972

Genus: Micromelerpeton BULMAN & WHITTARD, 1926
M. credneri BULMAN & WHITTARD, 1926

ORDER: STEREOSPONDYLOMORPHA YATES & WARREN, 2000

SUBORDER: ARCHEGOSAURIFORMES SCHOCH & MILNER, 2000

Superfamily: ARCHEGOSAUROIDEA Meyer, 1857 (sensu Efremov, 1932)

Family: ACTINODONTIDAE Lydekker, 1885.

Genus: Sclerocephalus GOLDFUSS, 1847
Sp

Biblography

ORDER: CAPITOSAURIA ROMER, 1947

Superfamily: CAPITOSAUROIDEA Watson, 1919 (sensu Romer, 1947)

Incertae sedis

Gen. sp indet.

Biblography

SUBCLASS: CHELONOMORPHA KUHN, 1960

PANTESTUDINES JOYCE, PARHAM & GAUTHIER, 2004

SUPERCOHORT: TESTUNINATA KLEIN, 1760 (sensu JOYCE et al, 2004)

Incertae sedis

Gen. sp indet.

Family: ODONTOCHELYIDAE Li, Wu, Rieppel, Wang, & Zhao, 2008

Genus: Odontochelys LI, WU, RIEPPEL, WANG & ZHAO, 2008
O. semitestacea LI, WU, RIEPPEL, WANG & ZHAO, 2008 (Type)

SUBCLASS: EUREPTILIA OLSON, 1947

ORDER: CAPTORHINOMORPHA WATSON, 1917

SUBORDER: CAPTORHINOMORPHA WATSON, 1917

Family: CAPTORHINIDAE Case, 1911

Subfamily: CAPTORHININAE Case, 1911 (sensu De Ricqles, et AL 1982)

Genus: Labidosaurus COPE, 1896
L. hamatus (COPE, 1896)

SUBCLASS: DIAPSIDA

NEODIAPSIDA: INCERTAE SEDIS

OR AMNIOTA: INCERTAE SEDIS

MAGANORDER: ICHTHYOPTERYGIA BAUR, 1887 emend nova

SUPERORDER: EOICHTHYOSAURIA MOTANI, 1999

GRANDORDER: ICHTHYOSAURIA EMEND BLAINVILLE, 1835

MICROORDER: MERRIAMOSAURIA MONTANI, 1999

ORDER: EUICHTHYOSAURIA MONTANI, 1999

SUBORDER: PARVIPELVIA MONTANI, 1999

INFRAORDER: THUNNOSAURIA MONTANI, 1999

Family: STENOPTERYGIIDAE Jaekel, 1904

Genus: Platypterygius KUHN, 1946
Subgenus: Longirostria ARKHANGELSKY, 1998
(Longirosria) longmani (WADE, 1988) ARKHANGELSKY, 1998 (Type)

Family: OPHTHALMOSAURIDAE Seeley, 1874

Genus: Ophthalmosaurus SEELEY, 1874
O. natans (MARSH, 1879) HUENE, 1922

Bibliography

INFRACLASS: NEODIAPSIDA BENTON, 1985

Incertae sedis

Family: PALIGUANIDAE Broom, 1926

Genus: Heleosuchus BROOM, 1913
H. griesbachii (OWEN, 1876)
= Saurosternon greisbachi OWEN, 1876

Bibliography

ORDER: THALATTOSAURIDAE MERRIAM, 1904

SUBORDER: THALATTOSAUROIDEA MERRIAM, 1904

Family: ENDENNASAURIDAE Rensto, 1984

Genus: Endennasaurus RENESTO, 1984
E. acutirostris RENESTO, 1984 (Type)

Bibliography

 

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

INFRACLASS: EURYAPSIDA COLBERT, 1945

SUPERORDER: SAUROPTERYGIA OWEN, 1860

PARVORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994

GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994

ORDER: PACHYPLEUROSAURIA SANZ, 1980

Family: PACHYPLEUROSAURIDAE Nopcsa, 1928

Genus: Neusticosaurus SEELEY, 1882
N. pusillus (FRAAS, 1881) SEELEY, 1881
N. edwardsi (CORNALIA, 1854) SANDER, 1989

ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835

INFRAORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835

Superfamily: PLESIOSAUROIDEA Wells, 1943

Family: PLESIOSAURIDAE Gray, 1825

Incertae sedis

The following are listed as being pathologic (avascular necrosis) by ROTHSCHILDS & STORRS, 2003 but no other locality or other information is listed.

Severial sauropterygia genera

Genus: Plesiosaurus non CONYBEARE, 1821
P. sp

Superfamily: ELASMOSAUROIDEA nova

Family: ELASMOSAURIDAE Cope, 1869

Genus: Muraenosaurus SEELEY, 1874
M. leedsii SEELEY, 1874
= Cimoliosaurus durobrivensis LYDEKKER, 1889

Genus: Styxosaurus WELLES, 1943
S. snowii (WILLISTON, 1890) WELLES, 1943

Genus: Nova SACHS, HORNUG, WOHLSEIN & KEAR, 2015

Family: LEPTOCLEIDIDAE White, 1940

Genus: Leptocleidus ANDREWS, 1922
L. sp

Incertae sedis

Genus: Ogmodirus WILLISTON & MOODIE, 1913 (nomen dubium)
O. martinii WILLISTON & MOODIE, 1913

Superfamily: PLIOSAUROIDEA Seeley, 1874

Family: PLIOSAURIDAE Seeley, 1874

Genus: Liopleurodon SAUVAGE, 1873
L. macromerus (PHILLIPS, 1871) TARLO, 1959

Genus: Megalneusaurus KNIGHT, 1898
M. rex KNIGHT, 1898

Genus: Peloneustus LYDEKKER, 1889
P. philarchus (SEELEY, 1869) LYDEKKER, 1889
= Pliosaurus evansi SEELEY, 1869

Genus: Pliosaurus OWEN, 1841
P. sp

Genus: Polycotylus COPE, 1869
P. latipinnis COPE, 1869

Bibliography

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

COHORT: SPHENODONTIFORMES Hay, 1930

ORDER: SPHENODONTIA WILLISTON, 1925

Family: BRACHYRHINODONTIDAE Huene, 1948

Genus: Clevosaurus SWINTON, 1939
C. sectumsemper KLEIN, WHITESIDE, LUCAS, VIEGAS, & BENTON, 2015

Family: GEPHYROSAURIDAE Evans, 1980

Genus: Gephyrosaurus EVANS, 1980
G. bridensis EVANS, 1980

Bibliography

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: LACERTILIA OWEN, 1842

Incertae sedis

Gen. sp indet.

GRANDORDER: ASCALABOTA GILMORE, 1928
= DIVISION: ASCALABOTA GILMORE, 1928

ORDER: IGUANIA LATREILLE, 1825

Incertae sedis

Genus: Scandensia EVANS & BARBADILLO, 1998
S. ciervensis EVANS & BARBADILLO, 1998 (Type)

ORDER: LEPTOGLOSSA COPE, 1900

SUBORDER: ZONUROMORPHA ALIFANOV, 2000

INFRAORDER: ZONURIA FURBRINGER, 1900

Superfamily: CORDYLOIDEA Fritzinger, 1826

Family: PARAMACELLODIDAE Estes, 1983

Genus: Becklesius ESTES, 1983
B. hoffstetteri (SEIFFERT, 1973)
= Becklesisaurus hoffstetteri SEIFFERT, 1973

SUBORDER: PLATYNOTA GILMORE, 1928


INFRAORDER: MOSASAURIA MARSH, 1860

Superfamily: MOSASAUROIDEA Gervais, 1853 emend Camp, 1923

Family: Incertae sedis

Subfamily: TETHYSAURINAE Makadi, Caldwell, & Osi, 2012

Genus: Pannoniasaurus MAKADI, CALDWELL & OSI, 2012
P. inexpectatus MAKADI, CALDWELL & OSI, 2012

Family: AIGIALOSAURIDAE Gorjanovic-Kramberger, 18922

Genus: Carentonosaurus RAGE & NERAUDEAU, 2004
C. mineauorum emend RAGE & NERAUDEAU, 2004 (Type)
= Carentonosaurus mineaui RAGE & NERAUDEAU, 2004

Family: MOSASAURIDAE Gervais, 1853

NATANIA

Subfamily: RUSSELLOSAURINAE Bell, 1993

Tribe: PLIOPLATECARPINI Russell, 1967

Genus: Platecarpus COPE, 1869
P. ictericus (COPE, 1871)

Genus: Plioplatecarpus DOLLO, 1882
P. marshi DOLLO, 1882

Gen sp indet.

Subfamily: TYLOSAURINAE Williston, 1897

Genus: Tylosaurus MARSH, 1872
T. kansasensis EVERHART, 2005

Subfamily: MOSASAURINAE Gervais, 1853 (sensu Williston, 1897)

Tribe: MOSASAURINI Gervais, 1853 (sensu Russell, 1967)

Genus: Mosasaurus CONYBEARE, 1822
M. hoffmanni MANTELL, 1829 (Type)

Bibliography

SUPERORDER: LACERTILIA OWEN, 1842

GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961

MICRORDER: SCINCOMORPHA CAMP, 1923

ORDER: ANGUIMORPHA FURBRINGER, 1900

SUBORDER: PYTHONOMORPHA COPE, 1898

INFRAORDER: OPHIDIOMORPHA PALCI & CLADWELL, 2007

PARVORDER: OPHIDIA BRONGNIART, 1800

Superfamily: SERPENTES Linnaeus, 1759

Family: ANILIIDAE Fitzenger, 1826

Subfamily: BOOIDINAE Gray, 1825

Infrafamily: BOOINIEA Gray, 1925

Tribe: MATSOINI Hoffstetter, 1961

Genus: Madtsoia SIMPSON, 1933
M. madagascariensis HOFFSTETTER, 1961

LEGION: ARCHOSAUROMORPHA HUENE, 1946

Incertae sedis

Gen. sp indet.

SUPERCOHORT: AVEMETATARSALIA BENTON, 1999

COHORT: PROLACERTIFORMES CAMP, 1945 nova

MAGNORDER: PROLACERTIFORMES CAMP, 1945 NOVA

SUPERORDER: TAPINOPLATIA PETERS, 2000

GRANDORDER: CHARACIOPODA PETERS, 2000

MICRORDER: FENESTRASAURIA PETERS, 2000

ORDER: PTEROSAURIA KAUP, 1834

Incertae sedis

Gen. sp indet.

SUBORDER: PTERODACTYLOIDEA PLIENINGER, 1901

Superfamily: PTERODACTYLOIDEI Orlov, 1964

Family: PTERODACTYLIDAE Bonaparte, 1838

Genus: Pterodactylus CUVIER, 1809
P. kochi (WAGNER, 1837)

SUBORDER: EUPTERODACTYLOIDEA BENNETT, 1994

Family: ANHANGUERIDAE Campos & Kellner, 1985

Genus: Anhanguera CAMPOS & KELLNER 1985
A. araripensis (WELLNHOFER, 1985) KELLNER & TOMIDA, 2000
= Santanadactylus araripensis WELLNHOFER, 1985

Genus: Siroccopteryx MADER & KELLNER, 1999
S. moroccensis MADER & KELLNER, 1999 (Type)
= Coloborhynchus moroccensis (MADER & KELLNER, 1999) UNWIN, 2001

Superfamily: AZHDARCHOIDEA Nesov, 1984

NEOAZHDARCHIA UNWIN, 2003

Family: TAPEJARIDAE Kellner, 1989

Subfamily: TAPEJARINAE Kellner, 1989

Genus: Caupedactylus KELLNER, 2013 (for 2012)
C. ybaka KELLNER, 2013 (for 2012) (Type)

Superfamily: PTERANODONTIDEA Marsh, 1876

Family: PTERANODONTIDAE Marsh, 1876

Genus: Pteranodon MARSH, 1876
P. ingens (MARSH, 1872) MARSH, 1876
P. longiceps MARSH, 1876
= Pteranodon marshi (MILLER, 1972)

Bibliography

SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990

COHORT: CROCODYLOTARSI BENTON & CLARK, 1988

SUPERORDER: PARASUCHIMORPHA NOVA

ORDER: PARASUCHIA HUXLEY, 1875

Family: ANGISTORHINIDAE Huene, 1936

Subfamily: PSEUDOPALATINAE, Long & Murry, 1995

Genus: Pseudopalatus MEHL, 1928
P. buceros (COPE, 1881)

Bibliography

MAGNORDER: SUCHIA KREBS, 1974

SUPERORDER: CROCODYLOMORPHA WALKER, 1968

ORDER: MESOSUCHIA emend HUXLEY, 1875

SUBORDER: THALATTOSUCHIA emend FRAAS, 1901

Family: METRIORHYNCHIDAE Fitzinger, 1843

Subfamily: METRIORHYNCHINAE Fitzinger, 1843

Genus: Metriorhynchus MEYER, 1830
M. superciliosum (De BLAINVILLE, 1853) DESLONGCHAMPS, 1869

Biblography:

PARVORDER: METASUCHIA BENTON & CLARK, 1988

NEOSUCHIA CLARK, IN BENTON & CLARK, 1988

INFRAORDER: TETHYSUCHIA BUFFETAUT, 1982

Family: DYROSAURIDAE de STEFANO, 1903

Gen. sp indet.

MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983

ORDER: MESOSUCHIA EMEND HUXLEY, 1875
PARVORDER: METASUCHIA BENTON & CLARK, 1988
INFRAORDER: SEBECOSUCHIA SIMPSON, 1937 (emend. GASPARINI, 1971)
Family: BAURUSCHIDAE Price, 1945

Genus: Stratiotosuchus CAMPOS, SUAREZ, RIFF & KELLNER, 2001
S. maxhechti CAMPOS, SUAREZ, RIFF & KELLNER, 2001

ORDER: EUSUCHIA emend HUXLEY, 1875

Family: BOREALOSUCHIDAE emend BROCHU, 1999

Genus: Borealosuchus BROCHU, 1997
B. formidabilis (ERICKSON, 1976) BROCHU, 1997
= Leidyosuchus formidabilis ERICKSON, 1976

SUBORDER: CROCODYLIA emend GMELIN, 1877

Family: GAVIALIDAE Cuvier, 1807

Subfamily: THORACOSAURINAE Cope, 1875 (sensu Nopcsa, 1928)

Genus: Toyotamphiamela AOKI, 1983
T. machikanense (KAMOI & MATSUMOTO, 1965) AOKI, 1983

INFRAORDER: BREVIROSTRES

Superfamily: ALLIGATORIDEA Gray, 1844 ( sensu Buffetaut, 1985)

Family: DIPLOCYNODONTIDAE emend Brouchu, 1999

Genus: Diplocynodon POMEL, 1847
= Diplyocynodus POMEL, 1847 (sic)
= Hispanochampsa KALIN, 1936
= Orthosaurus GREFFROY, 1833
= Plerodon MEYER, 1839
= Pleurodon MEYER, 1839 (sic)
= Saurocainus GERVIAS 1952
D. hantoniensis (WOOD, 1844)

Superfamily: CROCODILOIDEA Fitzinger, 1826

Family: CROCODYLIDAE Cuvier, 1807

Gen sp indet

Subfamily: TOMISTOMINAE Kalin, 1955

Genus: Gavialosuchus TOULA & KAIL, 1885
= Garialosuchus BOULENGER, 1886 (sic)
G. carolinensis ERICKSON & SAWYER, 1996

Subfamily: OSTEOLAEMINAE

Genus: Voay BROCHU, 2007
V. robustus (GRANDIDIER & VAILLANT, 1872) BROCHU, 2007 (Type)
= Crocodilus robustus GRANDIDIER & VAILLANT, 1872

Bibliography

LEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985

SUBLEGION: DINOSAURIFORMES NOVAS, 1992

SUPERCOHORT: DINOSAURIA OWEN, 1842

Gen. Sp indet.

SUPERORDER: HERRASAURA NOVA

ORDER: HERRERASAURIA GALTON, 1985

Family: HERRERASAURIDAE Benedetto, 1973

Genus: Herrerasaurus REIG, 1963
H. ischigualastensis REIG, 1963

COHORT: THEROPODA EMEND MARSH, 1881

Incertae sedis

Gen. sp indet.

SUPERORDER: NEOTHEROPODA BAKKER, 1986

GRANDORDER: CERATOSAURIA NOVAS, 1992

ORDER: CERATOSAURIA MARSH, 1884

Superfamily: CERATOSAUROIDEA Bonaparte, 1991

Family: CERATOSAURIDAE Marsh, 1884 emend. Gilmore, 1920

Genus: Ceratosaurus MARSH, 1884
C. nasicornis MARSH, 1884
C. dentisulcatus MADSEN & WELLES, 2000

Superfamily: ABELISAUROIDEA Novas, 1989

Family: ABELISAURIDAE Bonaparte & Novas, 1985

Subfamily: CARNOTAURINAE Sereno, 1998

Tribe: CARNOTAURINI Coria, Chiappe & Dingus, 2002

Genus: Aucasaurus CORIA, CHAIPPE & DINGUS, 2002
A. garridoi CORIA, CHIAPPE & DINGUS, 2002

Genus: Majungatholus SUES & TAQUET, 1979
M. atopus SUES & TAQUET, 1979

ORDER: COELOPHYSIA NOVA

SUBORDER: COELOPHYSOIDEA (NOPCSA, 1928) HOLTZ, 1994

Superfamily: PODOKESAUROIDEA Madsen & Welles, 2000

Family: PODOKEOSAURIDAE Huene, 1914

Subfamily: COELOPHYSINAE Nopcsa, 1928

Genus: Syntarsus RAATH, 1969
S. rhodesiensis RAATH, 1969

Subfamily: PROCOMPSOGNATHINAE (HUENE, 1929) Nopcsa, 1923

Genus: Procompsognathus E. FRAAS, 1913
P. triassicus E. FRAAS, 1913

Family: DILOPHOSAURIDAE Charig & Milner, 1990

Subfamily: DILOPHOSAURINAE Madsen & Welles, 2000

Genus: Dilophosaurus WELLES, 1970
D. wetherilli (WELLES, 1954) WELLES, 1970

ORDER: NEOTETANURAE SERENO, WILSON, LARSSON, DUTHEIL & SUES, 1994

GRANDORDER: TETANURAE GAUTHIER, 1986

ORDER: SPINOSAURIA OLSHEVSKY, 1991

Superfamily: SPINOSAUROIDEA Stromer, 1915

Family: SPINOSAURIDAE Stromer, 1915

Genus: Spinosaurus STROMER, 1915
S. maroccanus RUSSELL 1996

Superfamily: MEGALOSAUROIDEA Huxley, 1889

Family: MEGALOSAURIDAE Huxley, 1889

Genus: Megalosaurus BUCKLAND, 1824
M. bucklandii RITGEN, 1826

Genus: Monolophosaurus ZHAO & CURRIE, 1994
M. jiangi ZHAO & CURRIE, 1994

Genus: Poekilopleuron EUDES-DESLONGCHAMPS, 1838
P. bucklandii EUDES-DESLONGICHAMPS, 1838

SUBORDER: AVETHEROPODA PAUL, 1988

Superfamily: ALLOSAUROIDEA (Marsh, 1878) Currie & Zhao, 1994

Family?

Genus: Megaraptor NOVAS, 1998
M. namunhuaiquii NOVAS, 1998

Family: SINRAPTORIDAE Currie & Zhao, 1994

Genus: Sinraptor CURRIE & ZHAO, , 1994
S. hepingensis (GAO, 1992)
= Yangchuanosaurus hepingensis GAO, 1992
S. dongi CURRIE & ZHAO, 1994

Family: CARCHARODONTOSAURIDAE Stromer, 1931

Genus: Carcharodontosaurus STROMER, 1931
C. saharicus (DEPERET & SAVORNIN, 1925) STROMER, 1931

Genus: Mapusaurus CORIA & CURRIE, 2006
M. roseae CORIA & CURRIE, 2006 (Type)

Genus: Veterupristisaurus RAUHUT, 2011
V. milneri RAUHUT, 2011 (Type)

Gen. sp indet.

Family: ALLOSAURIDAE Marsh, 1878

Genus: Allosaurus MARSH, 1877
A. fragilis MARSH, 1877
= Antrodemus fragilis (MARSH, 1877)
= Labrosaurus fragilis (MARSH, 1877)
= Labrosaurus ferox MARSH, 1884

Genus: Neovenator HUTT, MARTILL & BARKER, 1996
N. salerii HUTT, MARTILL & BARKER, 1996

Family: ACROCANTHOSAURIDAE Molnar, 2001

Genus: Acrocanthosaurus STOVALL & LANGSTON, 1955
A. atokensis STOVALL & LANGSTON, 1955

MICRORDER: COELUROSAURIA HUENE, 1914

Family?

Genus: Nova: CORIA, CURRIE, EBERTH, GARRIDO & KOPPELHUS, 2001

ORDER: MANIRAPTORFORMES HOLTZ, 1996

AVETHEROPODA PAUL, 2002

SUBORDER: COELURIA MARSH, 1881

Family: COMPSOGNATHIDAE Cope, 1875

Genus: Compsognathus WAGNER, 1861
C. longipes WAGNER, 1861 (Type)
= Compsognathus corallestris BIDAR, DEMAY & THOMEL, 1972

Family: COELURIDAE Marsh, 1881

Genus: Marshosaurus MADSEN, 1976
M. bicentesimus MADSEN, 1976

Superfamily: TYRANNOSAUROIDEA (Osborn, 1905) Bonaparte ET AL, 1990

Incertae sedis

Genus: Appalachiosaurus CARR, WILLIAMSON, & SCHWIMMER, 2005
= Genus: Nova? KING, ABBOTT-KING, BELL, LAMB, DOBIE & WOMOCHEL, 1988
A. montgomeriensis CARR, WILLIAMSON, & SCHWIMMER, 2005 (Type)

Family: TYRANNOSAURIDAE Osborn, 1906

Gen. sp indet

Subfamily: ALBERTOSAURINAE Currie, Hurum & Sabath, 2003

Genus: Albertosaurus OSBORN, 1905
A. sarcophagus OSBORN, 1905
= Albertosaurus arctunguis PARKS, 1928
A. sp

Genus: Gorgosaurus LAMBE, 1914
G. libratus LAMBE, 1914
G. sp

Tribe: TARBOSAURINI Olshevsky, Ford & Yamamoto, 1995

Genus: Jenghizkhan (MALEEV, 1955) OLSHEVSKY, FORD & YAMAMOTO, 1995
J. bataar (MALEEV, 1955) OLSHEVSKY, FORD & YAMAMOTO, 1995

Tribe: TYRANNOSAURINI Olshevsky, Ford & Yamamoto, 1995

Genus: Daspletosaurus RUSSELL, 1970
D. torosus RUSSELL, 1970
D. Species Nova CARR & WILLIAMSON, 1999
D. Species Nova WILLIAMSON & CARR, 1999
D. sp
= Aublysodon sp LEHMAN & CARPENTER 1990

Genus: Nanotyrannus BAKKER, WILLIAMS & CURRIE, 1988
= Clevelanotyrannus BAKKER, WILLIAMS & CURRIE in CURRIE, 1987 (nomen nudum)
Note: Clevelanotyrannus was a name in a citation by BAKKER, et al in CURRIE, 1987 (later it was described as Nanotyrannus).
N. lancensis (GILMORE, 1946) BAKKER, WILLIAMS & CURRIE, 1988

Genus: Tyrannosaurus OSBORN, 1905
T. rex OSBORN, 1905

SUBORDER: ORNITHOMIMOSAURIA BARSBOLD, 1976

Family: ORNITHOMIMIDAE Marsh, 1890

Genus: Timimus RICH & RICH 1994
T. hermani RICH & RICH, 1994

Gen sp indet.

Family: DEINOCHERIDAE Osmolska & Roniewicz, 1970

Genus: Deinocheirus OSMOLOSKA & RONIEWICZ, 1970
D. mirificus OSMOLOSKA & RONIEWICZ, 1970

INFRAORDER: OVIRAPTOROSAURIA BARSBOLD, 1976

Family: OVIRAPTORIDAE Barsbold, 1976

Subfamily: OVIRAPTORINAE Barsbold, 1981

Genus: Citipati CLARK, NORELL & BARSBOLD, 2001
C. osmolskae CLARK, NORELL & BARSBOLD, 2001

SUBORDER: MANIRAPTORA GAUTHIER, 1986

INFRAORDER: PARVES SERENO, 1997

PARVORDER: DEINONYCHOSAURIA COLBERT & D. A. RUSSELL, 1969

Family: DROMAEOSAURIDAE D. A. Russell, 1969

Subfamily: MICROPTORINI Senter, Barsbold, Britt, & Burnham, 2004

Genus: Bambiraptor BURNHAM, DERSTLER, CURRIE, BAKKER, ZHOU & OSTROM, 2000
B. feinbergorum emend BURNHAM, DERSTLER, CURRIE, BAKKER, ZHOU & OSTROM, 2000

Genus: Hesperonychus LONGRICH & CURRIE, 2009
H. elizabethae LONGRICH & CURRIE, 2009 (Type)

Subfamily: VELOCIRPATORINAE Barsbold, 1978

Genus: Deinonychus OSTROM, 1969
D. antirrhopus OSTROM, 1969

Genus: Saurornitholestes SUES, 1978
S. langstoni SUES, 1978

Genus: Velociraptor OSBORN, 1924
V. mongoliensis OSBORN, 1924

Family: TROODONTIDAE Gilmore, 1924

Genus: Sinornithoides RUSSELL & DONG, 1993
S. youngi RUSSELL & DONG, 1993

Genus: Talos ZANNO, VARRICCHIO, O’CONNOR, TITUS, & KNELL, 2011
T. sampsoni ZANNO, VARRICCHIO, O’CONNOR, TITUS, & KNELL, 2011 (Type)

Genus: Troodon LEIDY, 1856
T. formosus LEIDY, 1856
= Stenonychosaurus inequalis STERNBERG, 1932

COHORT: ORNITHISCHIFORMES COOPER, 1985

SUPERORDER: PROSAUROPODAMORPHA Nova

ORDER: PROSAUROPODA HUENE, 1920 (emend)

SUBORDER: PLATEOSAURIA TORNIER, 1913

Family: PLATEOSAURIDAE Marsh, 1895

Genus: Plateosaurus MEYER, 1837
P. sp

SUPERORDER: SAUROPODOMORPHA HUENE, 1932

Family: MELANOROSAURIDAE Huene, 1929

Genus: Camelotia GALTON, 1985
C. borealis GALTON, 1985

GRANDORDER: BRONTOSAURIA OLSHEVSKY, 1991 (emend)

Family: MASSOSPONDYLIDAE Huene, 1914

Genus: Massospondylus OWEN, 1854
M. sp

MICRORDER: SAUROPODA MARSH, 1878

Incertae sedis

Genus: Coloradisaurus LAMBERT, 1983
C. brevis (BONAPARTE, 1978) LAMBERT, 1983
= Coloradia brevis BONAPARTE, 1978

Gen. sp indet.

Family: VULCANODONTIDAE Cooper, 1984

Genus: Vulcanodon RAATH, 1972
V. karibaensis RAATH, 1972

ORDER: EUSAUROPODA

SUBORDER: NEOSAUROPODA BONAPARTE, 1986

INFRAORDER: HOMALOSAUROPIA JANENSCH, 1929

Family: CAMARASAURIDAE Cope, 1877

Incertae sedis

Genus: Nurosaurus DONG, 1992 (nomen nudum)
N. qaganensis DONG, 1992 (Type)

Subfamily: CAMARASAURINAE Cope, 1877 (sensu Nopcsa, 1928)

Genus: Camarasaurus COPE, 1877
C. supremus COPE, 1877
C. grandis (MARSH, 1877) GILMORE, 1925
C. sp

PARVORDER: DIPLODOCIMORPHA CALVO & SALGADO, 1995

Superfamily: DIPLODOCOIDEA Upchurch, 1995

Family: DIPLOCODIDAE Marsh, 1884

Genus: Apatosaurus MARSH, 1877
A. sp

Genus: Brontosaurus MARSH, 1879
B. louisae (HOLLAND, 1915)
B. sp

Genus: Barosaurus MARSH, 1890
B. sp

Genus: Diplodocus MARSH, 1878
D. longus MARSH, 1878
D. hallorum (GILLETTE, 1991) (CURTICE, in press as per LUCAS & HUNT, 2000)
= Seismosaurus halli
GILLETTE, 1991
= Seismosaurus hallorum
(GILLETTE, 1991) (emend OLSHEVSKY, 1998)
D. sp

Gen. sp indet.

INFRAORDER: MACRONARIA WILLSON & SERENO, 1998

PARVORDER: TITANOSAURIFORMES SALGADO ET AL, 1997

Family: BRACHIOSAURIDAE Riggs, 1904

Subfamily: BRACHIOSAURINAE Riggs, 1904 (sensu Janensch, 1929)

Genus: Brachiosaurus RIGGS, 1903
B. sp

Superfamily: SOMPHOSPONDYLI Wilson & Sereno, 1998

Family: TITANOSAURIDAE Lydekker, 1885

Incertae sedis

Pathologic eggs

Subfamily: ANDESAURINAE Calvo & Bonaparte, 1991

Genus: Argentinosaurus BONAPARTE & CORIA, 1993
A. huinculensis BONAPARTE & CORIA, 1993

MAGNORDER: ORNITHISCHIA SEELEY, 1888

SUPERORDER: NEORNITHISCHIA COOPER, 1985

GRANDORDER: ORNITHOPODA MARSH, 1871

MICRORDER: EUORNITHOPODA SERENO, 1986

ORDER: HYPSILOPHODONTIA COOPER, 1983

Family: HYPSILOPHODONTIDAE Dollo, 1882

Incertae sedis

Gen. sp indet.

ORDER: IGUANODONTIA DOLLO, 1888

SUBORDER: DRYOMORPHA SERNEO, 1986

Family: DRYOSAURIDAE Milner & Norman, 1984

Dryosaurus or Valdosaurus?

Drysalatosaurus lettowvorbecki

SUBORDER: ANKYLOPOLLEXIA SERENO, 1986

Family: CAMPTOSAURIDAE Marsh, 1885

Genus: Camptosaurus MARSH, 1885
C. dispar
C. sp

PARVORDER: HADROSAURIFORMES SERENO, 1997

Superfamily: IGUANODONTIDEA Sereno, 1986 (emend)

Incertae sedis

Genus: Iguanodon non MANTELL, 1825
I. hollingtoniensis LYDEKKER, 1889

Family: IGUANODONTIDAE Cope, 1869

Genus: Iguanodon MANTELL, 1825
I. bernissartensis BOULENGER vide van BENEDEN, 1881
I? atherfieldensis HOOLEY, 1925
I. sp

Genus: Lurdusaurus TAQUET & RUSSELL, 1999
L. arenatus TAQUET & RUSSELL, 1999

Superfamily: HADROSAUROIDEA Sereno, 1986

Family: HADROSAURIDAE COPE, 1869

Incertae sedis

Gen. sp indet.

Tribe: EDMONTOSAURINI Brett-Surman vide Chapman & Brett-Surman, 1990

Genus: Edmontosaurus LAMBE, 1917
E. annectens (MARSH, 1892) BRETT-SURMAN vide CHAPMAN & BRETT-SURMAN, 1990
= Thespesius edmontonensis GILMORE, 1924
E. sp

Tribe: SAUROLOPHINI Brett-Surman vide Chapman & Brett-Surman, 1990

Genus: Prosaurolophus BROWN, 1916
P. maximus BROWN, 1916

Tribe: KRITOSAURINI nova

Genus: Gryposaurus LAMBE, 1914
G. notabilis LAMBE, 1914

Genus: Naashoibitosaurus HUNT & LUCAS, 1993
N. ostromi HUNT & LUCAS, 1993

Tribe: MAIASAURINI nova

Genus: Maiasaura HORNER & MAKELA, 1979
M. peeblesorum HORNER & MAKELA, 1979

Subfamily: LAMBEOSAURINAE Huene, 1948

Genus: Barsboldia MARYANSKA & OSMOLSKA, 1981
B. sicinskii MARYANSKA & OSMOLSKA, 1981

Gen. sp indet.

Tribe: LAMBEOSAURINI Parks, 1923

Genus: Arenysaurus PEREDA-SUBERBIOLA, CANUDO, CRUZADO-CAGALLERO, BARCO, LOPEZ-MARTINEZ, OMS, & RUIZ-OMENACA, 2009
A. ardevoli PEREDA-SUBERBIOLA, CANUDO, CRUZADO-CAGALLERO, BARCO, LOPEZ-MARTINEZ, OMS, & RUIZ-OMENACA, 2009 (Type)

Genus: Lambeosaurus PARKS, 1923
L? laticaudus MORRIS, 1981

Genus: Nipponosaurus NAGAO, 1936
N. sachaliensis NAGAO, 1936

Tribe: PARASAUROLOPHINI Brett-Surman vide Chapman & Brett-Surman, 1990

Genus: Parasaurolophus PARKS, 1922
P. walkeri PARKS, 1922 (Male)
P. cyrtocristatus OSTROM, 1961 (Female)

MICRORDER: MARGINOCEPHALIA SERENO, 1986

ORDER: PACHYCEPHALOSAURIA MARYANSKA & OSMOLSKA, 1974

SUBORDER: DOMOCEPHALINIA SERENO, 1986

INFRAORDER: PACHYCEPHALOSAURIA (MARYANSKA & OSMOLSKA, 1974) COOPER, 1985

Family: PACHYCEPHALOSAURIDAE Sternberg, 1945

Subfamily: PACHYCEPHALOSAURINAE Sternberg, 1945 (sensu Sereno, 1998

Genus: Pachycephalosaurus BROWN & SCHLAIKJER, 1943 (nomen conservandum)
P. wyomingensis (GILMORE, 1931) BROWN & SCHLAIKJER, 1943

Subfamily: STEGOCERINAE Sereno, 1998 (emend)

Genus: Stegoceras LAMBE, 1902
S. sp

ORDER: CERATOPIA MARSH, 1890

SUBORDER: NEOCERATOPIA SERENO, 1986

Family: PSITTACOSAURIDAE Osborn, 1923

Genus: Psittacosaurus OSBORN, 1923
P. neimongoliensis RUSSELL & ZHAO, 1996
P. sp

PARVORDER: CERATOPOMORPHA Wolfe & Kirkland, 1998 (emend)

Superfamily: CERATOPOIDEA Hay, 1902 (emend)

Incertae sedis

Gen. sp indet.

Family: CERATOPIDAE Marsh, 1888 (emend)

Subfamily: PACHYRHINOSAURINAE C. M. Sternberg, 1950

Genus: Pachyrhinosaurus STERNBERG, 1950
P. nova?

Genus: Nova? TANKE & FARKE, 2003

Subfamily: CERATOPIINAE Abel, 1919 (emend)

Subfamily: CENTROSAURINAE Lambe, 1915

Genus: Centrosaurus LAMBE, 1904
C. apertus LAMBE, 1904
C. sp

Genus: Einiosaurus SAMPSON, 1995
E. procurvicornis SAMPSON, 1995

Genus: Eoceratops LAMBE, 1915
E. canadensis (LAMBE, 1902) LAMBE, 1915

Genus: Monoclonius COPE, 1876 (nomen dubium)
M. ? lowei STERNBERG, 1940

Genus: Styracosaurus LAMBE, 1913
S. albertensis LAMBE, 1913
= Styracosaurus parksi BROWN & SCHLAIKJER, 1937
S. sp

Genus: Wendiceratops EVANS & RYAN, 2015
W. pinhornensis EVANS & RYAN, 2015 (Type)

Tribe: CHASMOSAURINI nova

Genus: Anchiceratops BROWN, 1914
A. ornatus BROWN, 1914 (Type)
A. sp

Genus: Arrhinoceratops PARKS, 1925
A. brachyops PARKS, 1925

Genus: Chasmosaurus LAMBE, 1914
C. belli (LAMBE, 1902) LAMBE, 1914
C? russelli STERNBEG, 1940
C. irviensis HOLMES, FORSTER, RYAN & SHEPHERD, 2001
C. sp

Genus: Eotriceratops WU, BRINKMAN, EBERTH & BRAMAN, 2007
E, xerinsularis WU, BRINKMAN, EBERTH & BRAMAN, 2007

Genus: Pentaceratops OSBORN, 1923
P. fenestratus WIMAN, 1930

Gen. sp indet.

Tribe: TOROSAURINI nova

Genus: Torosaurus MARSH, 1889
T. latus MARSH, 1891

Tribe: TRICERATOPINI nova

Genus: Triceratops MARSH, 1889
T. horridus (MARSH, 1889) MARSH, 1889
= Triceratops serratus MARSH, 1890
= Triceratops elatus MARSH, 1891
= Triceratops sulcatus MARSH, 1890 (nomen dubium)
= Diceratops hatcheri LULL, 1907
T. prorsus MARSH, 1890
T. sp

SUPERORDER: EURYOPDA SERENO, 1986

GRANDORDER: ARMATOSAURIA ZHAO, 1983 (emend)

ORDER: ANKYLOSAURIA OSBORN, 1923

INFRAORDER: ANKYLOSAURA OSBORN, 1923

Family: NODOSAURIDAE Marsh, 1890

Subfamily: EDMONTONIINAE Russell, 1940

Genus: Chassternbergia (BAKKER, 1988) OLSHEVSKY, 1991
C. rugosidens (GILMORE, 1930 ) OLSHEVSKY, 1991
C. sp
= Edmontonia rugosidens or new species WATABE, NAKAYA, & NAKAHARA, 2000

Genus: Edmontonia C. M. STERNBERG, 1928
E. sp

Subfamily: SAUROPELTINAE Ford, 2000

Genus: Sauropelta OSTROM, 1970
S. edwardsorum OSTROM, 1970 (Type)

Family: POLACANTHIDAE Weiland, 1911

Genus: Gastonia KIRKLAND, 1998
G. burgei KIRKLAND, 1998

Genus: Polacanthus FOX, 1865 (OWEN, vide HULKE, 1881)
P. foxii HULKE, 1881 (Type)

Family: ANKYLOSAURIDAE Brown, 1908

Incertae sedis

Genus: Heishansaurus BOHLIN, 1953 (nomen dubium)
H. pachycephalus BOHLIN, 1953 (Type)

Subfamily: ANKYLOSAURINAE Brown, 1908

Genus: Euoplocephalus LAMBE, 1910
E. tutus (LAMBE, 1902) LAMBE, 1910

Genus: Scolosaurus NOPCSA, 1928
S. cutleri NOPCSA, 1928

Subfamily: SYRMOSAURINAE Maleev, 1952

Genus: Talarurus MALEEV, 1952
T. plicatospineus MALEEV, 1952 (Type)

Genus: Tarchia MARYANSKA, 1977
T. kielanae MARYANSKA, 1977

Subfamily: SHASMOSAURINAE Tumanova, 1983

Genus: Shamosaurus TUMANOVA, 1983
S. scutatus TUMANOVA, 1983

Subfamily: STEGOPELTINAE Ford, 2000

Genus: Aletopelta FORD & KIRKLAND, 2001
A. coombsi FORD & KIRKLAND, 2001 (Type)

Genus: Stegopelta WILLISTON, 1905
S. landerensis WILLISTON, 1905

GRANDORDER: STEGOSAUROMORPHA COOPER, 1985 (emend)

ORDER: STEGOSAURIA MARSH, 1877

Family: HUAYANGOSAURIDAE Galton, 1990

Genus: Huayangosaurus DONG, TANG & ZHOU, 1982
H. taibaii DONG, TANG & ZHOU, 1982

Family: STEGOSAURIDAE Marsh, 1877

Subfamily: STEGOSAURINAE ABEL, 1919

Genus: Stegosaurus MARSH, 1877
S. ungulatus MARSH, 1879
S. stenops MARSH, 1887

Biblography

CLASS: AVES LINNAEUS, 1758

AVEBREVICAUDA PAUL, 2001
SUBCLASS: CARINATE MERREM, 1813
INFRACLASS: NEORNITHES GADOW, 1893
PARVCLASS: NEOGNATHAE PYCRAFT 1900
ORDER: GAVIIFORMES WETMORE & MILLER, 1926
Family: GAVIIDAE Allen, 1897

Genus: Polarornis CHATTERJEE, 2002
P. gregorii CHATTERJEE, 2002

VETEROVATA
(Parataxonomy of fossil eggs and shells)

OOCOHORT: THEROPODA NOVA
(Eggshells of the ornithoid basic type [Theropoda])

(EGGS WITH DINOSAUROID-PRISMATIC BASIC SHELL TYPE)

oofamily: PRISMATOOLITHIDAE Hirsch, 1994

Oogenus: Preprismatoolithus ZELENITSKY & HILLS, 1996
P. coloradensis (HIRSCH, 1994) ZELENITSKY & HILLS, 1996

OOCOHORT: ORNITHISCHIFORMES NOVA

OOMAGNORDER: ORNITHISCHIA SEELEY, 1888
oofamily: SPHEROOLITHIDAE Zhao, 1979

Incerate sedis

Oogenus: Triprismatoolithus JACKSON & VARRICCHIO, 2010
T. stephensi JACKSON & VARRICCHIO, 2010

OOCOHORT: ORNITHISCHIFORMES NOVA

OOSUPERORDER: SAUROPODOMORPHA NOVA
(TUBOSPHERULITIC STRUCTURAL MORPHOTYPE)

oofamily: "MEGALOOLITHIDAE" Zhao, 1979
[Sauropoda]

Oogenus: Megaloolithus VIANEY-LIAUD, MALLAN, BUSCALI & MONTGELARD, 1994
M. mammilare VIANEY-LIAUD, MALLAN, BUSCALI & MONTGELARD, 1994

Bibliography

SUBCLASS: SYNAPSIDA OSBORN, 1903

ORDER: PELYCOSAURIA COPE, 1878

SUBORDER: EUPELYCOSAURIA KEMP, 1982

INFRAORDER: DIMETRODONTOMORPHA emend NOPCSA, 1925

Family: SPHENACODONTIDAE Marsh, 1878

Subfamily: SPHENACODONTINAE Nopcsa, 1925

Tribe: DIMETERODONTINI nova

Genus: Dimetrodon COPE, 1878
D. gigashomogenes CASE, 1907 (as per REGA, NORIEGA, SUMIDA, HUTTENLOCKER, LEE & KENNEDY, 2012)
D. sp

Bibliography

SUPERORDER: THERASPID BROOM, 1905

GRANDORDER: EUTHERAPSIDA

MICRORDER: NEOTHERAPSIDA

ORDER: ANOMODONTIA WATSON & ROMER, 1956

SUBORDER: DICYNODONTIA OWEN, 1860

Superfamily: EODICYNODONTIA Barry, 1974 (sensu Cluver & King, 1983)

Family: EODICYNODONTIDAE Barry, 1974

Genus: Eodicynodon BARRY, 1974
E. oosthuizeni BARRY, 1964

Superfamily: PRISTERODONTIA Cluver & King, 1983

Family: KANNEMEYERIDAE Huene, 1948

SUBFAMILY: STAHLECKERIINAE HUENE, 1935

TRIBE: STAHLECKERINI HUENE, 1935 (SENSU LEHMAN, 1961 NON SENSU KEYESER & CRUCKSHANK, 1979)

Genus: Jachaleria BONAPARTE, 1971
J. candelariensis ARAUJO & GONZASA, 1978

Superfamily: DIICTODONTIA Cluver & King, 1983

Family: EMYDOPOIDEA Cluver & King, 1983

Subfamily: EMYDOPINAE Cluver & King, 1983

Genus: Emydops BROOM, 1912
E. oweni FRÖBISH & RESIZ, 2008

Bibliography

ICHNOSTEM: TETRAPODIPIDA NOVA

ICHNOLEGION: DINOSAUROMORPHIPEDA NOVA

ICHNOSUBLEGION: DINOSAURIFORMEIPEDIDII NOVA

ICHNOSUPERCOHORT: DINOSAUROPEDIDA NOVA

ICHNOCOHORT: THEROPODIPEDIA NOVA

Incertae sedis

Gen. sp indet.

Ichnofamily: GRALLATORIDAE emend Lull, 1904

Ichnogenus: Anchisauripus LULL, 1904
A. sp

ICHNOSUPERORDER: NEOTHEROPODOPEDIDA NOVA

ICHNOORDER: NEOTETANURIPEDIDII NOVA

ICHNOSUBORDER: AVETHEROPODIPIDA NOVA

ICHNOMICRORDER: COELUROSAURIPIDIDA NOVA

ICHNOORDER: MANIRAPTORFORMIPIDA NOVA

ICHNOSUBORDER: MANIRAPTORIPIDA NOVA

ICHNOINFRAORDER: PARVEPIDA NOVA

ICHNOPARVORDER: DEINONYCHOSAURIPEDII NOVA

ICHNOFAMILY: DROMAEOPODIDAE LI, LOCKLEY, MAKOVICKY, MATSUKAWA, NORELL, HARRIS & LIU, 2008

Gen. sp indet.

ICHNOCOHORT: ORNITHISCHIFORMEIPIDA NOVA

ICHNOSUPERORDER: PROSAUROPODAMORPHIPIDA NOVA

ICHNOORDER: PROSAUROPODINA VYLOV, 1988

Incertae sedis

Ichnogenus: Pseudotetrasauropus ELLENBERGER, 1972
P-like track GASTON, LOCKLEY, LUCAS & HUNT, 2005
Note: Digit II is extremly short, I propose the toe is pathologic and is missing from the foot.

ICHNOINFRAORDER: STYRACOSTERNIPIDA NOVA

ICHNOPARVORDER: HADROSAURIFORMIPEDIDA NOVA

Ichnosuperfamily: IGUANODONTIDEA Nova

Ichnofamily: IGUANODONTIPODIDAE emend Vylov, 1988

Ichnogenus: Caririchnium LEONARDI, 1984
C. leonardii LOCKLEY, 1988

INCHNOCLASS: AVIPEDIA VIALOV, 1966

Incertae sedis

Ichnogenus: Aquatilavips CURRIE, 1981
A. curriei McCREA & SARJEANT, 2001

Trias, bipede indet

Ichnogenus: Typopus E. HITCHCOCK, 1845
T. abnormis (E. HITCHCOCK, 1845) E. HITCHCOCK, 1845

Bibliography